Fire ecology of Scots pine in Northwest Europe

Keywords: biodiversity, fire ecology, fuel modelling, succession, tree regenerationIn this thesis the ecological consequences of forest fire are studied in North-west European Scots pine {Pinus sylvestris) forests. The focus is on post-fire succession, and the factors and mechanisms that influence the successional pathways after fire. Fuel load and fuel moisture determine the intensity of forest fire and thus the degree of humus consumption. In a controlled laboratory' experiment humus consumption was determined for different moisture levels. Experimental fires showed evidence that variation in precipitation throughfall causes spatial variation in humus consumption in the stand through differences in humus moisture with respect to tree crowns. Humus consumption influences tree mortality, growth of remaining trees and re-colonization. Surface fires in Scots pine plantations caused a partial reduction of the litter and humus layers and a high mortality in the smaller trees. Reduction in radial growth after surface fire was variable, and was less in large diameter trees and in trees that experienced less humus consumption around their stem bases. Experimental burning of the humus layer showed that increased removal of organic material by fire resulted in an increase in seedling numbers. Earlier studies have suggested that the charcoal produced by fire improves germination conditions by absorbing phytotoxins produced by ericaceous species. All such studies have used activated carbon as a standardized model for charcoal. Bioassays with pine seeds in aqueous extracts of Vaccinium myrtillus and Calluna vulgaris showed toxic effects of the two species, but charcoal reduced toxicity less than activated carbon. Therefore, those previous studies have overestimated the effect of charcoal on germination, likely because of the considerably higher active surface area of activated carbon. The post-fire tree cohort after severe but small-scaled fires in Scots pine stands mainly consisted of Scots pine, but also birch and aspen. Compared to succession after other disturbance types in Scots pine stands, such as windthrow or soil scarification, seedling numbers are higher after small-scale fires by a magnitude often.Based on the good regeneration and for the purpose of fuel load reduction in areas with increased fire hazard, the prescribed burning of Scots pine stands should be reconsidered. Controlled forest fires could be used as an additional silvicultural technique to regenerate and transform single-species pine stands into mixed and more natural forests

Effecten van brand op bodem en vegetatie in dennenbossen van voedselarme zandgronden bij Kootwijk

Door atmosferische stikstofdepositie zijn veel bossen van de arme zandgronden vermest en in hun verdere successie verstoord. In een voedselarm dennenbos op zandgrond is onderzocht of door brand de overmaat aan stikstof die in de strooisellaag ligt opgeslagen kan worden teruggedrongen en de bosontwikkeling kan worden teruggezet naar een voedselarme pionierfase. Het onderzoek werd uitgevoerd bij Kootwijk waar in 1995 een bosbrand woedde. De vegetatie- en bodemkundige toestand van voor de brand, één en negen jaar na de brand werd geanalyseerd. Een belangrijke conclusie is dat brand eerder tot een verrijking van de bodem leidt dan tot een verschraling. Door brand vindt een verplaatsing van elementen van ectorganische naar endorganische horizonten plaats. Weliswaar verdwijnt er in zijn totaliteit stikstof, maar direct na de brand is de bodem sterk verrijkt met kalium, ammonium en fosfaat en in zuurgraad gedaald. Bovendien wordt het beschikbaar gekomen ammonium door micro-organismen vastgelegd in organische stof wat daardoor een rijk karakter krijgt. Dit effect is negen jaar na de brand nog steeds aanwezig. Het laten staan van de necromassa na de brand, zoals in Kootwijk plaatsvond, draagt niet bij aan het terugzetten van de vegetatiesuccessie naar een initieel schraal stadium, maar wel aan de diversiteit van bodemeigenschappe

Anisotropic flow of charged hadrons, pions and (anti-)protons measured at high transverse momentum in Pb-Pb collisions at sNN=2.76\sqrt{s_{\rm NN}}=2.76 TeV

The elliptic, $v_2$, triangular, $v_3$, and quadrangular, $v_4$, azimuthal anisotropic flow coefficients are measured for unidentified charged particles, pions and (anti-)protons in Pb-Pb collisions at $\sqrt{s_{\rm NN}} = 2.76$ TeV with the ALICE detector at the Large Hadron Collider. Results obtained with the event plane and four-particle cumulant methods are reported for the pseudo-rapidity range $|\eta|<0.8$ at different collision centralities and as a function of transverse momentum, $p_{\rm T}$, out to $p_{\rm T}=20$ GeV/$c$. The observed non-zero elliptic and triangular flow depends only weakly on transverse momentum for $p_{\rm T}>8$ GeV/$c$. The small $p_{\rm T}$ dependence of the difference between elliptic flow results obtained from the event plane and four-particle cumulant methods suggests a common origin of flow fluctuations up to $p_{\rm T}=8$ GeV/$c$. The magnitude of the (anti-)proton elliptic and triangular flow is larger than that of pions out to at least $p_{\rm T}=8$ GeV/$c$ indicating that the particle type dependence persists out to high $p_{\rm T}$.Comment: 16 pages, 5 captioned figures, authors from page 11, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/186

Centrality dependence of charged particle production at large transverse momentum in Pb-Pb collisions at sNN=2.76\sqrt{s_{\rm{NN}}} = 2.76 TeV

The inclusive transverse momentum ($p_{\rm T}$) distributions of primary charged particles are measured in the pseudo-rapidity range $|\eta|<0.8$ as a function of event centrality in Pb-Pb collisions at $\sqrt{s_{\rm{NN}}}=2.76$ TeV with ALICE at the LHC. The data are presented in the $p_{\rm T}$ range $0.15<p_{\rm T}<50$ GeV/$c$ for nine centrality intervals from 70-80% to 0-5%. The Pb-Pb spectra are presented in terms of the nuclear modification factor $R_{\rm{AA}}$ using a pp reference spectrum measured at the same collision energy. We observe that the suppression of high-$p_{\rm T}$ particles strongly depends on event centrality. In central collisions (0-5%) the yield is most suppressed with $R_{\rm{AA}}\approx0.13$ at $p_{\rm T}=6$-7 GeV/$c$. Above $p_{\rm T}=7$ GeV/$c$, there is a significant rise in the nuclear modification factor, which reaches $R_{\rm{AA}} \approx0.4$ for $p_{\rm T}>30$ GeV/$c$. In peripheral collisions (70-80%), the suppression is weaker with $R_{\rm{AA}} \approx 0.7$ almost independently of $p_{\rm T}$. The measured nuclear modification factors are compared to other measurements and model calculations.Comment: 17 pages, 4 captioned figures, 2 tables, authors from page 12, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/284

Measurement of charm production at central rapidity in proton-proton collisions at s=2.76\sqrt{s} = 2.76 TeV

The $p_{\rm T}$-differential production cross sections of the prompt (B feed-down subtracted) charmed mesons D$^0$, D$^+$, and D$^{*+}$ in the rapidity range $|y|<0.5$, and for transverse momentum $1< p_{\rm T} <12$ GeV/$c$, were measured in proton-proton collisions at $\sqrt{s} = 2.76$ TeV with the ALICE detector at the Large Hadron Collider. The analysis exploited the hadronic decays D$^0 \rightarrow$K$\pi$, D$^+ \rightarrow$K$\pi\pi$, D$^{*+} \rightarrow$D$^0\pi$, and their charge conjugates, and was performed on a $L_{\rm int} = 1.1$ nb$^{-1}$ event sample collected in 2011 with a minimum-bias trigger. The total charm production cross section at $\sqrt{s} = 2.76$ TeV and at 7 TeV was evaluated by extrapolating to the full phase space the $p_{\rm T}$-differential production cross sections at $\sqrt{s} = 2.76$ TeV and our previous measurements at $\sqrt{s} = 7$ TeV. The results were compared to existing measurements and to perturbative-QCD calculations. The fraction of cdbar D mesons produced in a vector state was also determined.Comment: 20 pages, 5 captioned figures, 4 tables, authors from page 15, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/307

Two-pion Bose-Einstein correlations in central Pb-Pb collisions at sNN\sqrt{s_{\rm NN}} = 2.76 TeV

The first measurement of two-pion Bose-Einstein correlations in central Pb-Pb collisions at $\sqrt{s_{\rm NN}} = 2.76$ TeV at the Large Hadron Collider is presented. We observe a growing trend with energy now not only for the longitudinal and the outward but also for the sideward pion source radius. The pion homogeneity volume and the decoupling time are significantly larger than those measured at RHIC.Comment: 17 pages, 5 captioned figures, 1 table, authors from page 12, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/388

Suppression of charged particle production at large transverse momentum in central Pb-Pb collisions at sNN=2.76\sqrt{s_{\rm NN}} = 2.76 TeV

Inclusive transverse momentum spectra of primary charged particles in Pb-Pb collisions at $\sqrt{s_{_{\rm NN}}}$ = 2.76 TeV have been measured by the ALICE Collaboration at the LHC. The data are presented for central and peripheral collisions, corresponding to 0-5% and 70-80% of the hadronic Pb-Pb cross section. The measured charged particle spectra in $|\eta|<0.8$ and $0.3 < p_T < 20$ GeV/$c$ are compared to the expectation in pp collisions at the same $\sqrt{s_{\rm NN}}$, scaled by the number of underlying nucleon-nucleon collisions. The comparison is expressed in terms of the nuclear modification factor $R_{\rm AA}$. The result indicates only weak medium effects ($R_{\rm AA} \approx$ 0.7) in peripheral collisions. In central collisions, $R_{\rm AA}$ reaches a minimum of about 0.14 at $p_{\rm T}=6$-7GeV/$c$ and increases significantly at larger $p_{\rm T}$. The measured suppression of high-$p_{\rm T}$ particles is stronger than that observed at lower collision energies, indicating that a very dense medium is formed in central Pb-Pb collisions at the LHC.Comment: 15 pages, 5 captioned figures, 3 tables, authors from page 10, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/98

Particle-yield modification in jet-like azimuthal di-hadron correlations in Pb-Pb collisions at sNN\sqrt{s_{\rm NN}} = 2.76 TeV

The yield of charged particles associated with high-$p_{\rm T}$ trigger particles ($8 < p_{\rm T} < 15$ GeV/$c$) is measured with the ALICE detector in Pb-Pb collisions at $\sqrt{s_{\rm NN}}$ = 2.76 TeV relative to proton-proton collisions at the same energy. The conditional per-trigger yields are extracted from the narrow jet-like correlation peaks in azimuthal di-hadron correlations. In the 5% most central collisions, we observe that the yield of associated charged particles with transverse momenta $p_{\rm T}> 3$ GeV/$c$ on the away-side drops to about 60% of that observed in pp collisions, while on the near-side a moderate enhancement of 20-30% is found.Comment: 15 pages, 2 captioned figures, 1 table, authors from page 10, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/350

Transverse momentum spectra of charged particles in proton-proton collisions at s=900\sqrt{s} = 900 GeV with ALICE at the LHC

The inclusive charged particle transverse momentum distribution is measured in proton-proton collisions at $\sqrt{s} = 900$ GeV at the LHC using the ALICE detector. The measurement is performed in the central pseudorapidity region $(|\eta|<0.8)$ over the transverse momentum range $0.15<p_{\rm T}<10$ GeV/$c$. The correlation between transverse momentum and particle multiplicity is also studied. Results are presented for inelastic (INEL) and non-single-diffractive (NSD) events. The average transverse momentum for $|\eta|<0.8$ is $\left<p_{\rm T}\right>_{\rm INEL}=0.483\pm0.001$ (stat.) $\pm0.007$ (syst.) GeV/$c$ and \left_{\rm NSD}=0.489\pm0.001 (stat.) $\pm0.007$ (syst.) GeV/$c$, respectively. The data exhibit a slightly larger $\left<p_{\rm T}\right>$ than measurements in wider pseudorapidity intervals. The results are compared to simulations with the Monte Carlo event generators PYTHIA and PHOJET.Comment: 20 pages, 8 figures, 2 tables, published version, figures at http://aliceinfo.cern.ch/ArtSubmission/node/390

Inclusive J/psi production in pp collisions at sqrt(s) = 2.76 TeV

The ALICE Collaboration has measured inclusive J/psi production in pp collisions at a center of mass energy sqrt(s)=2.76 TeV at the LHC. The results presented in this Letter refer to the rapidity ranges |y|<0.9 and 2.5<y<4 and have been obtained by measuring the electron and muon pair decay channels, respectively. The integrated luminosities for the two channels are L^e_int=1.1 nb^-1 and L^mu_int=19.9 nb^-1, and the corresponding signal statistics are N_J/psi^e+e-=59 +/- 14 and N_J/psi^mu+mu-=1364 +/- 53. We present dsigma_J/psi/dy for the two rapidity regions under study and, for the forward-y range, d^2sigma_J/psi/dydp_t in the transverse momentum domain 0<p_t<8 GeV/c. The results are compared with previously published results at sqrt(s)=7 TeV and with theoretical calculations.Comment: 7 figures, 3 tables, accepted for publication in Phys. Lett.