Cheating and the evolutionary stability of mutualisms

Interspecific mutualisms have been playing a central role in the functioning of all ecosystems since the early history of life. Yet the theory of coevolution of mutualists is virtually nonexistent, by contrast with well-developed coevolutionary theories of competition, predator–prey and host–parasite interactions. This has prevented resolution of a basic puzzle posed by mutualisms: their persistence in spite of apparent evolutionary instability. The selective advantage of 'cheating', that is, reaping mutualistic benefits while providing fewer commodities to the partner species, is commonly believed to erode a mutualistic interaction, leading to its dissolution or reciprocal extinction. However, recent empirical findings indicate that stable associations of mutualists and cheaters have existed over long evolutionary periods. Here, we show that asymmetrical competition within species for the commodities offered by mutualistic partners provides a simple and testable ecological mechanism that can account for the long-term persistence of mutualisms. Cheating, in effect, establishes a background against which better mutualists can display any competitive superiority. This can lead to the coexistence and divergence of mutualist and cheater phenotypes, as well as to the coexistence of ecologically similar, but unrelated mutualists and cheaters

Magnetic Flux Expulsion in the Powerful Superbubble Explosions and the Alpha-Omega Dynamo

The possibility of the magnetic flux expulsion from the Galaxy in the superbubble (SB) explosions, important for the Alpha-Omega dynamo, is considered. Special emphasis is put on the investigation of the downsliding of the matter from the top of the shell formed by the SB explosion which is able to influence the kinematics of the shell. It is shown that either Galactic gravity or the development of the Rayleigh-Taylor instabilities in the shell limit the SB expansion, thus, making impossible magnetic flux expulsion. The effect of the cosmic rays in the shell on the sliding is considered and it is shown that it is negligible compared to Galactic gravity. Thus, the question of possible mechanism of flux expulsion in the Alpha-Omega dynamo remains open.Comment: MNRAS, in press, 11 pages, 9 figure

Shocked Quartz in Polymict Impact Breccia from the Upper Cretaceous Yallalie Impact Structure in Western Australia

Yallalie is a ~12 km diameter circular structure located ~200 km north of Perth, Australia. Previous studies have proposed that the buried structure is a complex impact crater based on geophysical data. Allochthonous breccia exposed near the structure has previously been interpreted as proximal impact ejecta; however, no diagnostic indicators of shock metamorphism have been found. Here we report multiple (27) shocked quartz grains containing planar fractures (PFs) and planar deformation features (PDFs) in the breccia. The PFs occur in up to five sets per grain, while the PDFs occur in up to four sets per grain. Universal stage measurements of all 27 shocked quartz grains confirms that the planar microstructures occur in known crystallographic orientations in quartz corresponding to shock compression from 5 to 20 GPa. Proximity to the buried structure (~4 km) and occurrence of shocked quartz indicates that the breccia represents either primary or reworked ejecta. Ejecta distribution simulated using iSALE hydrocode predicts the same distribution of shock levels at the site as those found in the breccia, which supports a primary ejecta interpretation, although local reworking cannot be excluded. The Yallalie impact event is stratigraphically constrained to have occurred in the interval from 89.8 to 83.6 Ma based on the occurrence of Coniacian clasts in the breccia and undisturbed overlying Santonian to Campanian sedimentary rocks. Yallalie is thus the first confirmed Upper Cretaceous impact structure in Australia

The 511 keV emission from positron annihilation in the Galaxy

The first gamma-ray line originating from outside the solar system that was ever detected is the 511 keV emission from positron annihilation in the Galaxy. Despite 30 years of intense theoretical and observational investigation, the main sources of positrons have not been identified up to now. Observations in the 1990's with OSSE/CGRO showed that the emission is strongly concentrated towards the Galactic bulge. In the 2000's, the SPI instrument aboard ESA's INTEGRAL gamma-ray observatory allowed scientists to measure that emission across the entire Galaxy, revealing that the bulge/disk luminosity ratio is larger than observed in any other wavelength. This mapping prompted a number of novel explanations, including rather "exotic ones (e.g. dark matter annihilation). However, conventional astrophysical sources, like type Ia supernovae, microquasars or X-ray binaries, are still plausible candidates for a large fraction of the observed total 511 keV emission of the bulge. A closer study of the subject reveals new layers of complexity, since positrons may propagate far away from their production sites, making it difficult to infer the underlying source distribution from the observed map of 511 keV emission. However, contrary to the rather well understood propagation of high energy (>GeV) particles of Galactic cosmic rays, understanding the propagation of low energy (~MeV) positrons in the turbulent, magnetized interstellar medium, still remains a formidable challenge. We review the spectral and imaging properties of the observed 511 keV emission and we critically discuss candidate positron sources and models of positron propagation in the Galaxy.Comment: 62 pages, 35 figures. Review paper to appear in Reviews of Modern Physic

The Evolution of Adiabatic Supernova Remnants in a Turbulent, Magnetized Medium

(Abridged) We present the results of three dimensional calculations for the MHD evolution of an adiabatic supernova remnant in both a uniform and turbulent interstellar medium using the RIEMANN framework of Balsara. In the uniform case, which contains an initially uniform magnetic field, the density structure of the shell remains largely spherical, while the magnetic pressure and synchrotron emissivity are enhanced along the plane perpendicular to the field direction. This produces a bilateral or barrel-type morphology in synchrotron emission for certain viewing angles. We then consider a case with a turbulent external medium as in Balsara & Pouquet, characterized by $v_{A}(rms)/c_{s}=2$. Several important changes are found. First, despite the presence of a uniform field, the overall synchrotron emissivity becomes approximately spherically symmetric, on the whole, but is extremely patchy and time-variable, with flickering on the order of a few computational time steps. We suggest that the time and spatial variability of emission in early phase SNR evolution provides information on the turbulent medium surrounding the remnant. The shock-turbulence interaction is also shown to be a strong source of helicity-generation and, therefore, has important consequences for magnetic field generation. We compare our calculations to the Sedov-phase evolution, and discuss how the emission characteristics of SNR may provide a diagnostic on the nature of turbulence in the pre-supernova environment.Comment: ApJ, in press, 5 color figure

Evidence for dark matter in the inner Milky Way

The ubiquitous presence of dark matter in the universe is today a central tenet in modern cosmology and astrophysics. Ranging from the smallest galaxies to the observable universe, the evidence for dark matter is compelling in dwarfs, spiral galaxies, galaxy clusters as well as at cosmological scales. However, it has been historically difficult to pin down the dark matter contribution to the total mass density in the Milky Way, particularly in the innermost regions of the Galaxy and in the solar neighbourhood. Here we present an up-to-date compilation of Milky Way rotation curve measurements, and compare it with state-of-the-art baryonic mass distribution models. We show that current data strongly disfavour baryons as the sole contribution to the galactic mass budget, even inside the solar circle. Our findings demonstrate the existence of dark matter in the inner Galaxy while making no assumptions on its distribution. We anticipate that this result will compel new model-independent constraints on the dark matter local density and profile, thus reducing uncertainties on direct and indirect dark matter searches, and will shed new light on the structure and evolution of the Galaxy.Comment: First submitted version of letter published in Nature Physics on Febuary 9, 2015: http://www.nature.com/nphys/journal/vaop/ncurrent/full/nphys3237.htm

Magnetogenesis from Cosmic String Loops

Large-scale coherent magnetic fields are observed in galaxies and clusters, but their ultimate origin remains a mystery. We reconsider the prospects for primordial magnetogenesis by a cosmic string network. We show that the magnetic flux produced by long strings has been overestimated in the past, and give improved estimates. We also compute the fields created by the loop population, and find that it gives the dominant contribution to the total magnetic field strength on present-day galactic scales. We present numerical results obtained by evolving semi-analytic models of string networks (including both one-scale and velocity-dependent one-scale models) in a Lambda-CDM cosmology, including the forces and torques on loops from Hubble redshifting, dynamical friction, and gravitational wave emission. Our predictions include the magnetic field strength as a function of correlation length, as well as the volume covered by magnetic fields. We conclude that string networks could account for magnetic fields on galactic scales, but only if coupled with an efficient dynamo amplification mechanism.Comment: 10 figures; v3: small typos corrected to match published version. MagnetiCS, the code described in paper, is available at http://markcwyman.com/ and http://www.damtp.cam.ac.uk/user/dhw22/code/index.htm

The supernova rate-velocity dispersion relation in the interstellar medium

We investigate the relationship between the velocity dispersion of the gas and the SN rate and feedback efficiency in the ISM. We explore the constancy of the velocity dispersion profiles in the outer parts of galactic disks at~6-8 km s^-1, and the transition to the starburst regime. Our results show that a) SN driving leads to constant velocity dispersions of sig~6 km s^-1 for the total gas and sigHI~3 km s^-1 for the HI gas, independent of the SN rate, for values of the rate between 0.01-0.5 the Galactic rate R_{G},b) the position of the transition to the starburst regime at SFR/Area~5*10^-3-10^-2 M_sol yr^-1 kpc^-2 observed in the simulations, is in good agreement with the transition to the starburst regime in the observations, c) for the high SN rates, no HI gas is present in the simulations box, however, for the total gas velocity dispersion, there is good agreement between the models and the observations,d) at the intermediate SN rates R/R_{G}~0.5-1, taking into account the thermal broadening of the HI line helps reach a good agreement in that regime between the models and the observations,e) for R/R_{G}<0.5, sig and sigHI fall below the observed values by a factor of~2. However, a set of simulation with different values of epsilon indicates that for larger values of the supernova feedback efficiencies, velocity dispersions of the HI gas of the order of 5-6 km s^{-1} can be obtained, in closer agreement with the observations. The fact that for R/R_{G}<0.5, the HI gas velocity dispersions are a factor ~2 smaller than the observed values could result from the fact that we might have underestimated the SN feedback efficiency. It might also be an indication that other physical processes couple to the stellar feedback in order to produce the observed level of turbulence in galactic disks.Comment: 44 pages, 22 figures. Accepted to Ap

A Broadband Study of Galactic Dust Emission

We have combined infrared data with HI, H2 and HII surveys in order to spatially decompose the observed dust emission into components associated with different phases of the gas. An inversion technique is applied. For the decomposition, we use the IRAS 60 and 100 micron bands, the DIRBE 140 and 240 micron bands, as well as Archeops 850 and 2096 micron wavelengths. In addition, we apply the decomposition to all five WMAP bands. We obtain longitude and latitude profiles for each wavelength and for each gas component in carefully selected Galactic radius bins.We also derive emissivity coefficients for dust in atomic, molecular and ionized gas in each of the bins.The HI emissivity appears to decrease with increasing Galactic radius indicating that dust associated with atomic gas is heated by the ambient interstellar radiation field (ISRF). By contrast, we find evidence that dust mixed with molecular clouds is significantly heated by O/B stars still embedded in their progenitor clouds. By assuming a modified black-body with emissivity law lambda^(-1.5), we also derive the radial distribution of temperature for each phase of the gas. All of the WMAP bands except W appear to be dominated by emission from something other than normal dust, most likely a mixture of thermal bremstrahlung from diffuse ionized gas, synchrotron emission and spinning dust. Furthermore, we find indications of an emissivity excess at long wavelengths (lambda > 850 micron) in the outer Galaxy (R > 8.9 kpc). This suggests either the existence of a very cold dust component in the outer Galaxy or a temperature dependence of the spectral emissivity index. Finally, it is shown that ~ 80% of the total FIR luminosity is produced by dust associated with atomic hydrogen, in agreement with earlier findings by Sodroski et al. (1997).Comment: accepted for publication by A&