Redescription and biology of Limonia badia (Walker) (Diptera: Limoniidae)

Limonia badia (Walker, 1848), previously known from the Nearctic Region only, is redescribed from Finland. Its systematic position and biology are currently debated. Limonia indigenoides (Alexander, 1920), another Nearctic species, is synonymized with L. badia. The species is distinguished by several distinctive characters, such as the wing pattern and the structure of the male terminalia. These characters are illustrated for the first time. Immature stages are probably associated with decayingwood of aspen (Populus tremula L.). A list of species of the Tipuloidea associated with L. badia in Finland – including Gnophomyia acheron Alexander, 1950, a species new to Europe – is presented

First records of some species of Diptera (Insecta) from the Azores

During a collecting trip undertaken by J. Roháček and M. Vála in the São Miguel Island (Azores) in August and September 2006, mainly devoted to acalyptrate flies, three distinctive species of Diptera were found, two of which proved to be hitherto unrecorded from the Azorean archipelago. These additions to the regional fauna are presented below with a discussion of their origin. The voucher specimens of the species recorded below are deposited in the following collections: JSO – collection of J. Starý, Olomouc, Czech Republic, SMOC – Silesian Museum, Opava, Czech Republic, ZMAN – Zoological Museum, Amsterdam, Netherlands

A laboratory for conceiving Essential Biodiversity Variables (EBVs)—The ‘Data pool initiative for the Bohemian Forest Ecosystem’

Effects of climate change-induced events on forest ecosystem dynamics of composition, function and structure call for increased long-term, interdisciplinary and integrated research on biodiversity indicators, in particular within strictly protected areas with extensive non-intervention zones. The long-established concept of forest supersites generally relies on long-term funds from national agencies and goes beyond the logistic and financial capabilities of state-or region-wide protected area administrations, universities and research institutes. We introduce the concept of data pools as a smaller-scale, user-driven and reasonable alternative to co-develop remote sensing and forest ecosystem science to validated products, biodiversity indicators and management plans. We demonstrate this concept with the Bohemian Forest Ecosystem Data Pool, which has been established as an interdisciplinary, international data pool within the strictly protected Bavarian Forest and Šumava National Parks and currently comprises 10 active partners. We demonstrate how the structure and impact of the data pool differs from comparable cases. We assessed the international influence and visibility of the data pool with the help of a systematic literature search and a brief analysis of the results. Results primarily suggest an increase in the impact and visibility of published material during the life span of the data pool, with highest visibilities achieved by research conducted on leaf traits, vegetation phenology and 3D-based forest inventory. We conclude that the data pool results in an efficient contribution to the concept of global biodiversity observatory by evolving towards a training platform, functioning as a pool of data and algorithms, directly communicating with management for implementation and providing test fields for feasibility studies on earth observation missions.publishedVersio

The state of research into children with cancer across Europe : new policies for a new decade

Overcoming childhood cancers is critically dependent on the state of research. Understanding how, with whom and what the research community is doing with childhood cancers is essential for ensuring the evidence-based policies at national and European level to support children, their families and researchers. As part of the European Union funded EUROCANCERCOMS project to study and integrate cancer communications across Europe, we have carried out new research into the state of research in childhood cancers. We are very grateful for all the support we have received from colleagues in the European paediatric oncology community, and in particular from Edel Fitzgerald and Samira Essiaf from the SIOP Europe office. This report and the evidence-based policies that arise from it come at a important junction for Europe and its Member States. They provide a timely reminder that research into childhood cancers is critical and needs sustainable long-term support.peer-reviewe

Molophilus (Molophilus) maurus Lackschewitz 1925

Molophilus (Molophilus) maurus Lackschewitz, 1925 Material examined. Austria: Steiermark: Admont env., Hall-Grieshof, boggy meadows, 15.vi. 2006, 10 3 (J. Starý leg.) (JSO). Distribution. Czech Republic, Latvia, Lithuania, Poland, Slovakia, Switzerland; Russia: European part. First record for Austria.Published as part of Starý, Jaroslav, 2011, Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae), pp. 45-62 in Zootaxa 2999 on page 60, DOI: 10.5281/zenodo.20328

Limonia opacipennis Starý 2017, sp. nov.

Limonia opacipennis sp. nov. (Figs 3–4) Type material. HOLOTYPE: ♂ (RMNH), ALGERIA: Petite Kabylie Mts, 32 km S El Aouana, 1300 m, 25.v.1986 (P. Oosterbroek leg.), labelled ‘ALGÉRIE / PETITE KABYLIE / P.Oosterbroek’ // ‘ 32 km S EL AOUANA / 1300 m / 25.V.1986 ’ [both printed, white labels] // ‘HOLOTYPE / Limonia / opacipennis sp. n. ♂/ J. Starý 2017’ [printed, red label]. The specimen is pinned, with left fore leg missing; apex of abdomen cut off, terminalia dissected and placed in a sealed plastic tube with glycerine, pinned with the specimen. PARATYPES: 2 ♂♂ 2 ♀♀, same data as for holotype (RMNH, JSOC). TUNISIA: Oued ed Demene, 7 km S of Aïn Draham, 600 m, along brook, 24.iv.1980, 1 ♀ (E. v.Nieukerken, G. Bryan & P. Oosterbroek leg.); Hotel les Chenes, 5 km S of Aïn Draham, 750 m, at light, Quercus faginea & suber veg., 23.-25.iv.1980, 1 ♀ (collector(s) not given, but most probably the same as for the preceding specimen) (RMNH). All specimens pinned or micro-pinned on a stage of polyporus; terminalia, if dissected, placed as for holotype. Diagnosis. Medium-sized species. Body colouration in general brown, restrictedly dark brown on dorsum of thorax and obscure yellow on pleuron. Wing membrane tinged brownish. Wing pattern diffuse, indistinct. Male terminalia with aedeagus of moderate length and breadth and paramere narrowly emarginated at posterior margin, with its inner process short, slender, rounded at tip. Body length 8.3–9.8 mm, wing length 8.2–10.6 mm. Description. Male. Head dark brown to almost black, suffused with grey pruinosity on frons and vertex, somewhat shiny on rostrum. Palpus black. Antenna 14-segmented, short, not reaching to base of wing. Scape black, pedicel and flagellomeres paler, brown to yellowish brown. Flagellomeres elongate, subcylindrical. Longest verticils very long, about five times as long as their respective flagellomeres. Thorax generally obscure yellow to yellowish brown. Pronotum brown dorsally, yellowed laterally. Prescutum and scutum with broad, dark brown, median area demarcated with prescutal setae, darker anteriorly, slightly paler posteriorly, sides of prescutum yellowish brown. Scutal lobes similarly dark as median prescutal area, obscure yellow in between. Scutellum pale yellow anteriorly, darker posteriorly. Mediotergite generally yellowish brown, paler anterolaterally. Pleuron essentially obscure yellow, darker on katepisternum. Wing moderately broad, with width-length ratio about 1: 3.5. Wing membrane conspicuously tinged brownish. Wing pattern consisting of three darker spots at anterior margin, diffuse and little-distinct, at origin of Rs, at tip of Sc1, and over R2, added with slightly indicated, diffuse, darker seams especially along Cu and so-called outer cord. Venation usual for Limonia, with discal cell moderately long; M3+4 and M4 subequal in length. Halter with pale stem and darker knob. Legs generally yellow, including coxae and trochanters, tips of femora slightly darkened, distal tarsomeres dark brown; tarsomeres 1 slightly longer than rest of tarsi. Abdomen brown dorsally, paler ventrally. Male terminalia (Figs 3–4) yellowish brown. Tergite 9 essentially semicircular in outline. Its posterior margin broadly rounded, formed by chitinized bar, with distinct U-shaped median notch. Gonocoxite usual in length and breadth. Gonostylus darkened distally, conical, moderate in length, evenly moderately arched and tapered to narrowly rounded tip, only slightly swollen in proximal half. Aedeagus of moderate length and breadth, considerably broad in proximal half, with another, narrower enlargement before apex. Paramere narrowly emarginated at posterior margin, with its inner process short, generally slender, rounded at tip. Female resembling male in general appearance. Female terminalia with cercus slightly upturned, subacute at tip, subequal in length to tergite 10. Hypogynial valve straight, reaching beyond middle of cercus. Differential diagnosis. This new species, probably a member of the L. phragmitidis (= L. tripunctata) species group (cf. SAVCHENKO 1985), is distinctive by having its wing membrane conspicuously tinged brownish. A significant feature of the male terminalia is the shape of the paramere which is narrowly emarginated at its posterior margin, with its inner process slender, and rounded at the tip. A similar condition, with other features quite different, is known in L. sylvicola and L. eos Starý & Savchenko, 1976. Etymology. The name of this new species, opacipennis, a combination of opacus (= dark) and penna (= wing), refers to its brownish wing membrane. An adjective in nominative singular.Published as part of Starý, Jaroslav, 2017, Four new species of Limonia from the Mediterranean (Diptera: Limoniidae), pp. 713-721 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 57 (2) on pages 716-717, DOI: 10.1515/aemnp-2017-0096, http://zenodo.org/record/531659

Molophilus (Molophilus) creticola Starý, 2011, sp. n.

Molophilus (Molophilus) creticola sp. n. Figures 1–3 Diagnosis. Medium-sized species within Molophilus. Body dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Gonocoxite of male terminalia without dorsal and lateral lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, slender, glabrous, pointed at tip. Inner gonostylus much stouter, obtuse at apex. Wing length 3.7–5.8 mm. Description. Male. Head. Antenna of moderate length, extending about to wing base, dark brown throughout. Flagellomeres generally ovoid to long-ovoid, with longest verticils slightly exceeding length of their respective segments. Thorax dark brown, suffused with dense greyish pruinosity, restrictedly patterned with yellow. Prescutum and scutum dark greyish brown, with yellow area lateral to prescutal pit; scutal lobe yellowed anteromedially and posterolaterally. Scutellum mostly yellow, dark greyish brown anteriorly. Postscutellum dark greyish brown, narrowly yellowed laterally. Pleuron dark greyish brown, yellowed on dorsopleural membrane and basalare. Wing membrane infuscated; venation generally as for genus. Halter pale yellow throughout. Legs with coxae, trochanters and bases of femora yellowish brown, rest of legs darkened, brown to dark brown. Abdomen dark brown. Male terminalia (Figs 1–3) dark brown. Dorsal portion of gonocoxite rather short, without dorsal and lateral lobe, subquadrangular in lateral view, slightly narrowed distally, obliquely truncate at apex. Short, darkened, rounded tooth at posteroventral edge of gonocoxite, directed mesally. Lateral excision rather wide. Ventral lobe of gonocoxite generally short and stout, conical, with subacute tip, much shorter than dorsal portion of gonocoxite. Both gonostyli mostly pale, only weakly pigmented distally. Outer gonostylus slender, glabrous, generally straight or slightly curved dorsally, pointed at tip, reaching to about three fourths length of inner gonostylus. The latter much stouter, exceeding ventral lobe by about two thirds its length, more or less straight, narrowing distally to obtuse apex. Aedeagus of moderate length and breadth, expanded before mid-length in lateral view and bent ventrally, then gradually tapered to slightly upturned tip. Aedeagal plate large, generally oval in ventral/ventrocaudal aspect, with short but comparatively stout, curved, blackened median spine. Female resembling male in general appearance. Material examined. Holotype 3: Greece, Crete, Khania Region, Kakopetros, brook, 23.v. 2004 (J. Starý leg.) (SMOC). Paratypes (35 3, 23 Ƥ): Greece: Crete: Khania Region: Kakopetros (400 m), 13.v. 1979, 1 3, 1 Ƥ; Kotsifiana (500 m), 20.v. 1977, 2 3, 1 Ƥ; Fassas valley W Chliaro (260–310 m), 18.– 20.v. 1977, 1 3, 3 Ƥ (all H. Malicky leg.) (all ZFMK; in ethanol); Kakopetros, brook, 23.v. 2004, 10 3, 4 Ƥ; Voukolies, brook, 23.v. 2004, 1 3; Alikianos, Xekollimenos River, 14.v. 2004, 3 Ƥ, 17.v. 2004, 1 3; Prases, 2 km W, 21.v. 2004, 8 3, 3 Ƥ; Prases, 4 km SW, 18.v. 2004, 2 Ƥ, 21.v. 2004, 2 3, 1 Ƥ; Samaria gorge, 14.v. 2004, 1 3; Stilos, Kiliaris River, 13.v. 2004, 5 3, 1 Ƥ; Episkopi, Petres River, 16.v. 2004, 3 3, 4 Ƥ (all J. Starý leg.) (JSO, SMOC). Etymology. The compound name of this new species, creticola, refers to its occurrence in Crete; the final component of the name derives from the Latin incola (= the dweller, inhabitant). A noun in nominative singular, standing in apposition to the generic name. Discussion. The male terminalia of this new species have the dorsal portion of the gonocoxite of generally simple structure, without the dorsal and lateral lobes, thus fitting in the group of species more or less closely related to M. (M.) undulatus Tonnoir in Goetghebuer & Tonnoir, 1920; most of them having been described relatively recently (Caspers 1980, Mendl 1986, Savchenko 1986, Starý 1978, 1992, Starý & Freidberg 2007). Besides the body colouration and some less conspicuous external traits they differ from each other by various details of the male terminalia, the most indicative of these being the structure of the aedeagus and aedeagal plate (Starý 1992). M. (M.) creticola sp. n. appears closest to M. (M.) cypricola sp. n. below, being practically identical with it externally. In M. (M.) creticola sp. n. the dorsal portion of the gonocoxite is shorter and somewhat narrowed distally (longer and parallel-sided in cypricola sp. n.), and the ventral lobe is broader at its base and more sharpened at its tip. The gonostyli of the two species differ in many traits: they are almost pale and generally shorter in M. (M.) creticola sp. n. (darkly pigmented in cypricola sp. n.), exhibiting further differences in shape. In addition, the outer gonostylus is glabrous in M. (M.) creticola sp. n., whereas it is microscopically setulose in M. (M.) cypricola sp. n., which appears to be a diagnostic difference. The aedeagal complex likewise shows differences, chiefly in the median spine of the aedeagal plate, which is distinctly shorter in M. (M.) creticola sp. n. (cf. Figs 1–3 and 4–6). For other details, see Discussion of M. (M.) cypricola sp. n. Distribution. Greece (Crete).Published as part of Starý, Jaroslav, 2011, Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae), pp. 45-62 in Zootaxa 2999 on pages 46-47, DOI: 10.5281/zenodo.20328

Molophilus (Molophilus) klementi Mendl 1973

Molophilus (Molophilus) klementi Mendl, 1973 Material examined. France: Alpes-Maritimes, Le Bar-sur-Loup, 5.5 km NNE (N Saut-du-Loup) (380 m), 14.vi. 1994, 2 3, 1 Ƥ (J.-P. Haenni & C. Dufour leg.) (JSO). Distribution. Austria, Germany, Romania, Switzerland. First record for France.Published as part of Starý, Jaroslav, 2011, Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae), pp. 45-62 in Zootaxa 2999 on page 60, DOI: 10.5281/zenodo.20328

Molophilus (Molophilus) ibericus

Molophilus (Molophilus) ibericus nom. n., stat. n. Figures 17, 19, 20 Molophilus gladius var. obscura Lackschewitz 1940 a: 23 (diagnosis). Diagnosis. Rather small species within Molophilus. Body dark brown, restrictedly patterned with yellow, suffused with greyish pruinosity. Gonocoxite of male terminalia without dorsal lobe. Outer (dorsal) gonostylus shorter than inner (ventral) gonostylus, spike-shaped, subacute at tip. Inner gonostylus tapered and curved downwards distally, pointed at apex. Wing length 3.3–4.6 mm. Redescription. Male. Head. Antenna (Fig. 17) short, not reaching wing base, dark brown throughout. Flagellomeres ovoid, with longest verticils slightly exceeding their respective segments; pubescence short and sparse. Thorax dark brown, restrictedly patterned with yellow, suffused with dense greyish pruinosity. Prescutum and scutum dark greyish brown, with yellow area lateral to prescutal pit, including adjacent part of paratergite; scutal lobe obscure yellow posterolaterally. Scutellum bright yellow, dark greyish brown on margins. Postscutellum dark greyish brown, yellowed anterolaterally. Pleuron dark greyish brown, yellowed on dorsopleural membrane and basalare. Wing membrane slightly infuscated; venation generally as for genus. Halter dark yellow, with paler knob. Legs with coxae, trochanters and bases of femora brown, rest of legs dark brown. Abdomen dark greyish brown. Male terminalia (Figs 19, 20) light brown. Dorsal portion of gonocoxite rather short, without dorsal lobe, only expanded posterodorsally and obliquely truncate apically in lateral view, with distinct, setiferous lobule at posteroventral edge. Lateral lobe short, not extending beyond other lobes. Lateral excision rather wide and comparatively shallow. Ventral lobe of gonocoxite of moderate length and breadth, slightly extending beyond apex of dorsal portion of gonocoxite. Both gonostyli darkly pigmented. Outer gonostylus spikeshaped, gently curved ventrally, provided with a few spinules distally, subacute at tip, reaching to about three fourths length of inner gonostylus. The latter smooth, exceeding ventral lobe by about one third its length, tapered and hook-shaped distally, pointed at apex. Aedeagus comparatively short and stout, curved, with middle part arching dorsally, conspicuously expanded in lateral aspect, and with apex drawn out into slender point extending ventrocaudally. Aedeagal plate moderate in size, generally oval in ventral/ventrocaudal aspect, with three dark, slightly curved spines: a median and two posterolateral spines. Female resembling male in general appearance. Type material examined. Lackschewitz (1940 a: 24) based his “ var. obscura ” of M. gladius (= propinquus) on the following material: “ Spanien, Aragonien, Albarracin, 22.– 30. VI. 1924, 3 ƤƤ Typen (Zerny); Marokko, Gr. Atlas, Tachdirt (2200–2900 m), 11.– 19.VII. 1933, 4 3 2 Ƥ (Zerny)”. Since, however, Lackschewitz (1940 a) nominated as types only the one male (see note under Type material examined of testaceus for the interpretation of the single male symbol) and an unspecified number of females from Spain, only these are syntypes; the Moroccan specimens are excluded from the type series (ICZN 1999, Article 72.4.6). At present, 1 3 and 1 Ƥ from “ Spanien, Aragonien, Albarracin” and 4 3 and 2 Ƥ from “ Marokko, Gr. Atlas, Tachdirt” are deposited in NHMW (P. Sehnal, pers. comm.). I have examined the Spanish and two Moroccan males, and I here select the Spanish male as lectotype. Lectotype 3 (by present designation): Spain, Aragón, Albarracin, 22.– 30.vi. 1924 (H. Zerny leg.) (NHMW), labelled “Aragon Albarracin / 22.– 30.VI ’ 24 Zerny” (printed), “Mol. gladius Meij. / obscura n. var. / det. Lacksch.” (printed, orange). Labelled as lectotype by the present author (“ LECTOTYPE / Molophilus / obscurus Lacksch. stat. n. 3 / J. Starý 2010 ”; printed red label) and identified as M. (M.) ibericus nom. n. The specimen is micropinned on a piece of foam, with left hind and right fore and hind legs missing; apex of abdomen is cut off. Terminalia mounted by Lackschewitz in Canada balsam between celluloid slides, pinned with the specimen. The two nontype Moroccan males are listed below under Other material examined. The lectotype is designated here to stabilize the concept of the name with respect to M. (M.) propinquus. Other material examined (75 3, 1 Ƥ): Spain: Andalucía: Ronda, Rio Grande, 5.iv. 2005, 1 3; El Burgo, Rio Turón, 5.iv. 2005, 3 3; Guaro (nr. Coin), brook, 4.iv. 2005, 12 3; Coin, Rio Seco, 30.iii. 2005, 7 3, 4.iv. 2005, 7 3, 7.iv. 2005, 19 3; Alhaurin el Grande, 5 km E, 30.iii. 2005, 1 3; Fuengirola, Rio de Fuengirola, 6.iv. 2005, 2 3; Benalmádena, 28.iii. 2005, 13 3, 9.iv. 2005, 3 3; El. Romeral (nr. Cártama), watercourse, 31.iii. 2005, 1 3 (all J. Starý leg.) (all JSO); Sierra Nevada Mts, Veleta road (1,500–1,800 m), 10.vii. 1977, 1 3, 1 Ƥ (H. Aspöck, H. Rausch & P. Ressl leg.) (ZFMK; in ethanol); Baza env., 12.v. 1979, 1 3 (J. Roháček leg.) (JSO). Morocco: Gr. Atlas, Tachdirt (2,200–2,900 m), 11.– 19.vii. 1933, 2 3 (H. Zerny leg.) (NHMW) [listed as “ var. obscura ” by Lackschewitz 1940 a; non-types (see above)]; Ht Atlas, Massif Toubkal, Imlil, 17 km S Asni (1,700–1,900 m), 7.vii. 1977, 2 3 (H. & T. v. Oorschot, E. Houkes & P. Oosterbroek leg.) (ZMAN, JSO). Etymology. The name of this species, ibericus, was proposed by H. Mendl for his undescribed species, referring to the Iberian Peninsula where Mendl’s specimens (now in ZFMK, see Other material examined) come from, as well as most other material examined here. The name is to be deemed to be a latinized adjective in nominative singular Discussion. Lackschewitz (1940 a) established this species as “ var. obscura n. var. ” of M. gladius de Meijere, 1920 [= M. propinquus (Egger, 1863)]. The name obscura is subspecific in terms of Article 45.6. 4. of ICZN (1999), and the taxon is here raised to species rank. The name, however, is a junior secondary homonym of Erioptera obscura Meigen, 1818 (now in Molophilus). Hence, a new replacement name is proposed here, ibericus nom. n., based on Mendl’s manuscript name (see Etymology). H. Mendl considered his ibericus to be a valid species; similarly, Savchenko (1982: 212, footnote) suggested Lackschewitz’s “ var. obscura ” as being possibly distinct from M. (M.) propinquus. M. (M.) propinquus, a common and widely distributed Palaearctic species, and its allies constitute a distinctive species complex distinguished by the structure of the male terminalia, primarily the ventrally curved aedeagus and the aedeagal plate provided with three spines subequal in size. M. (M.) alexanderianus Nielsen, 1963 (Middle Asia), M. (M.) pseudopropinquus Mendl, 1973 (France, Germany, Romania, Slovakia, Switzerland), and M. (M.) ponticus Savchenko, 1982 (Crimea, North Caucasus, Transcaucasia) belong here and are considered distinct species based on details of the male terminalia. Since, however, a certain genital variation may occur within each species, in particular for the widespread M. (M.) propinquus, all the species are more significantly differentiated by their male antennae, which differ considerably among the species by their overall length and the length and density of both verticils and pubescence on the flagellomeres. Some other forms, such as M. triacanthus Alexander, 1934 (Russian Far East, North Korea, Japan) and M. debilispinus Alexander, 1957 (Japan), are insufficiently known and currently treated as subspecies of M. (M.) propinquus (see Oosterbroek 2010). A form illustrated by Savchenko (1982, Fig. 98 / 4) as „ M. (s. str.) sp. cf. occultus Meijere “ is of doubtful taxonomic status, but could represent a separate taxon, having in fact nothing in common with M. (M.) occultus. M. (M.) ibericus nom. n. clearly fits in this species complex. It differs from M. (M.) propinquus externally by its darker body colouration and somewhat smaller size, having the antennae shorter, with moderately long verticils and only sparse and short pubescence; the antennae of M. (M.) propinquus are longer, with somewhat longer verticils and the pubescence conspicuous, long, suberect and dense (cf. Figs 17 and 18) [the antennae of alexanderianus are still longer, with flagellomeres subcylindrical, verticils indistinct, and pubescence conspicuous (see Savchenko 1982, Fig. 87 / 4); those of pseudopropinquus are moderately long, with verticils twice the length of their respective segments and pubescence conspicuous; the antennae of ponticus are short, long-verticilate and inconspicuously pubescent (see Savchenko 1982, Fig. 87 / 3)]. In the structure of the male terminalia, M. (M.) ibericus nom. n. is distinguished by having the lateral lobe short, not protruding (Fig. 19), whereas, in M. (M.) propinquus, this part extends posteriorly beyond other lobes of the gonocoxite. The outer gonostylus is shorter and subacute at the tip in M. (M.) ibericus nom. n. (longer and rather obtuse at tip in propinquus, with abundant, black spinules or spinoid setulae present not only at the apex but also at the inner margin along most of the length of the gonostylus), and the inner gonostylus is more slender and less conspicuously curved distally. The aedeagus is more expanded in the lateral view in M. (M.) ibericus nom. n., with a longer narrowed point and the spines on the aedeagal plate may be somewhat less pronounced. Distribution. Spain, Morocco. If Lackschewitz’s (1940 a) M. gladius var. obscura is the only record for M. (M.) propinquus from Spain in the recent checklist (Eiroa & Báez 2002), then the latter species should be excluded from the list.Published as part of Starý, Jaroslav, 2011, Descriptions and records of the Palaearctic Molophilus Curtis (Diptera, Limoniidae), pp. 45-62 in Zootaxa 2999 on pages 52-53, DOI: 10.5281/zenodo.20328

Neolimnophila alaskana Starý 2019, stat. nov.

Neolimnophila alaskana (Alexander, 1924) stat. nov. (Figs 7–9, 12) Limnophila (Neolimnophila) ultima alaskana Alexander, 1924: 10 (description). ? Crypteriella Sverdrupi Soot-Ryen, 1928:6 (description), Figs 1 (wing), 2 (antenna), 3 (tibial spur), 4 (ovipositor). Type material examined: HOLOTYPE: ♁, U.S.A.: ALASKA: Healy, 24.vi.1921, specimen + slide (J. M.Aldrich leg.) (USNM) [the date given in the description (ALEXANDER 1924: 10) is “ July 24, 1921 “], labelled “Healy Alaska / VI-24-21” (partly printed, white label), “ HOLOTYPE / Limnophila / ultima / alaskana / C.P. Alexander ” (partly printed, red label). The specimen is pinned, without right wing, left fore and right hind legs, and apex of abdomen. The slide (Canada balsam) with wing (right) under one circular coverslip and male terminalia under another, labelled as for the specimen, with the inscription “ HOLOTYPE 2287”. PARATYPE: U.S.A.: ALASKA: Flat, 5.viii.1919, 1 ♀ (+ slide) (A. H. Twitchell leg.) (USNM), labelled “Flat Alaska / Aug 5 1919 ” (printed, white label), “A H Twitchell / Collector” (printed, white label), “ PARATYPE / Limnophila / ultima / alaskana / C.P. Alexander” (partly printed, blue label). The specimen is glued onto a triangular cardboard point, without antennae, right wing, and with only left mid femur + tibia, and left hind femur. The slide (Canada balsam) with wing (right) under a circular coverslip, labelled as for the specimen, with the inscription “ PARATYPE 2287”. Other material examined (3 ♁♁ 1 ♀): CANADA: NORTHWEST TERRITORIES: Aklavik, Mackenzie River, 4.ix.1929, 1 ♀ (+ slide), Aklavik, 27.viii.1931, 1 ♁ (+ slide) (both O. Bryant leg.) (both USNM). CZECH REPUBLIC: MORAVIA: Moravskoslezské Beskydy Mts, Dolní Bečva, “Kamenné” (650 m), 29.vi.1994, 1 ♁ (J. Starý leg., at light) (JSOC). RUSSIA: EAST SIBERIA: Yakutia, Srednekolymskiy District,Agrakhtakh, 20.vi.1971, 1 ♁ (P. Polyakova & Bobrova leg.) (JSOC). Material not examined by the author (4 ♁♁; for explanation see under N. placida): KAZAKHSTAN: AKMOLA REGION: Atbasar, 30.viii.1936, 1 ♁ (P. D. Rezvoy leg.) (ZISP). RUSSIA: CENTRAL EUROPEAN RUSSIA: Moscow Region, Orekhovo-Zuyevo District, Smolevo village env., 55.5788°N, 38.6662°E, 16.–20.viii.2011, 1 ♁ (in ethanol) (K. Tomkovich leg.) (ZMUM). NORTH EUROPEAN RUSSIA: Murmansk Region, Khibiny Mts, basin of Vudyavr Lakes, 26.vii.1933, 1 ♁ (V. Y. Fridolin leg.) (ZISP). EAST SIBERIA: Zabaykalskiy Krai, Sretenskiy District, between Ivanovka and Kara Rivers (tributaries of Shilka River), 27.viii.1926, 1 ♁ (R. F. Hecker leg.) (ZISP). Diagnosis. Antenna moderately long, with verticils rather long, longest ones subequal in length to their respective segments. Dorsum of thorax heavily suffused with bluish dark grey pruinosity, prescutum with four brown stripes. Wing rather broad, width-length ratio about 1: 3. Male terminalia with two spines at base of gonocoxite, ventral spine about two thirds length of dorsal one, outer gonostylus slender and rather long, paramere broad, its distal outer extension smaller than that in N. placida. Wing length 7.8–8.7 mm. Redescription. Male. Head dark brown, suffused with bluish dark grey pruinosity on frons and vertex. Palpus and rostrum dark brown. Antenna dark brown throughout, moderately long, reaching to about base of wing, scape greyish pruinose. Flagellum consisting of basal fusion element and subsequent 10 elongate flagellomeres, with verticils sparse, longest ones subequal in length to their respective segments. Thorax generally dark brown, heavily suffused with bluish dark grey pruinosity, especially on dorsum, indistinctly paler on anterior paratergite and postrerolaterally on scutal lobes. Prescutum with four brown stripes. Wing rather broad, width-length ratio about 1: 3, sometimes with faintly indicated stigma. Venation usual for Neolimnophila. Sc1 ending about opposite fork of Rs. R2 (cross-vein r) at or slightly before fork of R3+4. R3 slightly shorter than R3+4. Latter vein sinuous, upturned distally. Cross-vein m-cu at from one fourth to one third length of discal cell (Fig. 12). Halter pale throughout. Legs brown, with setae on femora comparatively long and projecting, about twice diameter of femur. Abdomen dark brown. Male terminalia (Figs 7–9). Tergite 9 interrupted medially. Gonocoxite with two spines at base: longer, positioned dorsally, subequal in length to corresponding spine in N. carteri and N. placida, about one third length of gonocoxite; shorter, about two thirds length of dorsal spine, positioned more ventrally and medially. Both spines with delicate membranous scales distally. Outer gonostylus long and slender, about three fifths length of gonocoxite, almost parallel-sided for about three fourths its length, with apical hook, microscopically serrate at outer margin. Paramere rather broad, truncate at apex, its distal outer corner less extended outwardly than in N. placida. Female resembling male in general appearance, including length and structure of antenna and outline of wing. Female terminalia not essentially different externally from those of other species. Distribution. So far only listed from Alaska, so probably new to Canada. First records from the Palaearctic Region.Published as part of Starý, Jaroslav, 2019, Neolimnophila alaskana (Alexander, 1924) stat. nov., a species new to the Palaearctic Region (Diptera: Limoniidae), pp. 53-58 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 59 (1) on page 56, DOI: 10.2478/aemnp-2019-0004, http://zenodo.org/record/450542