Hydrogen and Metal Line Absorption Around Low-Redshift Galaxies in Cosmological Hydrodynamic Simulations

We study the physical conditions of the circum-galactic medium (CGM) around z=0.25 galaxies as traced by HI and metal line absorption, using cosmological hydrodynamic simulations that include galactic outflows. Using lines of sight targeted at impact parameters from 10 kpc to 1 Mpc around galaxies with halo masses from 10^11-10^13 M_solar, we study the physical conditions and their variation with impact parameter b and line-of-sight velocity delta v in the CGM as traced by HI, MgII, SiIV, CIV, OVI, and NeVIII absorbers. All ions show a strong excess of absorption near galaxies compared to random lines of sight. The excess continues beyond 1 Mpc, reflecting the correlation of metal absorption with large-scale structure. Absorption is particularly enhanced within about v<300 km/sec and roughly 300 kpc of galaxies (with distances somewhat larger for the highest ion), approximately delineating the CGM; this range contains the majority of global metal absorption. Low ions like MgII and SiIV predominantly arise in denser gas closer to galaxies and drop more rapidly with b, while high ions OVI and NeVIII trace more diffusely distributed gas with a comparatively flat radial profile; CIV is intermediate. All ions predominantly trace T~10^4-4.5 K photo-ionised gas at all b, but when hot CGM gas is present (mostly in larger halos), we see strong collisionally-ionised OVI and NeVIII at b <= 100 kpc. Larger halo masses generally produce more absorption, though overall the trends are not as strong as that with impact parameter. These findings arise using our favoured outflow scalings as expected for momentum-driven winds; with no winds, the CGM gas remains mostly unenriched, while our outflow model with a constant velocity and mass loading factor produce hotter, more widely dispersed metals.Comment: 26 pages, 15 figures, published in MNRAS. Updates to citations from previous versio

Feedback and Recycled Wind Accretion: Assembling the z=0 Galaxy Mass Function

We analyse cosmological hydrodynamic simulations that include observationally-constrained prescriptions for galactic outflows. If these simulated winds accurately represent winds in the real Universe, then material previously ejected in winds provides the dominant source of gas infall for new star formation at redshifts z<1. This recycled wind accretion, or wind mode, provides a third physically distinct accretion channel in addition to the "hot" and "cold" modes emphasised in recent theoretical studies. Because of the interaction between outflows and gas in and around halos, the recycling timescale of wind material (t_rec) is shorter in higher-mass systems, which reside in denser gaseous environments. In these simulations, this differential recycling plays a central role in shaping the present-day galaxy stellar mass function (GSMF). If we remove all particles that were ever ejected in a wind, then the predicted GSMFs are much steeper than observed; galaxy masses are suppressed both by the direct removal of gas and by the hydrodynamic heating of their surroundings, which reduces subsequent infall. With wind recycling included, the simulation that incorporates our favoured momentum-driven wind scalings reproduces the observed GSMF for stellar masses 10^9 < M < 5x10^10 Msolar. At higher masses, wind recycling leads to excessive galaxy masses and excessive star formation rates relative to observations. In these massive systems, some quenching mechanism must suppress the re-accretion of gas ejected from star-forming galaxies. In short, as has long been anticipated, the form of the GSMF is governed by outflows; the unexpected twist here for our simulated winds is that it is not primarily the ejection of material but how the ejected material is re-accreted that governs the GSMF.Comment: 16 pages, 7 figures, accepted by MNRA

Compositional diversity of rehabilitated tropical lands supports multiple ecosystem services and buffers uncertainties

High landscape diversity is assumed to increase the number and level of ecosystem services. However, the interactions between ecosystem service provision, disturbance and landscape composition are poorly understood. Here we present a novel approach to include uncertainty in the optimization of land allocation for improving the provision of multiple ecosystem services. We refer to the rehabilitation of abandoned agricultural lands in Ecuador including two types of both afforestation and pasture rehabilitation, together with a succession option. Our results show that high compositional landscape diversity supports multiple ecosystem services (multifunction effect). This implicitly provides a buffer against uncertainty. Our work shows that active integration of uncertainty is only important when optimizing single or highly correlated ecosystem services and that the multifunction effect on landscape diversity is stronger than the uncertainty effect. This is an important insight to support a land-use planning based on ecosystem services

Chandra survey of nearby highly inclined disc galaxies - III. Comparison with hydrodynamical simulations of circumgalactic coronae

X-ray observations of circumgalactic coronae provide a valuable means by which to test galaxy formation theories. Two primary mechanisms are thought to be responsible for the establishment of such coronae: accretion of intergalactic gas and/or galactic feedback. In this paper, we first compare our Chandra sample of galactic coronae of 53 nearby highly-inclined disc galaxies to an analytical model considering only the accretion of intergalactic gas. We confirm the existing conclusion that this pure accretion model substantially over-predicts the coronal emission. We then select 30 field galaxies from our original sample, and correct their coronal luminosities to uniformly compare them to deep X-ray measurements of several massive disc galaxies from the literature, as well as to a comparable sample of simulated galaxies drawn from the Galaxies-Intergalactic Medium Interaction Calculation (GIMIC). These simulations explicitly model both accretion and supernovae feedback and yield galaxies that exhibit X-ray properties in broad agreement with our observational sample. However, notable and potentially instructive discrepancies exist between the slope and scatter of the LX −M200 and LX − SFR relations, highlighting some known shortcomings of GIMIC, for example, the absence of AGN feedback, and possibly the adoption of constant stellar feedback parameters. The simulated galaxies exhibit a tight correlation (with little scatter) between coronal luminosity and halo mass. Having inferredM200 for our observational sample via the Tully-Fisher relation, we find a weaker and more scattered correlation. In the simulated and observed samples alike, massive non-starburst galaxies above a typical transition mass of M∗ � 2×1011 M⊙ or M200 � 1013 M⊙ tend to have higher LX/M∗ and LX/M200 than low-mass counterparts, indicating that the accretion of intergalactic gas plays an increasingly important role in establishing the observable hot circumgalactic medium with increasing galaxy mass. Subject headings: galaxies: general—galaxies: halos—galaxies: normal—X-rays: galaxie

Intraspecific Correlations of Basal and Maximal Metabolic Rates in Birds and the Aerobic Capacity Model for the Evolution of Endothermy

The underlying assumption of the aerobic capacity model for the evolution of endothermy is that basal (BMR) and maximal aerobic metabolic rates are phenotypically linked. However, because BMR is largely a function of central organs whereas maximal metabolic output is largely a function of skeletal muscles, the mechanistic underpinnings for their linkage are not obvious. Interspecific studies in birds generally support a phenotypic correlation between BMR and maximal metabolic output. If the aerobic capacity model is valid, these phenotypic correlations should also extend to intraspecific comparisons. We measured BMR, Msum (maximum thermoregulatory metabolic rate) and MMR (maximum exercise metabolic rate in a hop-flutter chamber) in winter for dark-eyed juncos (Junco hyemalis), American goldfinches (Carduelis tristis; Msum and MMR only), and black-capped chickadees (Poecile atricapillus; BMR and Msum only) and examined correlations among these variables. We also measured BMR and Msum in individual house sparrows (Passer domesticus) in both summer, winter and spring. For both raw metabolic rates and residuals from allometric regressions, BMR was not significantly correlated with either Msum or MMR in juncos. Moreover, no significant correlation between Msum and MMR or their mass-independent residuals occurred for juncos or goldfinches. Raw BMR and Msum were significantly positively correlated for black-capped chickadees and house sparrows, but mass-independent residuals of BMR and Msum were not. These data suggest that central organ and exercise organ metabolic levels are not inextricably linked and that muscular capacities for exercise and shivering do not necessarily vary in tandem in individual birds. Why intraspecific and interspecific avian studies show differing results and the significance of these differences to the aerobic capacity model are unknown, and resolution of these questions will require additional studies of potential mechanistic links between minimal and maximal metabolic output

Group Decisions in Biodiversity Conservation: Implications from Game Theory

. This paper shows how game theory may be used to inform group decisions in biodiversity conservation scenarios by modeling conflicts between stakeholders to identify Pareto–inefficient Nash equilibria. These are cases in which each agent pursuing individual self–interest leads to a worse outcome for all, relative to other feasible outcomes. Three case studies from biodiversity conservation contexts showing this feature are modeled to demonstrate how game–theoretical representation can inform group decision-making.–agent fish and coral conservation scenario from the Philippines. In each case there is reason to believe that traditional mechanism–design solutions that appeal to material incentives may be inadequate, and the game–theoretical analysis recommends a resumption of further deliberation between agents and the initiation of trust—and confidence—building measures. that formal mechanism–design solutions may backfire in certain cases. Such scenarios demand a return to group deliberation and the creation of reciprocal relationships of trust

The Role of Sialyl Glycan Recognition in Host Tissue Tropism of the Avian Parasite Eimeria tenella

Eimeria spp. are a highly successful group of intracellular protozoan parasites that develop within intestinal epithelial cells of poultry, causing coccidiosis. As a result of resistance against anticoccidial drugs and the expense of manufacturing live vaccines, it is necessary to understand the relationship between Eimeria and its host more deeply, with a view to developing recombinant vaccines. Eimeria possesses a family of microneme lectins (MICs) that contain microneme adhesive repeat regions (MARR). We show that the major MARR protein from Eimeria tenella, EtMIC3, is deployed at the parasite-host interface during the early stages of invasion. EtMIC3 consists of seven tandem MAR1-type domains, which possess a high specificity for sialylated glycans as shown by cell-based assays and carbohydrate microarray analyses. The restricted tissue staining pattern observed for EtMIC3 in the chicken caecal epithelium indicates that EtMIC3 contributes to guiding the parasite to the site of invasion in the chicken gut. The microarray analyses also reveal a lack of recognition of glycan sequences terminating in the N-glycolyl form of sialic acid by EtMIC3. Thus the parasite is well adapted to the avian host which lacks N-glycolyl neuraminic acid. We provide new structural insight into the MAR1 family of domains and reveal the atomic resolution basis for the sialic acid-based carbohydrate recognition. Finally, a preliminary chicken immunization trial provides evidence that recombinant EtMIC3 protein and EtMIC3 DNA are effective vaccine candidates