180 research outputs found
The emergence of division of labour in forced associations of normally solitary ant queens
ABSTRACT We tested the hypothesis that division of labour is an inescapable property of social groups, rather than an evolutionary event separate from sociality. We propose a variance-based emergent property model in which division of labour can spontaneously emerge when (1) individuals in a group vary in their intrinsic sensitivity to stimuli for a given task, and (2) the performance of a task by individuals with higher task sensitivities reduces performance of the task by the other group members. Under these conditions, the individuals with higher sensitivity become the task specialists. To determine if division of labour can occur in the absence of direct selection, we created artificial foundress associations (groups of queens during nest establishment) of the ant Pogonomyrmex barbatus, which has no history of cooperative colony founding. We compared the incidence of task specialization in this species to that in Pogonomyrmex californicus, in which foundress associations commonly occur. Levels of task specialization for nest excavation were high in both species, arguing that division of labour emerges in groups regardless of evolutionary history. Consistent with the variance-based model, the role of nest excavation specialist in P. barbatus foundress associations could be predicted by both (1) differences in excavation roles in prior pairs and (2) variation in excavation activity while solitary. Furthermore, the assumption of the excavator role by one foundress dramatically reduced the performance of that task by the other foundress of the pair. We also found a strong negative relationship between excavation activity and foundress survival in P. barbatus, suggesting that division of labour in this context may act as a constraint on social evolution
Exact Hypersurface-Homogeneous Solutions in Cosmology and Astrophysics
A framework is introduced which explains the existence and similarities of
most exact solutions of the Einstein equations with a wide range of sources for
the class of hypersurface-homogeneous spacetimes which admit a Hamiltonian
formulation. This class includes the spatially homogeneous cosmological models
and the astrophysically interesting static spherically symmetric models as well
as the stationary cylindrically symmetric models. The framework involves
methods for finding and exploiting hidden symmetries and invariant submanifolds
of the Hamiltonian formulation of the field equations. It unifies, simplifies
and extends most known work on hypersurface-homogeneous exact solutions. It is
shown that the same framework is also relevant to gravitational theories with a
similar structure, like Brans-Dicke or higher-dimensional theories.Comment: 41 pages, REVTEX/LaTeX 2.09 file (don't use LaTeX2e !!!) Accepted for
publication in Phys. Rev.
The Atacama Cosmology Telescope: Cosmological parameters from three seasons of data
We present constraints on cosmological and astrophysical parameters from
high-resolution microwave background maps at 148 GHz and 218 GHz made by the
Atacama Cosmology Telescope (ACT) in three seasons of observations from 2008 to
2010. A model of primary cosmological and secondary foreground parameters is
fit to the map power spectra and lensing deflection power spectrum, including
contributions from both the thermal Sunyaev-Zeldovich (tSZ) effect and the
kinematic Sunyaev-Zeldovich (kSZ) effect, Poisson and correlated anisotropy
from unresolved infrared sources, radio sources, and the correlation between
the tSZ effect and infrared sources. The power ell^2 C_ell/2pi of the thermal
SZ power spectrum at 148 GHz is measured to be 3.4 +\- 1.4 muK^2 at ell=3000,
while the corresponding amplitude of the kinematic SZ power spectrum has a 95%
confidence level upper limit of 8.6 muK^2. Combining ACT power spectra with the
WMAP 7-year temperature and polarization power spectra, we find excellent
consistency with the LCDM model. We constrain the number of effective
relativistic degrees of freedom in the early universe to be Neff=2.79 +\- 0.56,
in agreement with the canonical value of Neff=3.046 for three massless
neutrinos. We constrain the sum of the neutrino masses to be Sigma m_nu < 0.39
eV at 95% confidence when combining ACT and WMAP 7-year data with BAO and
Hubble constant measurements. We constrain the amount of primordial helium to
be Yp = 0.225 +\- 0.034, and measure no variation in the fine structure
constant alpha since recombination, with alpha/alpha0 = 1.004 +/- 0.005. We
also find no evidence for any running of the scalar spectral index, dns/dlnk =
-0.004 +\- 0.012.Comment: 26 pages, 22 figures. This paper is a companion to Das et al. (2013)
and Dunkley et al. (2013). Matches published JCAP versio
Behavioral genomics of honeybee foraging and nest defense
The honeybee has been the most important insect species for study of social behavior. The recently released draft genomic sequence for the bee will accelerate honeybee behavioral genetics. Although we lack sufficient tools to manipulate this genome easily, quantitative trait loci (QTLs) that influence natural variation in behavior have been identified and tested for their effects on correlated behavioral traits. We review what is known about the genetics and physiology of two behavioral traits in honeybees, foraging specialization (pollen versus nectar), and defensive behavior, and present evidence that map-based cloning of genes is more feasible in the bee than in other metazoans. We also present bioinformatic analyses of candidate genes within QTL confidence intervals (CIs). The high recombination rate of the bee made it possible to narrow the search to regions containing only 17–61 predicted peptides for each QTL, although CIs covered large genetic distances. Knowledge of correlated behavioral traits, comparative bioinformatics, and expression assays facilitated evaluation of candidate genes. An overrepresentation of genes involved in ovarian development and insulin-like signaling components within pollen foraging QTL regions suggests that an ancestral reproductive gene network was co-opted during the evolution of foraging specialization. The major QTL influencing defensive/aggressive behavior contains orthologs of genes involved in central nervous system activity and neurogenesis. Candidates at the other two defensive-behavior QTLs include modulators of sensory signaling (Am5HT(7) serotonin receptor, AmArr4 arrestin, and GABA-B-R1 receptor). These studies are the first step in linking natural variation in honeybee social behavior to the identification of underlying genes
The Atacama Cosmology Telescope: Two-Season ACTPol Spectra and Parameters
We present the temperature and polarization angular power spectra measured by
the Atacama Cosmology Telescope Polarimeter (ACTPol). We analyze night-time
data collected during 2013-14 using two detector arrays at 149 GHz, from 548
deg of sky on the celestial equator. We use these spectra, and the spectra
measured with the MBAC camera on ACT from 2008-10, in combination with Planck
and WMAP data to estimate cosmological parameters from the temperature,
polarization, and temperature-polarization cross-correlations. We find the new
ACTPol data to be consistent with the LCDM model. The ACTPol
temperature-polarization cross-spectrum now provides stronger constraints on
multiple parameters than the ACTPol temperature spectrum, including the baryon
density, the acoustic peak angular scale, and the derived Hubble constant.
Adding the new data to planck temperature data tightens the limits on damping
tail parameters, for example reducing the joint uncertainty on the number of
neutrino species and the primordial helium fraction by 20%.Comment: 23 pages, 25 figure
Studying Parton Energy Loss in Heavy-Ion Collisions via Direct-Photon and Charged-Particle Azimuthal Correlations
Charged-particle spectra associated with direct photon () and
are measured in + and Au+Au collisions at center-of-mass energy
GeV with the STAR detector at RHIC. A hower-shape
analysis is used to partially discriminate between and .
Assuming no associated charged particles in the direction (near
side) and small contribution from fragmentation photons (), the
associated charged-particle yields opposite to (away side) are
extracted. At mid-rapidity () in central Au+Au collisions,
charged-particle yields associated with and at high
transverse momentum ( GeV/) are suppressed by a factor
of 3-5 compared with + collisions. The observed suppression of the
associated charged particles, in the kinematic range and GeV/, is similar for and , and
independent of the energy within uncertainties. These
measurements indicate that the parton energy loss, in the covered kinematic
range, is insensitive to the parton path length.Comment: submitted to Phys. Rev. Lett, 6 pages, 4 figure
Charged and strange hadron elliptic flow in Cu+Cu collisions at = 62.4 and 200 GeV
We present the results of an elliptic flow analysis of Cu+Cu collisions
recorded with the STAR detector at 62.4 and 200GeV. Elliptic flow as a function
of transverse momentum is reported for different collision centralities for
charged hadrons and strangeness containing hadrons , ,
, in the midrapidity region . Significant reduction in
systematic uncertainty of the measurement due to non-flow effects has been
achieved by correlating particles at midrapidity, , with those at
forward rapidity, . We also present azimuthal correlations in
p+p collisions at 200 GeV to help estimating non-flow effects. To study the
system-size dependence of elliptic flow, we present a detailed comparison with
previously published results from Au+Au collisions at 200 GeV. We observe that
() of strange hadrons has similar scaling properties as were
first observed in Au+Au collisions, i.e.: (i) at low transverse momenta,
, scales with transverse kinetic energy, , and
(ii) at intermediate , , it scales with the number of
constituent quarks, . We have found that ideal hydrodynamic calculations
fail to reproduce the centrality dependence of () for
and . Eccentricity scaled values, , are larger
in more central collisions, suggesting stronger collective flow develops in
more central collisions. The comparison with Au+Au collisions which go further
in density shows depend on the system size, number of
participants . This indicates that the ideal hydrodynamic limit is
not reached in Cu+Cu collisions, presumably because the assumption of
thermalization is not attained.Comment: 18 pages, 14 figure
Identified high- spectra in Cu+Cu collisions at =200 GeV
We report new results on identified (anti)proton and charged pion spectra at
large transverse momenta (3<<10 GeV/c) from Cu+Cu collisions at
=200 GeV using the STAR detector at the Relativistic Heavy Ion
Collider (RHIC). This study explores the system size dependence of two novel
features observed at RHIC with heavy ions: the hadron suppression at
high- and the anomalous baryon to meson enhancement at intermediate
transverse momenta. Both phenomena could be attributed to the creation of a new
form of QCD matter. The results presented here bridge the system size gap
between the available pp and Au+Au data, and allow the detailed exploration for
the on-set of the novel features. Comparative analysis of all available 200 GeV
data indicates that the system size is a major factor determining both the
magnitude of the hadron spectra suppression at large transverse momenta and the
relative baryon to meson enhancement.Comment: Submitted to Phys. Rev. C, 9 pages, 5 figure
Measurements of meson production in relativistic heavy-ion collisions at RHIC
We present results for the measurement of meson production via its
charged kaon decay channel in Au+Au collisions at
, 130, and 200 GeV, and in and +Au collisions
at GeV from the STAR experiment at the BNL Relativistic
Heavy Ion Collider (RHIC). The midrapidity () meson transverse
momentum () spectra in central Au+Au collisions are found to be well
described by a single exponential distribution. On the other hand, the
spectra from , +Au and peripheral Au+Au collisions show power-law tails
at intermediate and high and are described better by Levy
distributions. The constant yield ratio vs beam species, collision
centrality and colliding energy is in contradiction with expectations from
models having kaon coalescence as the dominant mechanism for production
at RHIC. The yield ratio as a function of is consistent
with a model based on the recombination of thermal quarks up to GeV/, but disagrees at higher transverse momenta. The measured nuclear
modification factor, , for the meson increases above unity at
intermediate , similar to that for pions and protons, while is
suppressed due to the energy loss effect in central Au+Au collisions. Number of
constituent quark scaling of both and for the meson
with respect to other hadrons in Au+Au collisions at =200 GeV
at intermediate is observed. These observations support quark
coalescence as being the dominant mechanism of hadronization in the
intermediate region at RHIC.Comment: 22 pages, 21 figures, 4 table
Observation of charge-dependent azimuthal correlations and possible local strong parity violation in heavy ion collisions
Parity-odd domains, corresponding to non-trivial topological solutions of the
QCD vacuum, might be created during relativistic heavy-ion collisions. These
domains are predicted to lead to charge separation of quarks along the orbital
momentum of the system created in non-central collisions. To study this effect,
we investigate a three particle mixed harmonics azimuthal correlator which is a
\P-even observable, but directly sensitive to the charge separation effect. We
report measurements of this observable using the STAR detector in Au+Au and
Cu+Cu collisions at =200 and 62~GeV. The results are presented
as a function of collision centrality, particle separation in rapidity, and
particle transverse momentum. A signal consistent with several of the
theoretical expectations is detected in all four data sets. We compare our
results to the predictions of existing event generators, and discuss in detail
possible contributions from other effects that are not related to parity
violation.Comment: 17 pages, 14 figures, as accepted for publication in Physical Review
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