Le forme della regalità nella Roma latino-sabina

The analysis of historical sources, onomastics data, and the festive calendar, shows that the most archaic Roman kingship was structured in the form of a diarchy between a lifelong king-priest and a temporary warrior-king: the latter ruled in the season of war, while in peacetime the city government was led in turn by the patres (interregnum). This diarchy could be the result of the influence, on Roman institutions, of the constitutional structures of the Greeks and of the Italic peoples, since among other Indo-European cultures, like Vedic India and the Celts, the pattern seems rather to be a triad — composed of a priest, a supreme king and a warrior king — surviving in Rome only at a theological level in the Pre-Capitoline triad Iuppiter Mars Quirinus. Given these premises, the subsequent Etruscan monarchy appears to be the result of the rising of the army commander to a tyrannical and lifelong power, and of the marginalization of the rex sacrorum, while the Republic seems a partial restoration of the oldest constitution

Preadaptation and post-introduction evolution facilitate the invasion of Phragmites australis in North America

Compared with non-invasive species, invasive plant species may benefit from certain advantageous traits, for example, higher photosynthesis capacity and resource/energy-use efficiency. These traits can be preadapted prior to introduction, but can also be acquired through evolution following introduction to the new range. Disentangling the origins of these advantageous traits is a fundamental and emerging question in invasion ecology. We conducted a multiple comparative experiment under identical environmental condition with the invasive haplotype M lineage of the wetland grass Phragmites australis and compared the ecophysiological traits of this invasive haplotype M in North America with those of the European ancestor and the conspecific North American native haplotype E lineage, P. australis ssp. americanus. The invasive haplotype M differed significantly from the native North American conspecific haplotype E in several ecophysiological and morphological traits, and the European haplotype M had a more efficient photosynthetic apparatus than the native North American P. australis ssp. americanus. Within the haplotype M lineage, the introduced North American P. australis exhibited different biomass allocation patterns and resource/energy-use strategies compared to its European ancestor group. A discriminant analysis of principal components separated the haplotype M and the haplotype E lineages completely along the first canonical axis, highly related to photosynthetic gas-exchange parameters, photosynthetic energy-use efficiency and payback time. The second canonical axis, highly related to photosynthetic nitrogen use efficiency and construction costs, significantly separated the introduced P. australis in North America from its European ancestor. Synthesis. We conclude that the European P. australis lineage was preadapted to be invasive prior to its introduction, and that the invasion in North America is further stimulated by rapid post-introduction evolution in several advantageous traits. The multicomparison approach used in this study could be an effective approach for distinguishing preadaptation and post-introduction evolution of invasive species. Further research is needed to link the observed changes in invasive traits to the genetic variation and the interaction with the environment. Disentangling the origins of the advantageous traits is a fundamental and emerging question in invasion ecology. We compared the ecophysiological traits of the invasive haplotype M of Phragmites australis in North America with those in Europe and the conspecific North American native haplotype E lineage, P. australis ssp. americanus. The invasive haplotype M differed significantly from the native North American conspecific haplotype E in several ecophysiological and morphological traits, and the European haplotype M had a more efficient photosynthetic apparatus than the native North American P. australis ssp. americanus. Within the haplotype M lineage, the introduced North American P. australis exhibited different biomass allocation patterns and resource/energy-use strategies compared to its European ancestor group. A discriminant analysis of principal components (DAPC) significantly separated the three groups. We conclude that the European P. australis lineage was preadapted to be invasive prior to its introduction, and that the invasion in North America is further stimulated by rapid post-introduction evolution in several advantageous traits