The only known cyclopygid–‘atheloptic’ trilobite fauna from North America: the upper Ordovician fauna of the Pyle Mountain Argillite and its palaeoenvironmental significance

The trilobite fauna of the upper Ordovician (middle Katian) Pyle Mountain Argillite comprises a mixture of abundant mesopelagic cyclopygids and other pelagic taxa and a benthic fauna dominated by trilobites lacking eyes. Such faunas were widespread in deep water environments around Gondwana and terranes derived from that continent throughout Ordovician time but this is the only known record of such a fauna from North America and thus from Laurentia. It probably reflects a major sea level rise (the ‘Linearis drowning events’) as does the development of coeval cyclopygid-dominated deep water trilobite faunas in terranes that were marginal to Laurentia and are now preserved in Ireland and Scotland. The Pyle Mountain Argillite trilobite fauna occurs with a deep water Foliomena brachiopod fauna and comprises 22 species. Pelagic trilobites (mostly cyclopygids) constitute 36% of the preserved sclerites, and 45% of the fauna is the remains of trilobites lacking eyes, including one new species, Dindymene whittingtoni sp. nov. Three species of cyclopygid are present, belonging in Cyclopyge, Symphysops and Microparia (Heterocyclopyge). Cyclopygids are widely thought to have been stratified in the water column in life and thus their taxonomic diversity reflects the relative depths of the sea-beds on which their remains accumulated. A tabulation of middle and upper Katian cyclopygid-bearing faunas from several palaeoplates and terranes arranged on the basis of increasing numbers of cyclopygid genera allows an assessment of the relative depth ranges of the associated benthic taxa. The Pyle Mountain Argillite fauna lies towards the deeper end of this depth spectrum

Numerical calculation of strong-field laser-atom interaction: An approach with perfect reflection-free radiation boundary conditions

The time-dependent, single-particle Schrodinger equation with a finite-range potential is solved numerically on a three-dimensional spherical domain. In order to correctly account for outgoing waves, perfect reflection-free radiation boundary conditions are used on the surface of a sphere. These are computationally most effective if the particle wavefunction is expanded in the set of spherical harmonics and computations are performed in the Kramers-Henneberger accelerated frame. The method allows one to solve the full ionization dynamics in intense laser fields within a small region of atomic dimensions

Multifunctional and robust composite materials comprising gold nanoparticles at a spherical polystyrene particle surface

The preparation of composite particles comprising gold nanoparticles (4.5–26 nm) assembled at a polystyrene (PS) surface with tunable loading is reported with wide ranging potentials from cellular studies to catalysis.</p

Fossil AGN jets as ultra high energy particle accelerators

Remnants of AGN jets and their surrounding cocoons leave colossal magnetohydrodynamic (MHD) fossil structures storing total energies ~10^{60} erg. The original active galacic nucleus (AGN) may be dead but the fossil will retain its stable configuration resembling the reversed-field pinch (RFP) encountered in laboratory MHD experiments. In an RFP the longitudinal magnetic field changes direction at a critical distance from the axis, leading to magnetic re-connection there, and to slow decay of the large-scale RFP field. We show that this field decay induces large-scale electric fields which can accelerate cosmic rays with an E^{-2} power-law up to ultra-high energies with a cut-off depending on the fossil parameters. The cut-off is expected to be rigidity dependent, implying the observed composition would change from light to heavy close to the cut-off if one or two nearby AGN fossils dominate. Given that several percent of the universe's volume may house such slowly decaying structures, these fossils may even re-energize ultra-high energy cosmic rays from distant/old sources, offsetting the ``GZK-losses'' due to interactions with photons of the cosmic microwave background radiation and giving evidence of otherwise undetectable fossils. In this case the composition would remain light to the highest energies if distant sources or fossils dominated, but otherwise would be mixed. It is hoped the new generation of cosmic ray experiments such as the Pierre Auger Observatory and ultra-high energy neutrino telescopes such as ANITA and lunar Cherenkov experiments will clarify this.Comment: 11 pages, 6 figures, additional references and explanations. Accepted for publication in MNRA

Trace metal fluxes to the ocean: The importance of high‐standing oceanic islands

Peer Reviewedhttp://deepblue.lib.umich.edu/bitstream/2027.42/94592/1/grl16149.pd

Charge exchange emission and cold clumps in multi-phase galactic outflows

Large-scale outflows from starburst galaxies are multi-phase, multi-component fluids. Charge-exchange lines which originate from the interfacing surface between the neutral and ionised components are a useful diagnostic of the cold dense structures in the galactic outflow. From the charge-exchange lines observed in the nearby starburst galaxy M82, we conduct surface-to-volume analyses and deduce that the cold dense clumps in its galactic outflow have flattened shapes, resembling a hamburger or a pancake morphology rather than elongated shapes. The observed filamentary H$\alpha$ features are therefore not prime charge-exchange line emitters. They are stripped material torn from the slow moving dense clumps by the faster moving ionised fluid which are subsequently warmed and stretched into elongated shapes. Our findings are consistent with numerical simulations which have shown that cold dense clumps in galactic outflows can be compressed by ram pressure, and also progressively ablated and stripped before complete disintegration. We have shown that some clumps could survive their passage along a galactic outflow. These are advected into the circumgalactic environment, where their remnants would seed condensation of the circumgalactic medium to form new clumps. The infall of these new clumps back into the galaxy and their subsequent re-entrainment into the galactic outflow form a loop process of galactic material recycling.Comment: 17 pages, 6 figures; accepted for publication in MNRA

Morphological evolution of the mammalian jaw adductor complex

The evolution of the mammalian jaw during the transition from non-mammalian synapsids to crown mammals is a key event in vertebrate history and characterised by the gradual reduction of its individual bones into a single element and the concomitant transformation of the jaw joint and its incorporation into the middle ear complex. This osteological transformation is accompanied by a rearrangement and modification of the jaw adductor musculature, which is thought to have allowed the evolution of a more-efficient masticatory system in comparison to the plesiomorphic synapsid condition. While osteological characters relating to this transition are well documented in the fossil record, the exact arrangement and modifications of the individual adductor muscles during the cynodont–mammaliaform transition have been debated for nearly a century. We review the existing knowledge about the musculoskeletal evolution of the mammalian jaw adductor complex and evaluate previous hypotheses in the light of recently documented fossils that represent new specimens of existing species, which are of central importance to the mammalian origins debate. By employing computed tomography (CT) and digital reconstruction techniques to create three-dimensional models of the jaw adductor musculature in a number of representative non-mammalian cynodonts and mammaliaforms, we provide an updated perspective on mammalian jaw muscle evolution. As an emerging consensus, current evidence suggests that the mammal-like division of the jaw adductor musculature (into deep and superficial components of the m. masseter, the m. temporalis and the m. pterygoideus) was completed in Eucynodontia. The arrangement of the jaw adductor musculature in a mammalian fashion, with the m. pterygoideus group inserting on the dentary was completed in basal Mammaliaformes as suggested by the muscle reconstruction of Morganucodon oehleri. Consequently, transformation of the jaw adductor musculature from the ancestral (‘reptilian’) to the mammalian condition must have preceded the emergence of Mammalia and the full formation of the mammalian jaw joint. This suggests that the modification of the jaw adductor system played a pivotal role in the functional morphology and biomechanical stability of the jaw joint

Approximate imposition of boundary conditions in immersed boundary methods

We analyze several possibilities to prescribe boundary conditions in the context of immersed boundary methods. As basic approximation technique we consider the finite element method with a mesh that does not match the boundary of the computational domain, and therefore Dirichlet boundary conditions need to be prescribed in an approximate way. As starting variational approach we consider Nitsche&#39;s methods, and we then move to two options that yield non‐symmetric problems but that turned out to be robust and efficient. The essential idea is to use the degrees of freedom of certain nodes of the finite element mesh to minimize the difference between the exact and the approximated boundary conditio

Temporal Ordering in Endocytic Clathrin-Coated Vesicle Formation via AP2 Phosphorylation.

Clathrin-mediated endocytosis (CME) is key to maintaining the transmembrane protein composition of cells' limiting membranes. During mammalian CME, a reversible phosphorylation event occurs on Thr156 of the μ2 subunit of the main endocytic clathrin adaptor, AP2. We show that this phosphorylation event starts during clathrin-coated pit (CCP) initiation and increases throughout CCP lifetime. μ2Thr156 phosphorylation favors a new, cargo-bound conformation of AP2 and simultaneously creates a binding platform for the endocytic NECAP proteins but without significantly altering AP2's cargo affinity in vitro. We describe the structural bases of both. NECAP arrival at CCPs parallels that of clathrin and increases with μ2Thr156 phosphorylation. In turn, NECAP recruits drivers of late stages of CCP formation, including SNX9, via a site distinct from where NECAP binds AP2. Disruption of the different modules of this phosphorylation-based temporal regulatory system results in CCP maturation being delayed and/or stalled, hence impairing global rates of CME