Intraspecific Variation in Prey Susceptibility Mediates the Consumptive Effect of Predation: A Case Study of Yellowstone Elk and Wolves

The reintroduction of wolves (Canis lupus) to Yellowstone National Park starting in 1995 is an important case study for understanding the consequences of predation on a prey population. Simulation studies conducted prior to and shortly after wolf reintroduction predicted that wolf predation of elk (Cervus canadensis) would have a modest influence on elk abundance. Predation of elk by wolves has been well documented and elk have remained the primary prey for wolves despite a decline in elk abundance. I used two quantitative approaches to estimate the influence of wolf predation on adult female elk survival and abundance in northern Yellowstone and adjacent Montana during 2000-2017. My results suggest that, while wolves did kill adult female elk aged 2-14 years old, these elk generally had high survival. Elk were more likely to be killed by wolves as they aged. Wolf predation of adult female elk was primarily restricted to older individuals that generally comprised a small proportion of the total elk population. Harsh environmental conditions, such as heavy snow, increased mortality of adult elk, but elk aged 2-9 years old retained high survival regardless of the environmental conditions. The observed decline in elk abundance across the 17-year study was primarily due to mortality of 2-14 year-old elk that died due to causes unrelated to wolves, including malnutrition, harvest, and other predators. I could not estimate the full impact of wolves on female elk abundance because of the lack of data on elk calf and yearling mortality. However, wolves likely had a smaller impact on the elk population than did non-wolf causes of elk mortality. These findings clarify how the impact of predation on a prey population may be limited by the age of the prey that are consumed and the relative importance of the predated individuals to the population (i.e., their reproductive potential)

Signatures of Young Star Formation Activity Within Two Parsecs of Sgr A*

We present radio and infrared observations indicating on-going star formation activity inside the $\sim2-5$ pc circumnuclear ring at the Galactic center. Collectively these measurements suggest a continued disk-based mode of on-going star formation has taken place near Sgr A* over the last few million years. First, VLA observations with spatial resolution 2.17$"\times0.81"$ reveal 13 water masers, several of which have multiple velocity components. The presence of interstellar water masers suggests gas densities that are sufficient for self-gravity to overcome the tidal shear of the 4$\times10^6$ \msol\, black hole. Second, SED modeling of stellar sources indicate massive YSO candidates interior to the molecular ring, supporting in-situ star formation near Sgr A* and appear to show a distribution similar to that of the counter-rotating disks of $\sim$100 OB stars orbiting Sgr A*. Some YSO candidates (e.g., IRS~5) have bow shock structures suggesting that they have have gaseous disks that are phototoevaporated and photoionized by the strong radiation field. Third, we detect clumps of SiO (2-1) and (5-4) line emission in the ring based on CARMA and SMA observations. The FWHM and luminosity of the SiO emission is consistent with shocked protostellar outflows. Fourth, two linear ionized features with an extent of $\sim0.8$ pc show blue and redshifted velocities between $+50$ and $-40$ \kms, suggesting protostellar jet driven outflows with mass loss rates of $\sim5\times10^{-5}$ solar mass yr$^{-1}$. Finally, we present the imprint of radio dark clouds at 44 GHz, representing a reservoir of molecular gas that feeds star formation activity close to Sgr A*.Comment: 38 pages, 10 figures, ApJ (in press

Mid Infrared Hydrogen Recombination Line Emission from the Maser Star MWC 349A

We have detected and spectrally resolved the mid-IR hydrogen recombination lines H6(alpha)(12.372 micrometers), H7(alpha)(19.062 micrometers), H7(beta)(l1.309 micrometers) and H8(gamma)(12.385 micrometers) from the star MWC349A. This object has strong hydrogen maser emission (reported in the millimeter and submillimeter hydrogen recombination lines from H36(alpha) to H21(alpha)) and laser emission (reported in the H15(alpha), H12(alpha) and H10(alpha) lines). The lasers/masers are thought to arise predominantly in a Keplerian disk around the star. The mid-IR lines do not show evident signs of lasing, and can be well modeled as arising from the strong stellar wind, with a component arising from a quasi-static atmosphere around the disk, similar to what is hypothesized for the near IR (less than or equal to 4 micrometers) recombination lines. Since populations inversions in the levels producing these mid-IR transitions are expected at densities up to approximately 10(exp 11)/cu cm, these results imply either that the disk does not contain high-density ionized gas over long enough path lengths to produce a gain approximately 1, and/or that any laser emission from such regions is small compared to the spontaneous background emission from the rest of the source as observed with a large beam. The results reinforce the interpretation of the far-IR lines as true lasers

The reddest ISO-2MASS quasar

In the course of the NIR/MIR AGN search combining the 6.7 mu ISOCAM Parallel Survey and 2MASS we have discovered 24 type-1 quasars about a third of which are too red to be discriminated by optical/UV search techniques. Here we report on a detailed case study of the reddest type-1 quasar of our sample (J2341) at redshift z=0.236 with M_K=-25.8 and J-K=1.95. We performed spectroscopy in the optical with VLT/FORS1 and in the MIR with Spitzer as well as NIR imaging with ISPI at CTIO. The optical and NIR observations reveal a star forming emission-line galaxy at the same redshift as the quasar with a projected linear separation of 1.8 arcsec (6.7 kpc). The quasar and its companion are embedded in diffuse extended continuum emission. Compared with its companion the quasar exhibits redder optical-NIR colours, which we attribute to hot nuclear dust. The MIR spectrum shows only few emission lines superimposed on a power-law spectral energy distribution. However, the lack of strong FIR emission suggests that our potentially interacting object contains much less gas and dust and is in a stage different from dust reddened ULIRG-AGN like Mrk 231. The optical spectrum shows signatures for reddening in the emission-lines and no post-starburst stellar population is detected in the host galaxy of the quasar. The optical continuum emission of the active nucleus appears absorbed and diluted. Even the combination of absorption and host dilution is not able to match J2341 with standard quasar templates. While the BLR shows only a rather moderate absorption of E_(B-V)=0.3, the continuum shorter than 4500 AA requires strong obscuration with E_(B-V)=0.7, exceeding the constraints from the low upper limit on the 9.7 mu silicate absorption. This leads us to conclude that the continuum of J2341 is intrinsically redder than that of typical quasars.Comment: 8 pages, 9 figure

The environments of z~1 Active Galactic Nuclei at 3.6um

We present an analysis of a large sample of AGN environments at z~1 using stacked Spitzer data at 3.6um. The sample contains type-1 and type-2 AGN in the form of quasars and radio galaxies, and spans a large range in both optical and radio luminosity. We find, on average, that 2 to 3 massive galaxies containing a substantial evolved stellar population lie within a 200-300 kpc radius of the AGN, constituting a >8-sigma excess relative to the field. Secondly, we find evidence for the environmental source density to increase with the radio luminosity of AGN, but not with black-hole mass. This is shown first by dividing the AGN into their classical AGN types, where we see more significant over-densities in the fields of the radio-loud AGN. If instead we dispense with the classical AGN definitions, we find that the source over-density as a function of radio luminosity for all our AGN exhibits a positive correlation. One interpretation of this result is that the Mpc-scale environment is in some way influencing the radio emission that we observe from AGN. This could be explained by the confinement of radio jets in dense environments leading to enhanced radio emission or, alternatively, may be linked to more rapid black-hole spin brought on by galaxy mergers.Comment: 13 pages, 12 figures, accepted by MNRA

Waterbird monitoring and habitat association modeling to inform tidal marsh restoration in an urbanized estuary

The San Francisco Bay (SFB), like many other urbanized estuaries, is a critical wintering and stop-over area for migratory waterbirds. More than a million wintering waterbirds annually rely on a mosaic of natural and managed habitats in SFB, including former salt ponds. The South Bay Salt Pond Restoration Project plans restore 50 to 90% of a 6,100 hectare former salt production pond complex to tidal marsh, while maintaining the rest as foraging and roosting areas for migratory birds. Since 2002, we have evaluated migratory waterbird use of pre- and post-restoration salt ponds in south SFB. Our approach has been to use monthly surveys, applied studies and modeling to evaluate avian response to this changing habitat. Trend analyses indicate that overall waterbird numbers have nearly doubled across the complex from 2003-2014. Dabbling ducks increased significantly during initial phases of the project, while diving ducks, small and medium shorebird abundances declined initially, but rebounded in recent years. We modeled relationships between abundances of multiple species and habitat characteristics at both landscape and micro-scales. Modeling results suggest that the importance of different habitat characteristics varies among waterbird guilds. For example, foraging diving duck abundances were higher in deep, un-breached ponds, located close to the edge of SFB, while foraging small shorebirds predominated in shallow, breached ponds containing islands. Our research highlights important habitat characteristics both within and across wetland areas, and provides a unique opportunity to guide future management and restoration decisions for multiple species across large landscapes in SFB and other urbanized estuaries

Accurate masses and radii of normal stars: modern results and applications

This paper presents and discusses a critical compilation of accurate, fundamental determinations of stellar masses and radii. We have identified 95 detached binary systems containing 190 stars (94 eclipsing systems, and alpha Centauri) that satisfy our criterion that the mass and radius of both stars be known to 3% or better. To these we add interstellar reddening, effective temperature, metal abundance, rotational velocity and apsidal motion determinations when available, and we compute a number of other physical parameters, notably luminosity and distance. We discuss the use of this information for testing models of stellar evolution. The amount and quality of the data also allow us to analyse the tidal evolution of the systems in considerable depth, testing prescriptions of rotational synchronisation and orbital circularisation in greater detail than possible before. The new data also enable us to derive empirical calibrations of M and R for single (post-) main-sequence stars above 0.6 M(Sun). Simple, polynomial functions of T(eff), log g and [Fe/H] yield M and R with errors of 6% and 3%, respectively. Excellent agreement is found with independent determinations for host stars of transiting extrasolar planets, and good agreement with determinations of M and R from stellar models as constrained by trigonometric parallaxes and spectroscopic values of T(eff) and [Fe/H]. Finally, we list a set of 23 interferometric binaries with masses known to better than 3%, but without fundamental radius determinations (except alpha Aur). We discuss the prospects for improving these and other stellar parameters in the near future.Comment: 56 pages including figures and tables. To appear in The Astronomy and Astrophysics Review. Ascii versions of the tables will appear in the online version of the articl

The ~0.9 mJy sample: A mid-infrared spectroscopic catalog of 150 infrared-luminous, 24 micron selected galaxies at 0.3<z<3.5

We present a catalog of mid-infrared (MIR) spectra of 150 infrared (IR) luminous galaxies in the Spitzer extragalactic first look survey obtained with IRS on board Spitzer. The sample is selected to be brighter than ~0.9 mJy at 24 micron and it has a z distribution in the range [0.3,3.5] with a peak at z=1. It primarily comprises ultraluminous IR galaxies at z>1 and luminous IR galaxies at z<1, as estimated from their monochromatic 14 micron luminosities. The number of sources with spectra that are dominated by an active galactic nucleus (AGN) continuum is 49, while 39 sources have strong, star-formation related features. For this classification, we used the equivalent width (EW) of the 11.3 micron polycyclic aromatic hydrocarbon (PAH) feature. Several intermediate/high z starbursts have higher PAH EW than local ULIRGs. An increase in the AGN activity is observed with increasing z and luminosity, based on the decreasing EW of PAHs and the increasing [NeIII]/[NeII] ratio. Spectral stacking leads to the detection of the 3.3 micron PAH, the H2 0-0 S(1) and S(3) lines, and the [NeV] line. We observe differences in the flux ratios of PAHs in the stacked spectra of IR-luminous galaxies with z or luminosity, which are not due to extinction effects. When placing the observed galaxies on IR color-color diagrams, we find that the wedge defining AGN comprises most sources with continuum-dominated spectra, but also contains many starbursts. The comparison of the 11.3 micron PAH EW and the H-band effective radius, measured from HST data, indicates that sources with EW>2 micron, are typically more extended than ~3 kpc. However, there is no strong correlation between the MIR spectral type and the near-IR extent of the sources. [Abridged].Comment: accepted for publication in the Astrophysical Journa

Physical Conditions in the Ionized Gas of 30 Doradus

We present a mid-infrared spectroscopic data cube of the central part of 30 Doradus, observed with Spitzer's IRS and MIPS/SED mode. Aromatic dust emission features and emission lines from molecular and atomic hydrogen are detected but not particularly strong. The dominant spectral features are emission lines from moderately ionized species of argon, neon, and sulphur, which are used to determine the physical conditions in the ionized gas. The ionized gas excitation shows strong variations on parsec scales, some of which can plausibly be associated with individual hot stars. We fit the ionic line strengths with photoionization and shock models, and find that photoionization dominates in the region. The ionization parameter U traces the rim of the central bubble, as well as highlighting isolated sources of ionization, and at least one quiescent clump. The hardness of the ionizing radiation field T_rad reveals several "hot spots" that are either the result of individual very hot stars or trace the propagation of the diffuse ionizing field through the surrounding neutral cloud. Consistent with other measurements of giant molecular hydrogen regions, log(U) ranges between -3 and -0.75, and T_rad between 30000 and 85000K.Comment: 32 pages, 26 figures, ApJ accepted. A version with high-resolution images can be found at http://www.astro.virginia.edu/~ged3j/indebetouw20090125.pd

The Spectral Energy Distributions of Red 2MASS AGN

We present infrared (IR) to X-ray spectral energy distributions (SEDs) for 44 red AGN selected from the 2MASS survey on the basis of their red J-K$_S$ color (>2 mag) and later observed by Chandra. In comparison with optically-, radio-, and X-ray selected AGN, their median SEDs are red in the optical and near-IR with little/no blue bump. It thus seems that near-IR color selection isolates the reddest subset of AGN that can be classified optically. The shape of the SEDs is generally consistent with modest absorption by gas (in the X-ray) and dust (in the optical-IR). The levels of obscuration, estimated from X-rays, far-IR and our detailed optical/near-IR color modeling are all consistent implying N_H < few*10^{22} cm^{-2}. We present SED models that show how the AGN optical/near-IR colors change due to differing amounts of reddening, AGN to host galaxy ratio, redshift and scattered light emission and apply them to the sources in the sample. We find that the 2MASS AGN optical color, B-R, and to a lesser extent the near-IR color, J-K$_S$, are strongly affected by reddening, host galaxy emission, redshift, and in few, highly polarized objects, also by scattered AGN light. The obscuration/inclination of the AGN allows us to see weaker emission components which are generally swamped by the AGN.Comment: 52 pages, 17 figures, accepted for publication in Ap