Mussels with Meat: Bivalve Tissue-Shell Radiocarbon Age Differences and Archaeological Implications

Local reservoir ages are often estimated from the difference between the radiocarbon ages of aquatic material and associated terrestrial samples for which no reservoir effect is expected. Frequently, the selected aquatic material consists of bivalve shells that are typically well preserved in the archaeological record. For instance, large shell middens attest to the importance of mussel consumption at both coastal and inland sites. However, different physiological mechanisms associated with tissue and shell growth may result in differences in reservoir effects between the surviving component (shell) and the component relevant to dietary reservoir effects in consumers (tissue). The current study examines bivalve tissue-shell age differences both from freshwater and marine contexts close to archaeological sites where human consumption of mollusks has been attested. Results exhibited significant 14C age differences between bivalve tissue and shell in a freshwater context. In a marine context, no significant bivalve tissue-shell age differences were observed. The results also showed that riverine and lacustrine shells show large and variable freshwater reservoir effects. The results have important implications for establishing local reservoir effects especially in a freshwater environment. For good a priori knowledge of expected 14C differences in organic and inorganic water, carbon is thus necessary. Furthermore, the high variability in freshwater shell 14C ages implies the need for representative sampling from the archaeological record

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TAFELBAND Das Wohnhaus des Arbeiters (-) Tafelband (2) (-) Einband (-) Titelseite (-) Tafel I: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel I) Tafel II: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel II) Tafel III: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel III) Tafel IV: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel IV) Tafel V: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel V) Tafel VI: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel VI) Tafel VII: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel VII) Tafel VIII: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel VIII) Tafel IX: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel IX) Tafel X: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel X) Tafel XI: Beispiel eines Arbeiterwohnhauses, Ansicht, Schnitt, Grundriss (Tafel XI) Tafel XII: Lageplan einer Arbeiterwohnsiedlung (Tafel XII) Farbinformation (-) Einband (-

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TEXTBEILAGE Das Wohnhaus des Arbeiters (-) Textbeilage (1) (-) Titelseite (-) Vorwort (-) Inhaltsverzeichnis (-) Erster Theil ([1]) I. Allgemeine Grundsätze, welche bei der Anlage von Arbeiterwohnungen maßgebend sind ([1]) 1. Die Lage der Arbeiterwohnungen ([1]) 2. Disposition, Einrichtung und Größe der einzelnen Räume (7) 3. Isolirung, Heizung und Ventilation (12) 4. Die Wahl der Baumaterialien und die Bau-Ausführung (20) Zweiter Theil (30) Massenberechung (32) Auszug der Maurermaterialien (37) Kosten-Anschlag (39

Adaptations and transformations of hunter-gatherers in forest environments: New archaeological and anthropological insights

Like any other living being, humans constantly influence their environment, be it intentionally or unintentionally. By extracting natural resources, they shape their environment and also that of plants and other animals. A great difference setting people apart from all other living beings is the ability to construct and develop their own niche intentionally, and the unique tool for this is cultural behaviour. Here, we discuss anthropogenic environmental changes of hunter-gatherers and present new palaeoecological and palynological data. The studies are framed with ethnoarchaeological data from Western Siberia to gain a better understanding of how different triggers lead to coping mechanisms. For archaeological implication, we use two Mesolithic case studies from Germany: One of them focuses on hazelnut economy around ancient Lake Duvensee, and the other broaches the issue of selective roe deer hunt and its consequences at the site of Friesack. We address the archaeological evidence from the perspective of active alteration and its consequences, starting our argumentation from a perspective of niche construction theory. This approach has rarely been applied to early Holocene hunter-gatherers in Northern Europe even though the available data render possible to discuss human–environment interaction from such a perspective. It is demonstrated that archaeological research has tools at hand that enables to detect anthropogenic niche construction. However, the ethnoarchaeological example shows limitations and archaeologically invisible triggers and consequent results of human adaptations. The critical revision of such perspectives based on empirical data provides a better understanding of social and environmental transformations in the early- and mid-Holocene

The prelude to industrial whaling:Identifying the targets of ancient European whaling using zooarchaeology and collagen mass-peptide fingerprinting

Taxonomic identification of whale bones found during archaeological excavations is problematic due to their typically fragmented state. This difficulty limits understanding of both the past spatio-temporal distributions of whale populations and of possible early whaling activities. To overcome this challenge, we performed zooarchaeology by mass spectrometry on an unprecedented 719 archaeological and palaeontological specimens of probable whale bone from Atlantic European contexts, predominantly dating from ca 3500 BCE to the eighteenth century CE. The results show high numbers of Balaenidae (many probably North Atlantic right whale (Eubalaena glacialis)) and grey whale (Eschrichtius robustus) specimens, two taxa no longer present in the eastern North Atlantic. This discovery matches expectations regarding the past utilization of North Atlantic right whales, but was unanticipated for grey whales, which have hitherto rarely been identified in the European zooarchaeological record. Many of these specimens derive from contexts associated with mediaeval cultures frequently linked to whaling: the Basques, northern Spaniards, Normans, Flemish, Frisians, Anglo-Saxons and Scandinavians. This association raises the likelihood that early whaling impacted these taxa, contributing to their extirpation and extinction. Much lower numbers of other large cetacean taxa were identified, suggesting that what are now the most depleted whales were once those most frequently used.</p

Historical Demographic Processes Dominate Genetic Variation in Ancient Atlantic Cod Mitogenomes

Ancient DNA (aDNA) approaches have been successfully used to infer the long-term impacts of climate change, domestication, and human exploitation in a range of terrestrial species. Nonetheless, studies investigating such impacts using aDNA in marine species are rare. Atlantic cod (Gadus morhua), is an economically important species that has experienced dramatic census population declines during the last century. Here, we investigated 48 ancient mitogenomes from historical specimens obtained from a range of archeological excavations in northern Europe dated up to 6,500 BCE. We compare these mitogenomes to those of 496 modern conspecifics sampled across the North Atlantic Ocean and adjacent seas. Our results confirm earlier observations of high levels of mitogenomic variation and a lack of mutation-drift equilibrium—suggestive of population expansion. Furthermore, our temporal comparison yields no evidence of measurable mitogenomic changes through time. Instead, our results indicate that mitogenomic variation in Atlantic cod reflects past demographic processes driven by major historical events (such as oscillations in sea level) and subsequent gene flow rather than contemporary fluctuations in stock abundance. Our results indicate that historical and contemporaneous anthropogenic pressures such as commercial fisheries have had little impact on mitogenomic diversity in a wide-spread marine species with high gene flow such as Atlantic cod. These observations do not contradict evidence that overfishing has had negative consequences for the abundance of Atlantic cod and the importance of genetic variation in implementing conservation strategies. Instead, these observations imply that any measures toward the demographic recovery of Atlantic cod in the eastern Atlantic, will not be constrained by recent loss of historical mitogenomic variation.</jats:p

Resurrection and redescription of Varestrongylus alces (Nematoda; Protostrongylidae), a lungworm of the Eurasian moose (Alces alces), with report on associated pathology

Varestrongylus alces, a lungworm in Eurasian moose from Europe has been considered a junior synonym of Varestrongylus capreoli, in European roe deer, due to a poorly detailed morphological description and the absence of a type-series. Methods Specimens used in the redescription were collected from lesions in the lungs of Eurasian moose, from Vestby, Norway. Specimens were described based on comparative morphology and integrated approaches. Molecular identification was based on PCR, cloning and sequencing of the ITS-2 region of the nuclear ribosomal DNA. Phylogenetic analysis compared V. alces ITS-2 sequences to these of other Varestrongylus species and other protostrongylids. Results Varestrongylus alces is resurrected for protostrongylid nematodes of Eurasian moose from Europe. Varestrongylus alces causes firm nodular lesions that are clearly differentiated from the adjacent lung tissue. Histologically, lesions are restricted to the parenchyma with adult, egg and larval parasites surrounded by multinucleated giant cells, macrophages, eosinophilic granulocytes, lymphocytes. The species is valid and distinct from others referred to Varestrongylus, and should be separated from V. capreoli. Morphologically, V. alces can be distinguished from other species by characters in the males that include a distally bifurcated gubernaculum, arched denticulate crura, spicules that are equal in length and relatively short, and a dorsal ray that is elongate and bifurcated. Females have a well-developed provagina, and are very similar to those of V. capreoli. Morphometrics of first-stage larvae largely overlap with those of other Varestrongylus. Sequences of the ITS-2 region strongly support mutual independence of V. alces, V. cf. capreoli, and the yet undescribed species of Varestrongylus from North American ungulates. These three taxa form a well-supported crown-clade as the putative sister of V. alpenae. The association of V. alces and Alces or its ancestors is discussed in light of host and parasite phylogeny and host historical biogeography. Varestrongylus alces is a valid species, and should be considered distinct from V. capreoli. Phylogenetic relationships among Varestrongylus spp. from Eurasia and North America are complex and consistent with faunal assembly involving recurrent events of geographic expansion, host switching and subsequent speciation. Cervidae, Cryptic species, Historical biogeography, ITS-2, Metastrongyloidea, Parasite biodiversity, Varestrongylinae, Varestrongylus capreoli, Verminous pneumoniapublishedVersio

The transition from foraging to farming (7000–500 cal BC) in the SE Baltic : A re-evaluation of chronological and palaeodietary evidence from human remains

Our knowledge of the timing and completeness of the transition from foraging, fishing and hunting to food production in boreal northeastern Europe is far from clear. Here, we present new bone collagen AMS 14C dates, and δ13C and δ15N isotope values for 20 humans and 17 animals from a 6500-year period dating from the Late Mesolithic to the Bronze Age in Lithuania. AMS 14C dates revealed large discrepancies in comparison to previously obtained radiocarbon dates, thus highlighting the need to re-date all prehistoric human remains where chronology was based on 14C dating of bone collagen. Stable isotope data indicate that inland Mesolithic-Subneolithic hunter-gatherers (7000–3000 cal BC) relied on a balance of freshwater food and game animals with regard to protein intake. The coastal Subneolithic groups (ca. 3000 cal BC) relied heavily on lagoon fishing, while seals and forest game were of lesser importance. Animal husbandry, most likely of sheep or goats, was a main source of protein for Neolithic Corded Ware Culture people (2900–2400 cal BC), although a significant contribution of freshwater food is also evident. Significant intra-individual variation in stable isotope values may demonstrate that a highly flexible subsistence strategy was adopted by the CWC people. Unusually high δ13C values indicate that millet had been already introduced into the farming economy of the Late Bronze Age around 1000 cal BC

Winter temperature and forest cover have shaped red deer distribution in Europe and the Ural Mountains since the Late Pleistocene

Aim: The Expansion-Contraction model has been used to explain the responses of species to climatic changes. During periods of unfavourable climatic conditions, species retreat to refugia from where they may later expand. This paper focuses on the palaeoecology of red deer over the past 54 ka across Europe and the Urals, to reveal patterns of change in their range and explore the role of environmental conditions in determining their distribution. Location: Europe and western Asia to 63°E. Taxon: Red deer (Cervus elaphus). Methods: We collected 984 records of radiocarbon-dated red deer subfossils from the Late Pleistocene and the Holocene, including 93 original dates. For each deer sample we compiled climatic and biome type data for the corresponding time intervals. Results: During the last 54 ka changes in red deer range in Europe and the Urals were asynchronous and differed between western and eastern Europe and western Asia due to different environmental conditions in those regions. The range of suitable areas for deer during the Last Glacial Maximum (LGM) was larger than previously thought and covered vast regions not only in southern but also in western and eastern Europe. Throughout the period investigated the majority of specimens inhabited forests in the temperate climatic zone. The contribution of forests in deer localities significantly decreased during the last 4 ka, due to deforestation of Europe caused by humans. Mean January temperature was the main limiting factor for species distribution. Over 90% of the samples were found in areas where mean January temperature was above −10°C. Main conclusions: Red deer response to climatic oscillations are in agreement with the Expansion-Contraction model but in contradiction to the statement of only the southernmost LGM refugia of the species. During the last 54 ka red deer occurred mostly in forests of the temperate climatic zone. © 2020 John Wiley & Sons Ltd.European Social Fund, Grant/Award Number: UDA-POKL.04.01.01-00-072/09-00; University of Wroclaw, Grant/Award Number: 0410/2990/18; Institute of Environmental Biology, University of Wrocław, Grant/Award Number: 0410/2990/18; Mammal Research Institute Polish Academy of Sciences; Narodowe Centrum Nauki , Grant/Award Number: DEC-2013/11/B/NZ8/00888 and UMO-2016/23/B/HS3/00387; Romanian National Authority for Scientific Research, UEFISCDI, Grant/Award Number: PN-IIIP4-ID-PCE-2016-0676; National Centre for Atmospheric Science and the Centre for Environmental Data Analysis, UK; Faunal Database of the Stage Three Project; Leverhulme Trust, Grant/Award Number: F00568W