The Dark Halo - Spheroid Conspiracy and the Origin of Elliptical Galaxies

Dynamical modeling and strong lensing data indicate that the total density profiles of early-type galaxies are close to isothermal, i.e., rho_tot ~ r^gamma with gamma approx -2. To understand the origin of this universal slope we study a set of simulated spheroids formed in isolated binary mergers as well as the formation within the cosmological framework. The total stellar plus dark matter density profiles can always be described by a power law with an index of gamma approx -2.1 with a tendency toward steeper slopes for more compact, lower-mass ellipticals. In the binary mergers the amount of gas involved in the merger determines the precise steepness of the slope. This agrees with results from the cosmological simulations where ellipticals with steeper slopes have a higher fraction of stars formed in situ. Each gas-poor merger event evolves the slope toward gamma ~ -2, once this slope is reached further merger events do not change it anymore. All our ellipticals have flat intrinsic combined stellar and dark matter velocity dispersion profiles. We conclude that flat velocity dispersion profiles and total density distributions with a slope of gamma ~ -2 for the combined system of stars and dark matter act as a natural attractor. The variety of complex formation histories as present in cosmological simulations, including major as well as minor merger events, is essential to generate the full range of observed density slopes seen for present-day elliptical galaxies.Comment: Accepted by the Astrophysical Journal, 17 pages, 12 figure

Short-lived star-forming giant clumps in cosmological simulations of z~2 disks

Many observed massive star-forming z\approx2 galaxies are large disks that exhibit irregular morphologies, with \sim1kpc, \sim10^(8-10)Msun clumps. We present the largest sample to date of high-resolution cosmological SPH simulations that zoom-in on the formation of individual M*\sim10^(10.5)Msun galaxies in \sim10^(12)Msun halos at z\approx2. Our code includes strong stellar feedback parameterized as momentum-driven galactic winds. This model reproduces many characteristic features of this observed class of galaxies, such as their clumpy morphologies, smooth and monotonic velocity gradients, high gas fractions (f_g\sim50%) and high specific star-formation rates (\gtrsim1Gyr^(-1)). In accord with recent models, giant clumps (Mclump\sim(5x10^8-10^9)Msun) form in-situ via gravitational instabilities. However, the galactic winds are critical for their subsequent evolution. The giant clumps we obtain are short-lived and are disrupted by wind-driven mass loss. They do not virialise or migrate to the galaxy centers as suggested in recent work neglecting strong winds. By phenomenologically implementing the winds that are observed from high-redshift galaxies and in particular from individual clumps, our simulations reproduce well new observational constraints on clump kinematics and clump ages. In particular, the observation that older clumps appear closer to their galaxy centers is reproduced in our simulations, as a result of inside-out formation of the disks rather than inward clump migration.Comment: 11 pages, 6 figures, 1 table. Accepted for publication in the Astrophysical Journa

Forming Early-Type Galaxies in LambdaCDM Simulations -I. Assembly histories

We present a sample of nine high resolution cosmological simulations in the mass range of M_vir=7x10^11-4x10^12 M_sun starting from LambdaCDM initial conditions. Our simulations include primordial radiative cooling, photoionization, star formation, supernova II feedback, but exclude supernova driven winds and AGN feedback. The simulated galaxies assemble in two phases, with the initial growth dominated by compact (r<r_eff) in situ star formation fueled by cold, low entropy gas streams resulting in a very similar mean assembly redshift of z_{f,ins}~2.5 for the in situ stellar component in all galaxies. The late growth is dominated by accretion of old stars formed in subunits outside the main galaxy (r>r_eff) resulting in an assembly redshift of z_{f,acc}~0.5-1.5 with much larger scatter. We find a positive correlation between the fraction of accreted stars and the final mass of our galaxies. We show that gravitational feedback strongly suppresses late star formation in massive galaxies contributing to the observed galaxy color bimodality. The accretion of stellar material is also responsible for the observed size growth of early-type galaxies. In addition, we find that the dark matter fractions within the stellar half-mass radii continuously increase towards lower redshift from about f_DM~0.05 at z~3 to f_DM~0.1-0.3 at z=0. Furthermore, the logarithmic slope of the total density profile is nearly isothermal at the present-day (gamma'~1.9-2.2). Finally, the input of gravitational heating lowers the central dark matter densities in the galaxies, with the effect being smaller compared to simulations without supernova feedback.Comment: 23 pages, 16 figures, accepted for publication in Ap

The Two Phases of Galaxy Formation

Cosmological simulations of galaxy formation appear to show a two-phase character with a rapid early phase at z>2 during which in-situ stars are formed within the galaxy from infalling cold gas followed by an extended phase since z<3 during which ex-situ stars are primarily accreted. In the latter phase massive systems grow considerably in mass and radius by accretion of smaller satellite stellar systems formed at quite early times (z>3) outside of the virial radius of the forming central galaxy. These tentative conclusions are obtained from high resolution re-simulations of 39 individual galaxies in a full cosmological context with present-day virial halo masses ranging from 7e11 M_sun h^-1 < M_vir < 2.7e13 M_sun h^-1 and central galaxy masses between 4.5e10 M_sun h^-1 < M_* < 3.6e11 M_sun h^-1. The simulations include the effects of a uniform UV background, radiative cooling, star formation and energetic feedback from SNII. The importance of stellar accretion increases with galaxy mass and towards lower redshift. In our simulations lower mass galaxies ($M_* < 9e10 M_sun h^-1) accrete about 60 per cent of their present-day stellar mass. High mass galaxy ($M_* > 1.7e11 M_sun h^-1) assembly is dominated by accretion and merging with about 80 per cent of the stars added by the present-day. In general the simulated galaxies approximately double their mass since z=1. For massive systems this mass growth is not accompanied by significant star formation. The majority of the in-situ created stars is formed at z>2, primarily out of cold gas flows. We recover the observational result of archaeological downsizing, where the most massive galaxies harbor the oldest stars. We find that this is not in contradiction with hierarchical structure formation. Most stars in the massive galaxies are formed early on in smaller structures, the galaxies themselves are assembled late.Comment: 13 pages, 13 figures, accepted for publication in Ap

Assembly of the Red Sequence in Infrared-Selected Galaxy Clusters from the IRAC Shallow Cluster Survey

We present results for the assembly and star formation histories of massive (~L*) red sequence galaxies in 11 spectroscopically confirmed, infrared-selected galaxy clusters at 1.0 < z < 1.5, the precursors to present-day massive clusters with M ~ 10^15 M_sun. Using rest-frame optical photometry, we investigate evolution in the color and scatter of the red sequence galaxy population, comparing with models of possible star formation histories. In contrast to studies of central cluster galaxies at lower redshift (z < 1), these data are clearly inconsistent with the continued evolution of stars formed and assembled primarily at a single, much-earlier time. Specifically, we find that the colors of massive cluster galaxies at z = 1.5 imply that the bulk of star formation occurred at z ~ 3, whereas by z = 1 their colors imply formation at z ~ 2; therefore these galaxies exhibit approximately the same luminosity-weighted stellar age at 1 < z < 1.5. This likely reflects star formation that occurs over an extended period, the effects of significant progenitor bias, or both. Our results generally indicate that massive cluster galaxy populations began forming a significant mass of stars at z >~ 4, contained some red spheroids by z ~ 1.5, and were actively assembling much of their final mass during 1 < z < 2 in the form of younger stars. Qualitatively, the slopes of the cluster color-magnitude relations are consistent with no significant evolution relative to local clusters.Comment: 24 pages, 9 figures, accepted to Ap

The cosmological size and velocity dispersion evolution of massive early-type galaxies

We analyze 40 cosmological re-simulations of individual massive galaxies with present-day stellar masses of $M_{*} > 6.3 \times 10^{10} M_{\odot}$ in order to investigate the physical origin of the observed strong increase in galaxy sizes and the decrease of the stellar velocity dispersions since redshift $z \approx 2$. At present 25 out of 40 galaxies are quiescent with structural parameters (sizes and velocity dispersions) in agreement with local early type galaxies. At z=2 all simulated galaxies with $M_* \gtrsim 10^{11}M_{\odot}$ (11 out of 40) at z=2 are compact with projected half-mass radii of $\approx$ 0.77 ($\pm$0.24) kpc and line-of-sight velocity dispersions within the projected half-mass radius of $\approx$ 262 ($\pm$28) kms$^{-1}$ (3 out of 11 are already quiescent). Similar to observed compact early-type galaxies at high redshift the simulated galaxies are clearly offset from the local mass-size and mass-velocity dispersion relations. Towards redshift zero the sizes increase by a factor of $\sim 5-6$, following $R_{1/2} \propto (1+z)^{\alpha}$ with $\alpha = -1.44$ for quiescent galaxies ($\alpha = -1.12$ for all galaxies). The velocity dispersions drop by about one-third since $z \approx 2$, following $\sigma_{1/2} \propto (1+z)^{\beta}$ with $\beta = 0.44$ for the quiescent galaxies ($\beta = 0.37$ for all galaxies). The simulated size and dispersion evolution is in good agreement with observations and results from the subsequent accretion and merging of stellar systems at $z\lesssim 2$ which is a natural consequence of the hierarchical structure formation. A significant number of the simulated massive galaxies (7 out of 40) experience no merger more massive than 1:4 (usually considered as major mergers). On average, the dominant accretion mode is stellar minor mergers with a mass-weighted mass-ratio of 1:5. (abridged)Comment: 9 pages, 6 figures, accepted for publication in Ap

CANDELS Observations of the Structural Properties and Evolution of Galaxies in a Cluster at z=1.62

We discuss the structural and morphological properties of galaxies in a z=1.62 proto-cluster using near-IR imaging data from Hubble Space Telescope Wide Field Camera 3 data of the Cosmic Assembly Near-IR Deep Extragalactic Legacy Survey (CANDELS). The cluster galaxies exhibit a clear color-morphology relation: galaxies with colors of quiescent stellar populations generally have morphologies consistent with spheroids, and galaxies with colors consistent with ongoing star formation have disk-like and irregular morphologies. The size distribution of the quiescent cluster galaxies shows a deficit of compact (< 1kpc), massive galaxies compared to CANDELS field galaxies at z=1.6. As a result the cluster quiescent galaxies have larger average effective sizes compared to field galaxies at fixed mass at greater than 90% significance. Combined with data from the literature, the size evolution of quiescent cluster galaxies is relatively slow from z~1.6 to the present, growing as (1+z)^(-0.6+/-0.1). If this result is generalizable, then it implies that physical processes associated with the denser cluster region seems to have caused accelerated size growth in quiescent galaxies prior to z=1.6 and slower subsequent growth at z<1.6 compared to galaxies in the lower density field. The quiescent cluster galaxies at z=1.6 have higher ellipticities compared to lower redshift samples at fixed mass, and their surface-brightness profiles suggest that they contain extended stellar disks. We argue the cluster galaxies require dissipationless (i.e., gas-poor or "dry") mergers to reorganize the disk material and to match the relations for ellipticity, stellar mass, size, and color of early-type galaxies in z<1 clusters.Comment: Accepted for publication in ApJ. 14 pages in emulateapj format. Replacement includes improvements from referee report, and updates and additions to reference

A High Statistics Search for Ultra-High Energy Gamma-Ray Emission from Cygnus X-3 and Hercules X-1

We have carried out a high statistics (2 Billion events) search for ultra-high energy gamma-ray emission from the X-ray binary sources Cygnus X-3 and Hercules X-1. Using data taken with the CASA-MIA detector over a five year period (1990-1995), we find no evidence for steady emission from either source at energies above 115 TeV. The derived upper limits on such emission are more than two orders of magnitude lower than earlier claimed detections. We also find no evidence for neutral particle or gamma-ray emission from either source on time scales of one day and 0.5 hr. For Cygnus X-3, there is no evidence for emission correlated with the 4.8 hr X-ray periodicity or with the occurrence of large radio flares. Unless one postulates that these sources were very active earlier and are now dormant, the limits presented here put into question the earlier results, and highlight the difficulties that possible future experiments will have in detecting gamma-ray signals at ultra-high energies.Comment: 26 LaTeX pages, 16 PostScript figures, uses psfig.sty to be published in Physical Review

Sussing merger trees: the impact of halo merger trees on galaxy properties in a semi-analytic model

A halo merger tree forms the essential backbone of a semi-analytic model for galaxy formation and evolution. Recent studies have pointed out that extracting merger trees from numerical simulations of structure formation is non-trivial; different tree building algorithms can give differing merger histories. These differences should be carefully understood before merger trees are used as input for models of galaxy formation. We investigate the impact of different halo merger trees on a semi-analytic model. We find that the z = 0 galaxy properties in our model show differences between trees when using a common parameter set. The star formation history of the universe and the properties of satellite galaxies can show marked differences between trees with different construction methods. Independently calibrating the semi-analytic model for each tree can reduce the discrepancies between the z = 0 global galaxy properties, at the cost of increasing the differences in the evolutionary histories of galaxies. Furthermore, the underlying physics implied can vary, resulting in key quantities such as the supernova feedback efficiency differing by factors of 2. Such a change alters the regimes where star formation is primarily suppressed by supernovae. Therefore, halo merger trees extracted from a common halo catalogue using different, but reliable, algorithms can result in a difference in the semi-analytic model. Given the uncertainties in galaxy formation physics, however, these differences may not necessarily be viewed as significant

Nck2 promotes human melanoma cell proliferation, migration and invasion in vitro and primary melanoma-derived tumor growth in vivo

<p>Abstract</p> <p>Background</p> <p>Nck1 and Nck2 adaptor proteins are involved in signaling pathways mediating proliferation, cytoskeleton organization and integrated stress response. Overexpression of Nck1 in fibroblasts has been shown to be oncogenic. Through the years this concept has been challenged and the consensus is now that overexpression of either Nck cooperates with strong oncogenes to transform cells. Therefore, variations in Nck expression levels in transformed cells could endorse cancer progression.</p> <p>Methods</p> <p>Expression of Nck1 and Nck2 proteins in various cancer cell lines at different stages of progression were analyzed by western blots. We created human primary melanoma cell lines overexpressing GFP-Nck2 and investigated their ability to proliferate along with metastatic characteristics such as migration and invasion. By western blot analysis, we compared levels of proteins phosphorylated on tyrosine as well as cadherins and integrins in human melanoma cells overexpressing or not Nck2. Finally, in mice we assessed tumor growth rate of human melanoma cells expressing increasing levels of Nck2.</p> <p>Results</p> <p>We found that expression of Nck2 is consistently increased in various metastatic cancer cell lines compared with primary counterparts. Particularly, we observed significant higher levels of Nck2 protein and mRNA, as opposed to no change in Nck1, in human metastatic melanoma cell lines compared with non-metastatic melanoma and normal melanocytes. We demonstrated the involvement of Nck2 in proliferation, migration and invasion in human melanoma cells. Moreover, we discovered that Nck2 overexpression in human primary melanoma cells correlates with higher levels of proteins phosphorylated on tyrosine residues, assembly of Nck2-dependent pY-proteins-containing molecular complexes and downregulation of cadherins and integrins. Importantly, we uncovered that injection of Nck2-overexpressing human primary melanoma cells into mice increases melanoma-derived tumor growth rate.</p> <p>Conclusions</p> <p>Collectively, our data indicate that Nck2 effectively influences human melanoma phenotype progression. At the molecular level, we propose that Nck2 in human primary melanoma promotes the formation of molecular complexes regulating proliferation and actin cytoskeleton dynamics by modulating kinases or phosphatases activities that results in increased levels of proteins phosphorylated on tyrosine residues. This study provides new insights regarding cancer progression that could impact on the therapeutic strategies targeting cancer.</p