Optical and ROSAT X-ray observations of the dwarf nova OY Carinae in superoutburst and quiescence

We present ROSAT X-ray and optical light curves of the 1994 February superoutburst of the eclipsing SU UMa dwarf nova OY Carinae. There is no eclipse of the flux in the ROSAT HRI light curve. Contemporaneous `wide B' band optical light curves show extensive superhump activity and dips at superhump maximum. Eclipse mapping of these optical light curves reveals a disc with a considerable physical flare, even three days into the superoutburst decline. We include a later (1994 July) ROSAT PSPC observation of OY Car that allows us to put constraints on the quiescent X-ray spectrum. We find that while there is little to choose between OY Car and its fellow high inclination systems with regard to the temperature of the emitting gas and the emission measure, we have difficulties reconciling the column density found from our X-ray observation with the column found in HST UV observations by Horne et al. (1994). The obvious option is to invoke time variability.Comment: 16 pages, 14 figures, accepted for publication in MNRA

IUE observations of the 1987 superoutburst of the dwarf nova Z Cha

Low resolution IUE observations of the dwarf nova Z Cha during superoutburst are presented. These cover most of the development of the outburst and have sufficient time resolution to probe continuum and line behavior on orbital phase. The observed modulation on this phase is very similar to that observed in the related object OY Car. The results imply the presence of a cool spot on the edge of the edge of the accretion disk, which periodically occults the brighter inner disk. Details of the line behavior suggest that the line originated in an extended wind-emitting region. In contrast to archive spectra obtained in normal outburst, the continuum is fainter and redder, indicating that the entire superoutburst disk may be geometrically thicker than during a normal outburst

The relationship between morphological and behavioral mimicry in hover flies (Diptera: Syrphidae).

Palatable (Batesian) mimics of unprofitable models could use behavioral mimicry to compensate for the ease with which they can be visually discriminated or to augment an already close morphological resemblance. We evaluated these contrasting predictions by assaying the behavior of 57 field-caught species of mimetic hover flies (Diptera: Syrphidae) and quantifying their morphological similarity to a range of potential hymenopteran models. A purpose-built phylogeny for the hover flies was used to control for potential lack of independence due to shared evolutionary history. Those hover fly species that engage in behavioral mimicry (mock stinging, leg waving, wing wagging) were all large wasp mimics within the genera Spilomyia and Temnostoma. While the behavioral mimics assayed were good morphological mimics, not all good mimics were behavioral mimics. Therefore, while the behaviors may have evolved to augment good morphological mimicry, they do not advantage all good mimics

Pathogens of Dikerogammarus haemobaphes regulate host activity and survival, but also threaten native amphipod populations in the UK

Dikerogammarus haemobaphes is a non-native amphipod in UK freshwaters. Studies have identified this species as a low-impact invader in the UK, relative to its cousin Dikerogammarus villosus. It has been suggested that regulation by symbionts (such as Microsporidia) could explain this difference in impact. The effect of parasitism on D. haemobaphes is largely unknown. This was explored herein using 2 behavioural assays measuring activity and aggregation. First, D. haemobaphes were screened histologically post-assay, identifying 2 novel viruses (D. haemobaphes bi-facies-like virus [DhbflV], D. haemobaphes bacilliform virus [DhBV]), Cucumispora ornata (Microsporidia), Apicomplexa, and Digenea, which could alter host behaviour. DhBV infection burden increased host activity, and C. ornata infection reduced host activity. Second, native invertebrates were collected from the invasion site at Carlton Brook, UK, and tested for the presence of C. ornata. PCR screening identified that Gammarus pulex and other native invertebrates were positive for C. ornata. The host range of this parasite, and its impact on host survival, was additionally explored using D. haemobaphes, D. villosus, and G. pulex in a laboratory trial. D. haemobaphes and G. pulex became infected by C. ornata, which also lowered survival rate. D. villosus did not become infected. A PCR protocol for DhbflV was also applied to D. haemobaphes after the survival trial, associating this virus with decreased host survival. In conclusion, D. haemobaphes has a complex relationship with parasites in the UK environment. C. ornata likely regulates populations by decreasing host survival and activity, but despite this benefit, the parasite threatens susceptible native wildlife

Erratum: The helium abundance in the ejecta of U Scorpii

U Scorpii (U Sco) is a recurrent nova which has been observed in outburst on 10 occasions, most recently in 2010. We present near-infrared (near-IR) and optical spectroscopy of the 2010 outburst of U Sco. The reddening of U Sco is found to be E(B − V) = 0.14 ± 0.12, consistent with previous determinations, from simultaneous optical and near-IR observations. The spectra show the evolution of the linewidths and profiles to be consistent with previous outbursts. Velocities are found to be up to 14 000 km s−1 in broad components and up to 1800 km s−1 in narrow-line components, which become visible around day 8 due to changes in the optical depth. From the spectra we derive a helium abundance of N(He)/N(H) = 0.073 ± 0.031 from the most reliable lines available; this is lower than most other estimates and indicates that the secondary is not helium-rich, as previous studies have suggested

Temporal dynamics of aquatic communities and implications for pond conservation

Conservation through the protection of particular habitats is predicated on the assumption that the conservation value of those habitats is stable. We test this assumption for ponds by investigating temporal variation in macroinvertebrate and macrophyte communities over a 10-year period in northwest England. We surveyed 51 ponds in northern England in 1995/6 and again in 2006, identifying all macrophytes (167 species) and all macroinvertebrates (221 species, excluding Diptera) to species. The alpha-diversity, beta-diversity and conservation value of these ponds were compared between surveys. We find that invertebrate species richness increased from an average of 29. 5 species to 39. 8 species between surveys. Invertebrate gamma-diversity also increased between the two surveys from 181 species to 201 species. However, this increase in diversity was accompanied by a decrease in beta-diversity. Plant alpha-, beta and gamma-diversity remained approximately constant between the two periods. However, increased proportions of grass species and a complete loss of charophytes suggests that the communities are undergoing succession. Conservation value was not correlated between sampling periods in either plants or invertebrates. This was confirmed by comparing ponds that had been disturbed with those that had no history of disturbance to demonstrate that levels of correlation between surveys were approximately equal in each group of ponds. This study has three important conservation implications: (i) a pond with high diversity or high conservation value may not remain that way and so it is unwise to base pond conservation measures upon protecting currently-speciose habitats; (ii) maximising pond gamma-diversity requires a combination of late and early succession ponds, especially for invertebrates; and (iii) invertebrate and plant communities in ponds may require different management strategies if succession occurs at varying rates in the two groups

Chandra Observations of the Dwarf Nova WX Hyi in Quiescence

We report Chandra observations of the dwarf nova WX Hyi in quiescence. The X-ray spectrum displays strong and narrow emission lines of N, O, Mg, Ne, Si, S and Fe. The various ionization states implied by the lines suggest that the emission is produced within a flow spanning a wide temperature range, from T ~ 10^6 K to T >~ 10^8 K. Line diagnostics indicate that most of the radiation originates from a very dense region, with n ~ 10^{13}-10^{14} cm^{-3}. The Chandra data allow the first tests of specific models proposed in the literature for the X-ray emission in quiescent dwarf novae. We have computed the spectra for a set of models ranging from hot boundary layers, to hot settling flows solutions, to X-ray emitting coronae. WX Hyi differs from other dwarf novae observed at minimum in having much stronger low temperature lines, which prove difficult to fit with existing models, and possibly a very strong, broad O VII line, perhaps produced in a wind moving at a few x 10^3 km/s. The accretion rate inferred from the X-rays is lower than the value inferred from the UV. The presence of high-velocity mass ejection could account for this discrepancy while at the same time explaining the presence of the broad O VII line. If this interpretation is correct, it would provide the first detection of a wind from a dwarf nova in quiescence.Comment: accepted to ApJ; 19 pages, 3 figures, 1 tabl

An XMM-Newton observation of the nova-like variable UX UMa: spatially and spectrally resolved two-component X-ray emission

In the optical and ultraviolet regions of the electromagnetic spectrum, UX Ursae Majoris is a deeply eclipsing cataclysmic variable. However, no soft X-ray eclipse was detected in ROSAT observations. We have obtained a 38 ksec XMM-Newton observation to further constrain the origin of the X-rays. The combination of spectral and timing information allows us to identify two components in the X-ray emission of the system. The soft component, dominant below photon energies of 2 keV, can be fitted with a multi-temperature plasma model and is uneclipsed. The hard component, dominant above 3 keV, can be fitted with a kT ~ 5 keV plasma model and appears to be deeply eclipsed. We suggest that the most likely source of the hard X-ray emission in UX UMa, and other systems in high mass transfer states, is the boundary layer.Comment: To appear in MNRAS Letter

Genetic diversity in nitrogen fertilizer responses and N gas emission in modern wheat

Crops assimilate nitrogen (N) as ammonium via the glutamine synthetase/glutamate synthase (GS/GOGAT) pathway which is of central importance for N uptake and potentially represents a bottle neck for N fertilizer-use efficiency. The aim of this study was to assess whether genetic diversity for N-assimilation capacity exists in wheat and could be exploited for breeding. Wheat plants rapidly, within 6h, responded to N application with an increase in GS activity. This was not accompanied by an increase in GS gene transcript abundance and a comparison of GS1 and GS2 protein models revealed a high degree of sequence conservation. N responsiveness amongst ten wheat varieties was assessed by measuring GS enzyme activity, leaf tissue ammonium, and by a leaf-disc assay as a proxy for apoplastic ammonia. Based on these data, a high-GS group showing an overall positive response to N could be distinguished from an inefficient, low-GS group. Subsequent gas emission measurements confirmed plant ammonia emission in response to N application and also revealed emission of N2O when N was provided as nitrate, which is in agreement with our current understanding that N2O is a by-product of nitrate reduction. Taken together, the data suggest that there is scope for improving N assimilation capacity in wheat and that further investigations into the regulation and role of GS-GOGAT in NH3 emission is justified. Likewise, emission of the climate gas N2O needs to be reduced, and future research should focus on assessing the nitrate reductase pathway in wheat and explore fertilizer management options

Hyperons analogous to the \Lambda(1405)

The low mass of the $\Lambda(1405)$ hyperon with $j^P = 1/2^-$, which is higher than the ground state $\Lambda(1116)$ mass by 290 MeV, is difficult to understand in quark models. We analyze the hyperon spectrum in the bound state approach of the Skyrme model that successfully describes both the $\Lambda(1116)$ and the $\Lambda(1405)$. This model predicts that several hyperon resonances of the same spin but with opposite parity form parity doublets that have a mass difference of around 300 MeV, which is indeed realized in the observed hyperon spectrum. Furthermore, the existence of the $\Xi(1620)$ and the $\Xi(1690)$ of $j^P=1/2^-$ is predicted by this model. Comments on the $\Omega$ baryons and heavy quark baryons are made as well.Comment: 4 pages, talk presented at the Fifth Asia-Pacific Conference on Few-Body Problems in Physics 2011 (APFB2011), Aug. 22-26, 2011, Seoul, Kore