Composition of Early Planetary Atmospheres II: Coupled Dust and Chemical Evolution in Protoplanetary Disks

We present the next step in a series of papers devoted to connecting the composition of the atmospheres of forming planets with the chemistry of their natal evolving protoplanetary disks. The model presented here computes the coupled chemical and dust evolution of the disk and the formation of three planets per disk model. Our three canonical planet traps produce a Jupiter near 1 AU, a Hot Jupiter and a Super-Earth. We study the dependency of the final orbital radius, mass, and atmospheric chemistry of planets forming in disk models with initial disk masses that vary by 0.02 $M_\odot$ above and below our fiducial model ($M_{disk,0} = 0.1 ~M_\odot$). We compute C/O and C/N for the atmospheres formed in our 3 models and find that C/O$_{\rm planet}\sim$ C/O$_{\rm disk}$, which does not vary strongly between different planets formed in our model. The nitrogen content of atmospheres can vary in planets that grow in different disk models. These differences are related to the formation history of the planet, the time and location that the planet accretes its atmosphere, and are encoded in the bulk abundance of NH$_3$. These results suggest that future observations of atmospheric NH$_3$ and an estimation of the planetary C/O and C/N can inform the formation history of particular planetary systems.Comment: Accepted for publication in MNRA

Testing particle trapping in transition disks with ALMA

We present new Atacama Large Millimeter/submillimeter Array (ALMA) continuum observations at 336GHz of two transition disks, SR21 and HD135344B. In combination with previous ALMA observations from Cycle 0 at 689GHz, we compare the visibility profiles at the two frequencies and calculate the spectral index ($\alpha_{\rm{mm}}$). The observations of SR21 show a clear shift in the visibility nulls, indicating radial variations of the inner edge of the cavity at the two wavelengths. Notable radial variations of the spectral index are also detected for SR21 with values of $\alpha_{\rm{mm}}{\sim}3.8-4.2$ in the inner region ($r<35$ AU) and $\alpha_{\rm{mm}}{\sim}2.6-3.0$ outside. An axisymmetric ring (which we call the ring model) or a ring with the addition of an azimuthal Gaussian profile, for mimicking a vortex structure (which we call the vortex model), is assumed for fitting the disk morphology. For SR21, the ring model better fits the emission at 336GHz, conversely the vortex model better fits the 689GHz emission. For HD135344B, neither a significant shift in the null of the visibilities nor radial variations of $\alpha_{\rm{mm}}$ are detected. Furthermore, for HD135344B, the vortex model fits both frequencies better than the ring model. However, the azimuthal extent of the vortex increases with wavelength, contrary to model predictions for particle trapping by anticyclonic vortices. For both disks, the azimuthal variations of $\alpha_{\rm{mm}}$ remain uncertain to confirm azimuthal trapping. The comparison of the current data with a generic model of dust evolution that includes planet-disk interaction suggests that particles in the outer disk of SR21 have grown to millimetre sizes and have accumulated in a radial pressure bump, whereas with the current resolution there is not clear evidence of radial trapping in HD135344B, although it cannot be excluded either.Comment: Minor changes after language edition. Accepted for publication in A&A (abstract slightly shortened for arXiv

A Multi-Wavelength Analysis of Dust and Gas in the SR 24S Transition Disk

We present new Atacama Large Millimeter/sub-millimeter Array (ALMA) 1.3 mm continuum observations of the SR 24S transition disk with an angular resolution $\lesssim0.18"$ (12 au radius). We perform a multi-wavelength investigation by combining new data with previous ALMA data at 0.45 mm. The visibilities and images of the continuum emission at the two wavelengths are well characterized by a ring-like emission. Visibility modeling finds that the ring-like emission is narrower at longer wavelengths, in good agreement with models of dust trapping in pressure bumps, although there are complex residuals that suggest potentially asymmetric structures. The 0.45 mm emission has a shallower profile inside the central cavity than the 1.3 mm emission. In addition, we find that the $^{13}$CO and C$^{18}$O (J=2-1) emission peaks at the center of the continuum cavity. We do not detect either continuum or gas emission from the northern companion to this system (SR 24N), which is itself a binary system. The upper limit for the dust disk mass of SR 24N is $\lesssim 0.12\,M_{\bigoplus}$, which gives a disk mass ratio in dust between the two components of $M_{\mathrm{dust, SR\,24S}}/M_{\mathrm{dust, SR\,24N}}\gtrsim840$. The current ALMA observations may imply that either planets have already formed in the SR 24N disk or that dust growth to mm-sizes is inhibited there and that only warm gas, as seen by ro-vibrational CO emission inside the truncation radii of the binary, is present.Comment: Accepted for publication in Ap

Dust Settling and Rapid Planetary Migration

Planetary migration is essential to explain the observed mass-period relation for exoplanets. Without some stopping mechanism, the tidal, resonant interaction between planets and their gaseous disc generally causes the planets to migrate inward so efficiently that they plunge into the host star within the gaseous disc lifetime ($\sim$ 1-3 Myrs). We investigate planetary migration by analytically calculating the migration rate and time within self-consistently computed, radiatively heated discs around M stars in which the effects of dust settling are included. We show that dust settling lowers the disc temperature and raises the gas density in the mid-plane. This inescapable evolution of disc structure speeds up type I planetary migration for lower mass bodies by up to a factor of about 2. We also examine the effects of dust settling on the gap-opening mass and type II migration, and find that the gap-opening mass is reduced by a factor of 2 and type II migration becomes slower by a factor of 2. While dust settling can somewhat alleviate the problem of planetary migration for more massive planets, the more rapid migration of low mass planets and planetary cores requires a robust slowing mechanism.Comment: 16 pages, 13 figures, 3 tables, accepted for publication in MNRA

Coupling of adenovirus to transferrin-polylysine/DNA complexes greatly enhances receptor-mediated gene delivery and expression of transfected genes.

We are developing efficient methods for gene transfer into tissue culture cells. We have previously shown that coupling of a chimeric adenovirus with polylysine allowed the construction of an adenovirus-polylysine-reporter-gene complex that transferred the transporter gene with great efficiency into HeLa cells. We have now explored simpler, biochemical means for coupling adenovirus to DNA/polylysine complexes and show that such complexes yield virtually 100% transfection in tissue culture cell lines. In these methods adenovirus is coupled to polylysine, either enzymatically through the action of transglutaminase or biochemically by biotinylating adenovirus and streptavidinylating the polylysine moiety. Combination complexes containing DNA, adenovirus-polylysine, and transferrin-polylysine have the capacity to transfer the reporter gene into adenovirus-receptor- and/or transferrin-receptor-rich cells

Debris disk size distributions: steady state collisional evolution with P-R drag and other loss processes

We present a new scheme for determining the shape of the size distribution, and its evolution, for collisional cascades of planetesimals undergoing destructive collisions and loss processes like Poynting-Robertson drag. The scheme treats the steady state portion of the cascade by equating mass loss and gain in each size bin; the smallest particles are expected to reach steady state on their collision timescale, while larger particles retain their primordial distribution. For collision-dominated disks, steady state means that mass loss rates in logarithmic size bins are independent of size. This prescription reproduces the expected two phase size distribution, with ripples above the blow-out size, and above the transition to gravity-dominated planetesimal strength. The scheme also reproduces the expected evolution of disk mass, and of dust mass, but is computationally much faster than evolving distributions forward in time. For low-mass disks, P-R drag causes a turnover at small sizes to a size distribution that is set by the redistribution function (the mass distribution of fragments produced in collisions). Thus information about the redistribution function may be recovered by measuring the size distribution of particles undergoing loss by P-R drag, such as that traced by particles accreted onto Earth. Although cross-sectional area drops with 1/age^2 in the PR-dominated regime, dust mass falls as 1/age^2.8, underlining the importance of understanding which particle sizes contribute to an observation when considering how disk detectability evolves. Other loss processes are readily incorporated; we also discuss generalised power law loss rates, dynamical depletion, realistic radiation forces and stellar wind drag.Comment: Accepted for publication by Celestial Mechanics and Dynamical Astronomy (special issue on EXOPLANETS

Planetary population synthesis

In stellar astrophysics, the technique of population synthesis has been successfully used for several decades. For planets, it is in contrast still a young method which only became important in recent years because of the rapid increase of the number of known extrasolar planets, and the associated growth of statistical observational constraints. With planetary population synthesis, the theory of planet formation and evolution can be put to the test against these constraints. In this review of planetary population synthesis, we first briefly list key observational constraints. Then, the work flow in the method and its two main components are presented, namely global end-to-end models that predict planetary system properties directly from protoplanetary disk properties and probability distributions for these initial conditions. An overview of various population synthesis models in the literature is given. The sub-models for the physical processes considered in global models are described: the evolution of the protoplanetary disk, the planets' accretion of solids and gas, orbital migration, and N-body interactions among concurrently growing protoplanets. Next, typical population synthesis results are illustrated in the form of new syntheses obtained with the latest generation of the Bern model. Planetary formation tracks, the distribution of planets in the mass-distance and radius-distance plane, the planetary mass function, and the distributions of planetary radii, semimajor axes, and luminosities are shown, linked to underlying physical processes, and compared with their observational counterparts. We finish by highlighting the most important predictions made by population synthesis models and discuss the lessons learned from these predictions - both those later observationally confirmed and those rejected.Comment: 47 pages, 12 figures. Invited review accepted for publication in the 'Handbook of Exoplanets', planet formation section, section editor: Ralph Pudritz, Springer reference works, Juan Antonio Belmonte and Hans Deeg, Ed

Circumstellar disks and planets. Science cases for next-generation optical/infrared long-baseline interferometers

We present a review of the interplay between the evolution of circumstellar disks and the formation of planets, both from the perspective of theoretical models and dedicated observations. Based on this, we identify and discuss fundamental questions concerning the formation and evolution of circumstellar disks and planets which can be addressed in the near future with optical and infrared long-baseline interferometers. Furthermore, the importance of complementary observations with long-baseline (sub)millimeter interferometers and high-sensitivity infrared observatories is outlined.Comment: 83 pages; Accepted for publication in "Astronomy and Astrophysics Review"; The final publication is available at http://www.springerlink.co

Insect small nuclear RNA gene promoters evolve rapidly yet retain conserved features involved in determining promoter activity and RNA polymerase specificity

In animals, most small nuclear RNAs (snRNAs) are synthesized by RNA polymerase II (Pol II), but U6 snRNA is synthesized by RNA polymerase III (Pol III). In Drosophila melanogaster, the promoters for the Pol II-transcribed snRNA genes consist of ∼21 bp PSEA and ∼8 bp PSEB. U6 genes utilize a PSEA but have a TATA box instead of the PSEB. The PSEAs of the two classes of genes bind the same protein complex, DmSNAPc. However, the PSEAs that recruit Pol II and Pol III differ in sequence at a few nucleotide positions that play an important role in determining RNA polymerase specificity. We have now performed a bioinformatic analysis to examine the conservation and divergence of the snRNA gene promoter elements in other species of insects. The 5′ half of the PSEA is well-conserved, but the 3′ half is divergent. Moreover, within each species positions exist where the PSEAs of the Pol III-transcribed genes differ from those of the Pol II-transcribed genes. Interestingly, the specific positions vary among species. Nevertheless, we speculate that these nucleotide differences within the 3′ half of the PSEA act similarly to induce conformational alterations in DNA-bound SNAPc that result in RNA polymerase specificity