PENINGKATAN KETERAMPILAN MENULIS TEKS EKSPOSISI MELALUI PENGGUNAAN STRATEGI PEMBELAJARAN THINK TALK WRITE DAN MEDIA AUDIO VISUAL ( Penelitian Tindakan Kelas pada Siswa Kelas X IPS 2 SMA N 1 Surakarta Tahun Pelajaran 2017/2018)

Yustina Dwinuryati.2017. Peningkatan Keterampilan menulis Teks Eksposisi melalui Penggunaan Strategi Pembelajaran Think, Talk, Write dan Media Audio Visual (Penelitian Tindakan Kelas pada Siswa Kelas X IPS 2 SMA N 1 Surakarta Tahun Pelajaran 2017/2018). Tesis. Pembimbing: Prof. Dr. Andayani, M.Pd. Kopembimbing: Prof. Dr. Retno Winarni, M.Pd. Program Studi Magister Pendidikan Bahasa Indonesia, Fakultas Keguruan dan Ilmu Pendidikan, Universitas Sebelas Maret Surakarta. ABSTRAK Keterampilan menulis merupakan keterampilan berbahasa tingkat tinggi dan harus diajarkan kepada siswa di Sekolah Menengah Atas. Salah satu keterampilan menulis yang harus diajarkan kepada siswa kelas X sesuai Kurikulum 2013 Revisi adalah menulis teks eksposisi. Pembelajaran keterampilan menulis teks eksposisi di kelas X IPS2 SMA N 1 Surakarta mengalami permasalahan baik dari sisi motivasi belajar maupun keterampilan menulis siswa. Penelitian ini bertujuan untuk meningkatkan: (1) motivasi belajar menulis teks eksposisi siswa kelas X IPS 2 SMA Negeri 1 Surakarta dengan strategi pembelajaran think talk write dan penggunaan media audio visual dan (2) keterampilan menulis teks eksposisi siswa kelas X IPS 2 SMA Negeri 1 Surakarta dengan strategi pembelajaran think talk write dan media audio visual. Strategi penelitian berupa Penelitian Tindakan Kelas. Data penelitian bersumber dari proses pembelajaran, informan, hasil tes menulis teks eksposisi, dan dokumen. Teknik pengumpulan data dengan pengamatan, kajian dokumen, wawancara, dan tes. Uji validitas data menggunakan teknik triangulasi sumber data dan triangulasi metode. Teknik analisis data menggunakan teknik deskriptif komparatif dan analisis kritis. Hasil penelitian menunjukkan bahwa penerapan strategi pembelajaran think, talk, write dan penggunaan media audio visual pada siswa kelas X IPS2 SMA N I Surakarta dapat meningkatkan motivasi belajar dan keterampilan menulis teks eksposisi dari siklus 1 ke siklus 2. Hal itu dibuktikan adanya perubahan dan peningkatan motivasi belajar dan keterampilan menulis teks eksposisi siswa: (1) motivasi siswa meningkat dari siklus 1 sebesar 68% menjadi 82% pada siklus 2 dan (2) keterampilan menulis teks eksposisi meningkat dari siklus 1 sebesar 76% meningkat menjadi 88% pada siklus 2. Kata kunci: teks eksposisi, motivasi, strategi think talk write, audio visua

Gravity modes as a way to distinguish between hydrogen- and helium-burning red giant stars

Red giants are evolved stars that have exhausted the supply of hydrogen in their cores and instead burn hydrogen in a surrounding shell. Once a red giant is sufficiently evolved, the helium in the core also undergoes fusion. Outstanding issues in our understanding of red giants include uncertainties in the amount of mass lost at the surface before helium ignition and the amount of internal mixing from rotation and other processes. Progress is hampered by our inability to distinguish between red giants burning helium in the core and those still only burning hydrogen in a shell. Asteroseismology offers a way forward, being a powerful tool for probing the internal structures of stars using their natural oscillation frequencies. Here we report observations of gravity-mode period spacings in red giants that permit a distinction between evolutionary stages to be made. We use high-precision photometry obtained with the Kepler spacecraft over more than a year to measure oscillations in several hundred red giants. We find many stars whose dipole modes show sequences with approximately regular period spacings. These stars fall into two clear groups, allowing us to distinguish unambiguously between hydrogen-shell-burning stars (period spacing mostly about 50 seconds) and those that are also burning helium (period spacing about 100 to 300 seconds).Comment: to appear as a Letter to Natur

The case for the continued use of the genus name Mimulus for all monkeyflowers

The genus Mimulus is a well-studied group of plant species, which has for decades allowed researchers to address a wide array of fundamental questions in biology (Wu & al. 2008; Twyford & al. 2015). Linnaeus named the type species of Mimulus (ringens L.), while Darwin (1876) used Mimulus (luteus L.) to answer key research questions. The incredible phenotypic diversity of this group has made it the focus of ecological and evolutionary study since the mid-20th century, initiated by the influential work of Clausen, Keck, and Hiesey as well as their students and collaborators (Clausen & Hiesey 1958; Hiesey & al. 1971, Vickery 1952, 1978). Research has continued on this group of diverse taxa throughout the 20th and into the 21st century (Bradshaw & al. 1995; Schemske & Bradshaw 1999; Wu & al. 2008; Twyford & al. 2015; Yuan 2019), and Mimulus guttatus was one of the first non-model plants to be selected for full genome sequencing (Hellsten & al. 2013). Mimulus has played a key role in advancing our general understanding of the evolution of pollinator shifts (Bradshaw & Schemske 2003; Cooley & al. 2011; Byers & al. 2014), adaptation (Lowry & Willis 2010; Kooyers & al. 2015; Peterson & al. 2016; Ferris & Willis 2018; Troth & al. 2018), speciation (Ramsey & al. 2003; Wright & al. 2013; Sobel & Streisfeld 2015; Zuellig & Sweigart 2018), meiotic drive (Fishman & Saunders 2008), polyploidy (Vallejo-Marín 2012; Vallejo-Marín & al. 2015), range limits (Angert 2009; Sexton et al. 2011; Grossenbacher & al. 2014; Sheth & Angert 2014), circadian rhythms (Greenham & al. 2017), genetic recombination (Hellsten & al. 2013), mating systems (Fenster & Ritland 1994; Dudash & Carr 1998; Brandvain & al. 2014) and developmental biology (Moody & al. 1999; Baker & al. 2011, 2012; Yuan 2019). This combination of a rich history of study coupled with sustained modern research activity is unparalleled among angiosperms. Across many interested parties, the name Mimulus therefore takes on tremendous biological significance and is recognizable not only by botanists, but also by zoologists, horticulturalists, naturalists, and members of the biomedical community. Names associated with a taxonomic group of this prominence should have substantial inertia, and disruptive name changes should be avoided. As members of the Mimulus community, we advocate retaining the genus name Mimulus to describe all monkeyflowers. This is despite recent nomenclature changes that have led to a renaming of most monkeyflower species to other genera.Additional co-authors: Jannice Friedman, Dena L Grossenbacher, Liza M Holeski, Christopher T Ivey, Kathleen M Kay, Vanessa A Koelling, Nicholas J Kooyers, Courtney J Murren, Christopher D Muir, Thomas C Nelson, Megan L Peterson, Joshua R Puzey, Michael C Rotter, Jeffrey R Seemann, Jason P Sexton, Seema N Sheth, Matthew A Streisfeld, Andrea L Sweigart, Alex D Twyford, John H Willis, Kevin M Wright, Carrie A Wu, Yao-Wu Yua

Simple Y-Autosomal Incompatibilities Cause Hybrid Male Sterility in Reciprocal Crosses Between Drosophila virilis and D. americana

Postzygotic reproductive isolation evolves when hybrid incompatibilities accumulate between diverging populations. Here, I examine the genetic basis of hybrid male sterility between two species of Drosophila, Drosophila virilis and D. americana. From these analyses, I reach several conclusions. First, neither species carries any autosomal dominant hybrid male sterility alleles: reciprocal F1 hybrid males are perfectly fertile. Second, later generation (backcross and F2) hybrid male sterility between D. virilis and D. americana is not polygenic. In fact, I identified only three genetically independent incompatibilities that cause hybrid male sterility. Remarkably, each of these incompatibilities involves the Y chromosome. In one direction of the cross, the D. americana Y is incompatible with recessive D. virilis alleles at loci on chromosomes 2 and 5. In the other direction, the D. virilis Y chromosome causes hybrid male sterility in combination with recessive D. americana alleles at a single QTL on chromosome 5. Finally, in contrast with findings from other Drosophila species pairs, the X chromosome has only a modest effect on hybrid male sterility between D. virilis and D. americana

Mechanisms of Transmission Ratio Distortion at Hybrid Sterility Loci Within and Between Mimulus Species

Hybrid incompatibilities are a common correlate of genomic divergence and a potentially important contributor to reproductive isolation. However, we do not yet have a detailed understanding of how hybrid incompatibility loci function and evolve within their native species, or why they are dysfunctional in hybrids. Here, we explore these issues for a well-studied, two-locus hybrid incompatibility between hybrid male sterility 1 (hms1) and hybrid male sterility 2 (hms2) in the closely related yellow monkeyflower species Mimulus guttatus and M. nasutus. By performing reciprocal backcrosses with introgression lines (ILs), we find evidence for gametic expression of the hms1-hms2 incompatibility. Surprisingly, however, hybrid transmission ratios at hms1 do not reflect this incompatibility, suggesting that additional mechanisms counteract the effects of gametic sterility. Indeed, our backcross experiment shows hybrid transmission bias toward M. guttatus through both pollen and ovules, an effect that is particularly strong when hms2 is homozygous for M. nasutus alleles. In contrast, we find little evidence for hms1 transmission bias in crosses within M. guttatus, providing no indication of selfish evolution at this locus. Although we do not yet have sufficient genetic resolution to determine if hybrid sterility and transmission ratio distortion (TRD) map to the same loci, our preliminary fine-mapping uncovers a genetically independent hybrid lethality system involving at least two loci linked to hms1. This fine-scale dissection of TRD at hms1 and hms2 provides insight into genomic differentiation between closely related Mimulus species and reveals multiple mechanisms of hybrid dysfunction

A Simple Genetic Incompatibility Causes Hybrid Male Sterility in Mimulus

Much evidence has shown that postzygotic reproductive isolation (hybrid inviability or sterility) evolves by the accumulation of interlocus incompatibilities between diverging populations. Although in theory only a single pair of incompatible loci is needed to isolate species, empirical work in Drosophila has revealed that hybrid fertility problems often are highly polygenic and complex. In this article we investigate the genetic basis of hybrid sterility between two closely related species of monkeyflower, Mimulus guttatus and M. nasutus. In striking contrast to Drosophila systems, we demonstrate that nearly complete hybrid male sterility in Mimulus results from a simple genetic incompatibility between a single pair of heterospecific loci. We have genetically mapped this sterility effect: the M. guttatus allele at the hybrid male sterility 1 (hms1) locus acts dominantly in combination with recessive M. nasutus alleles at the hybrid male sterility 2 (hms2) locus to cause nearly complete hybrid male sterility. In a preliminary screen to find additional small-effect male sterility factors, we identified one additional locus that also contributes to some of the variation in hybrid male fertility. Interestingly, hms1 and hms2 also cause a significant reduction in hybrid female fertility, suggesting that sex-specific hybrid defects might share a common genetic basis. This possibility is supported by our discovery that recombination is reduced dramatically in a cross involving a parent with the hms1–hms2 incompatibility

CAC.Mimulus.2013.photoperiod.experiment

This data file contains flowering data for Mimulus guttatus and Mimulus nasutus from a growthchamber, photoperiod experiment performed in 2013. Individuals from five maternal M. guttatus lines (N=11-12 per line, per treatment) and two M. nasutus lines (N=23 per line, per treatment) were grown under two daylength treatments: 12 or 16 hrs. Whether an individual flowered was recorded as yes or no after eight weeks of growth. Families were originally collected from Catherine Creek in Washington State (USA) in 2010 or 2012. Each row in the data sheet is one individual. See additional information and descriptions of their location of origin in Kenney and Sweigart 2016

Gene duplicates cause hybrid lethality between sympatric species of <i>Mimulus</i>

<div><p>Hybrid incompatibilities play a critical role in the evolution and maintenance of species. We have discovered a simple genetic incompatibility that causes lethality in hybrids between two closely related species of yellow monkeyflower (<i>Mimulus guttatus</i> and <i>M</i>. <i>nasutus</i>). This hybrid incompatibility, which causes one sixteenth of F₂ hybrid seedlings to lack chlorophyll and die shortly after germination, occurs between sympatric populations that are connected by ongoing interspecific gene flow. Using complimentary genetic mapping and gene expression analyses, we show that lethality occurs in hybrids that lack a functional copy of the critical photosynthetic gene <i>pTAC14</i>. In <i>M</i>. <i>guttatus</i>, this gene was duplicated, but the ancestral copy is no longer expressed. In <i>M</i>. <i>nasutus</i>, the duplication is missing altogether. As a result, hybrids die when they are homozygous for the nonfunctional <i>M</i>. <i>guttatus</i> copy and missing the duplicate from <i>M</i>. <i>nasutus</i>, apparently due to misregulated transcription of key photosynthetic genes. Our study indicates that neutral evolutionary processes may play an important role in the evolution of hybrid incompatibilities and opens the door to direct investigations of their contribution to reproductive isolation among naturally hybridizing species.</p></div