Seabed drag coefficient over natural beds of horse mussels (Atrina zelandica)

Measurements of seabed drag coefficient, C100, were made under tidal currents at four sites in Mahurangi Harbour, New Zealand. At the first three sites the dominant roughness element was the pinnid bivalve, Atrina zelandica (horse mussel). At the fourth site, which was devoid of horse mussels but covered in cockle shells, patches of seaweed and crab burrows, C100 was smallest (0.0055), but still twice as large as the value typically applied to abiotic, flat, cohesionless seabeds (0.0025). The mean drag coefficient plus-or-minus standard error at the three sites with horse mussels was: 0.0082 ± 0.0010 (site 1); 0.0096 ± 0.0009 (site 2); 0.0115 ± 0.0016 (site 3). There were no clear differences amongst sites 1, 2 and 3 in terms of the attributes of individual horse mussels (e.g. shell height, width or orientation), which could have been used to explain the ranking of the drag coefficients. There were, however, differences amongst the three sites in terms of spatial distribution of individual bivalves. The site with the highest density of horse mussels, site 1, had the lowest drag coefficient and an areal concentration (λ) of horse mussels higher than typical values cited for the critical concentration (λc) for the onset of skimming flow over various idealized, three-dimensional roughness elements. At sites 2 and 3, the drag coefficient was given by: C100=[κ/1n (300/mkλ)]2 which was valid for λ \u3c λ c, where κ is von Karman\u27s constant, k is the horse mussel height (i.e., protrusion above the seabed), m ≈ 100 and λc ≈ 0.2. The stable eddies that are hypothesized to lodge between roughness elements at concentrations greater than λc may influence benthic community dynamics

Intertidal habitat mapping for ecosystem goods and services: Tairua harbour

In January 2013, Waikato Regional Council (WRC) contracted the National Institute of Water and Atmospheric Research (NIWA), to develop rapid assessment techniques for mapping of intertidal habitats associated with the provision of ecosystem goods and services. Ecosystem goods and services are defined as ‘the direct and indirect benefits that humankind receives or values from natural or semi-natural habitats’ and include the provision of food and raw materials, waste treatment, processing and storage, disturbance prevention, sediment retention, water filtration and regulation, nutrient regulation, gas and climate regulation, habitat structure and cultural services such as spiritual heritage and leisure and recreation (Townsend et al. 2010). Assessment techniques were to be trialled and implemented for the intertidal area of the Tairua estuary with the view of providing: • Descriptions of habitat types that may be linked to levels of ecosystem goods and services. • Descriptions of the techniques involved in differentiating and mapping habitats, and an analysis of the precision and accuracy/validity of the methods. This report documents the evolution of the methodology and a proof of concept using Tairua estuary as a test case to build a habitat map. The goal is to generate precursor maps that will facilitate the mapping of ecosystem goods and services in the near future

Species and functional trait turnover in response to broad-scale change and an invasive species

While beta diversity has been implicated as a key factor in controlling resilience of communities to stressors, lack of long-term data sets has limited the study of temporal dynamics of beta diversity. With a time series at two sites in excess of 40yr, we investigated turnover of both species and functional traits in a system stressed by eutrophication and overfishing and undergoing climate change and invasion. The two sites, although located near to each other, differ in water depth (20 cf. 35m), but both sites have displayed increased abundances of an invasive polychaete since 1990. We tested two hypotheses related to the effect of an invasive species; that taxa richness and turnover would decrease, and trait richness would increase post invasion and that trait turnover would increase between arrival and establishment of the invasive. Generally, we observed different dynamics at the two sites and responses not consistent with our hypotheses. We detected an increase in taxa richness at both sites and an increase in taxa turnover and number of traits at one site only. Trait turnover was higher prior to the invasion, although again only at one site. Disjunctive responses between species and trait turnover occurred, with the invader contributing in a nonrandom fashion to trait turnover. The lack of strong, consistent responses to the arrival and establishment of the invasive, and the decrease in trait turnover, suggests that effects of invasives are not only system- and species-dependent, but also depend on community dynamics of the invaded site, in particular the assembly processes, and historical context.Peer reviewe

Template for using biological trait groupings when exploring large-scale variation in seafloor multifunctionality

Understanding large-scale spatial variation in ecosystem properties and associated functionality is key for successful conservation of ecosystems. This study provides a template for how to estimate differences in ecosystem functionality over large spatial scales by using groupings of biological traits. We focus on trait groupings that describe three important benthic ecosystem properties, namely bioturbation, community stability, and juvenile dispersal. Recognizing that groups of traits interact and are constrained within an organism, we statistically define important functional trait subgroups that describe each ecosystem property. The sub-groups are scored according to their weighted ecological impact to gain an overall estimation of the cumulative expression of each ecosystem property at individual sites. Furthermore, by assigning each property a value relative to its observed maximum, and by summing up the individual property values, we offer an estimate of benthic ecosystem multifunctionality. Based on a spatially extensive benthic data set, we were able to identify coastal areas with high and low potential for the considered benthic ecosystem properties and the measure of ecosystem multifunctionality. Importantly, we show that a large part of the spatial variation in functional trait sub-groups and in benthic ecosystem multifunctionality was explained by environmental change. Our results indicate that through this simplification it is possible to estimate the functionality of the seafloor. Such information is vital in marine spatial planning efforts striving to balance the utilization with the preservation of natural resources.Peer reviewe

ANALISIS PEMAHAMAN KONSEP MATEMATIKA SISWA DENGAN MODEL PEMBELAJARAN PROBLEM BASED LEARNING PADA POKOK BAHASAN SEGITIGA (Study deskriptif di Kelas VII SMP Negeri 2 Plered Kab. Cirebon)

Anggun Maya Sari. NIM 59451058. “Analisis Pemahaman Konsep Matematika Siswa dengan Model Pembelajaran Problem Based Learning pada Pokok Bahasan Segitiga”. (Studi Deskriptif di Kelas VII SMP Negeri 2 Plered). Pembelajaran yang efektif adalah pembelajaran yang menyediakan kesempatan kepada siswa untuk belajar mandiri, sehingga dalam prosesnya siswa dapat memperoleh pemahaman dan pengetahuan dengan lebih mendalam. Akan tetapi proses kegiatan pembelajaran matematika yang berlangsung disekolah, pada umumnya guru hanya sekedar penyampai informasi tanpa mempertimbangkan seberapa jauh pemahaman siswa terhadap konsep matematika dari pokok bahasan segitiga yang disampaikan. Tujuan penelitian ini adalah 1) mengetahui seberapa besar pemahaman konsep yang dimiliki peserta didik; 2) mengetahui factor pendukung dan penghambat peserta didik dalam memahami konsep yang diberikan; 3) mengetahui deskripsi aktivitas siswa dalam memahami konsep matematika melalui model pembelajaran Problem Based Learning; 4) mengetahui respon siswa setelah melaksanakan model pembelajaran Problem Based Learning; 5) mengetahui sejauhmana pemahaman konsep matematika siswa setelah mengikuti proses pembelajaran Problem Based Learning. Penerapan model pembelajaran Problem Based Learning (PBL) merupakan salah satu model pembelajaran yang dapat memberikan kondisi belajar aktif kepada siswa. Dengan diterapkannya pembelajaran matematika dengan model pembelajaran PBL, diharapkan dapat membantu siswa dalam memahami konsep matematika. Penelitian ini menggunakan pendekatan kualitatif dan kuantitatif dengan metode deskriptif. Pengumpulan data menggunakan lembar observasi, pedoman wawancara, angket dan tes. Informan dalam penelitian ini adalah kelas VII yang sudah dibentuk kelompok kelas, yaitu VII A, VII D dan VII F. dengan menggunakan proportionate stratified random sampling, maka dari masingmasing kelompok kelas didapat 18 siswa sebagai informan penelitian. Berdasarkan dari hasil wawancara tentang siswa dalam pemahaman konsep matematika, diperoleh siswa masih kurang dan masih perlu banyak bimbingan. Aktivitas siswa selama pembelajaran melalui model pembelajaran PBL diperoleh dari hasil lembar observasi dengan nilai rata-rata dari semua aspek sebesar 54,39% yang termasuk dalam kategori sedang. Berdasarkan angket respon siswa setelah dilaksanakan model pembelajaran PBL termasuk dalam kategori baik dengan nilai rata-rata 76,44%. Pemahaman konsep matematika siswa setelah mengikuti proses pembelajaran dengan model pembelajaran PBL diperoleh siswa lebih rajin dalam mencari bahan untuk menyelesaikan soal-soal yang diberikan dan menambah pemahaman siswa mengenai konsep matematika, meski masih belum 100% benar, dan hasil tes didapat nilai rata-rata 39% yang termasuk dalam kategori sangat kurang. Kata Kunci: Pemahaman konsep matematika, Problem Based Learning, Segitig

The Challenge of Implementing the Marine Ecosystem Service Concept

The concept of ecosystem services has gained traction as a means of linking societal benefits to the underlying ecology and functioning of ecosystems, and is now frequently included in decision-making and legislation. Moving the ecosystem service concept from theory into practice is now crucial. However, advancements in this area of research differ by ecosystem type, and marine systems lag significantly behind terrestrial counterparts in terms of understanding, implementation, and number of studies. In this paper we explore several reasons why ecosystem service research has been limited in marine systems and we outline the challenges that hinder progress. Marine systems suffer from a scarcity of spatial data relative to terrestrial counterparts. In terrestrial systems the spatial patterns of land-use/land-cover (LULC) are relatively straightforward to access via satellite and have been used as proxy indicators of service provisions. In contrast, remote sensing tools used to study the surface of the Earth are much less effective at capturing images of the seabed, and by extension marine habitats. Marine waters and their constituents are also frequently driven great distances by winds, tides, and currents. This creates a challenge for management as the identification and protection of areas where ecosystem services are exploited is not necessarily sufficient to ensure sustained service delivery. Further complications arise from the three-dimensional uses of marine systems, incorporating activities that use the sea surface, the water column and the benthic habitats below. Progress is being made as technological advancements are resulting in the acquisition of spatial data at faster rates and higher resolutions than previously possible. There is a growing capacity to map, model and value an increasing number of services with initiatives such as InVEST or principle-based modeling. We suggest that awareness is needed around the limited progress in marine systems as this could affect the way we value the biosphere and the relative proportion between biomes

Assessing Benthic Responses to Fishing Disturbance Over Broad Spatial Scales That Incorporate High Environmental Variation

Marine benthic habitats are modified by a number of human-related disturbances. When these disturbances occur at large scales over areas of high environmental variability, it is difficult to assess impacts using metrics such as species richness or individual species distributions because of varying species-specific responses to environmental drivers (e.g., exposure, sediment, temperature). Impact assessment can also be problematic when assessed at broad spatial scales because of regional heterogeneity of species pools. Even when effects on individual species can be detected, it is difficult to upscale from individual species to ecosystem scale effects. Here, we use a functional group approach to assess broad scale patterns in ecological processes with respect to fishing and environmental drivers. We used data from field surveys of benthic communities from two large, widely separated areas in New Zealand’s EEZ (Chatham Rise and Challenger Plateau). We assigned 828 taxonomic units (most identified to species) into functional groups related to important ecosystem processes and likely sensitivity to, and recovery from, fishing disturbance to the seafloor. These included: opportunistic early colonists; substrate stabilisers (e.g., tube mat formers); substrate destabilisers; shell hash-creating species; emergent epifauna; burrowers; and predators and scavengers. Effects of fishing disturbance on benthic functional composition were observed, even at this broad spatial scale. Responses varied between functional groups, with some being tolerant of fishing impacts and others showing rapid declines with minimal fishing effort. The use of a functional group approach facilitates assessment of impacts across regions and species, allowing for improved generalisations of impacts to inform management and decision making

Consequences of increasing hypoxic disturbance on benthic communities and ecosystem functioning

Disturbance-mediated species loss has prompted research considering how ecosystem functions are changed when biota is impaired. However, there is still limited empirical evidence from natural environments evaluating the direct and indirect (i.e. via biota) effects of disturbance on ecosystem functioning. Oxygen deficiency is a widespread threat to coastal and estuarine communities. While the negative impacts of hypoxia on benthic communities are well known, few studies have assessed in situ how benthic communities subjected to different degrees of hypoxic stress alter their contribution to ecosystem functioning. We studied changes in sediment ecosystem function (i.e. oxygen and nutrient fluxes across the sediment water-interface) by artificially inducing hypoxia of different durations (0, 3, 7 and 48 days) in a subtidal sandy habitat. Benthic chamber incubations were used for measuring responses in sediment oxygen and nutrient fluxes. Changes in benthic species richness, structure and traits were quantified, while stress-induced behavioral changes were documented by observing bivalve reburial rates. The initial change in faunal behavior was followed by non-linear degradation in benthic parameters (abundance, biomass, bioturbation potential), gradually impairing the structural and functional composition of the benthic community. In terms of ecosystem function, the increasing duration of hypoxia altered sediment oxygen consumption and enhanced sediment effluxes of NH4 + and dissolved Si. Although effluxes of PO4 were not altered significantly, changes were observed in sediment PO4 sorption capability. The duration of hypoxia (i.e. number of days of stress) explained a minor part of the changes in ecosystem function. Instead, the benthic community and disturbancedriven changes within the benthos explained a larger proportion of the variability in sediment oxygen- and nutrient fluxes. Our results emphasize that the level of stress to the benthic habitat matters, and that the link between biodiversity and ecosystem function is likely to be affected by a range of factors in complex, natural environments.Peer reviewe

Identifying "vital attributes" for assessing disturbance-recovery potential of seafloor communities

Despite a long history of disturbance–recovery research, we still lack a generalizable understanding of the attributes that drive community recovery potential in seafloor ecosystems. Marine soft‐sediment ecosystems encompass a range of heterogeneity from simple low‐diversity habitats with limited biogenic structure, to species‐rich systems with complex biogenic habitat structure. These differences in biological heterogeneity are a product of natural conditions and disturbance regimes. To search for unifying attributes, we explore whether a set of simple traits can characterize community disturbance–recovery potential using seafloor patch‐disturbance experiments conducted in two different soft‐sediment landscapes. The two landscapes represent two ends of a spectrum of landscape biotic heterogeneity in order to consider multi‐scale disturbance–recovery processes. We consider traits at different levels of biological organization, from the biological traits of individual species, to the traits of species at the landscape scale associated with their occurrence across the landscape and their ability to be dominant. We show that in a biotically heterogeneous landscape (Kawau Bay, New Zealand), seafloor community recovery is stochastic, there is high species turnover, and the landscape‐scale traits are good predictors of recovery. In contrast, in a biotically homogeneous landscape (Baltic Sea), the options for recovery are constrained, the recovery pathway is thus more deterministic and the scale of recovery traits important for determining recovery switches to the individual species biological traits within the disturbed patch. Our results imply that these simple, yet sophisticated, traits can be effectively used to characterize community recovery potential and highlight the role of landscapes in providing resilience to patch‐scale disturbances.Peer reviewe