566 research outputs found

    Evolution of flowering strategies in Oenothera glazioviana: an integral projection model approach

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    The timing of reproduction is a key determinant of fitness. Here, we develop parameterized integral projection models of size-related flowering for the monocarpic perennial Oenothera glazioviana and use these to predict the evolutionarily stable strategy (ESS) for flowering. For the most part there is excellent agreement between the model predictions and the results of quantitative field studies. However, the model predicts a much steeper relationship between plant size and the probability of flowering than observed in the field, indicating selection for a 'threshold size' flowering function. Elasticity and sensitivity analysis of population growth rate u and net reproductive rate R0 are used to identify the critical traits that determine fitness and control the ESS for flowering. Using the fitted model we calculate the fitness landscape for invading genotypes and show that this is characterized by a ridge of approximately equal fitness. The implications of these results for the maintenance of genetic variation are discussed

    Evolution of size-dependent flowering in a variable environment: partitioning the effects of fluctuating selection

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    In a stochastic environment, two distinct processes, namely nonlinear averaging and non-equilibrium dynamics, influence fitness. We develop methods for decomposing the effects of temporal variation in demography into contributions from nonlinear averaging and non-equilibrium dynamics. We illustrate the approach using Carlina vulgaris, a monocarpic species in which recruitment, growth and survival all vary from year to year. In Carlina the absolute effect of temporal variation on the evolutionarily stable flowering strategy is substantial (ca. 50% of the evolutionarily stable flowering size) but the net effect is much smaller (ca. 10%) because the effects of temporal variation do not influence the evolutionarily stable strategy in the same direction

    Evolution of complex flowering strategies: an age- and size-structured integral projection model

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    We explore the evolution of delayed age- and size-dependent flowering in the monocarpic perennial Carlina vulgaris, by extending the recently developed integral projection approach to include demographic rates that depend on size and age. The parameterized model has excellent descriptive properties both in terms of the population size and in terms of the distributions of sizes within each age class. In Carlina the probability of flowering depends on both plant size and age. We use the parameterized model to predict this relationship, using the evolutionarily stable strategy (ESS) approach. Despite accurately predicting the mean size of flowering individuals, the model predicts a step-function relationship between the probability of flowering and plant size, which has no age component. When the variance of the flowering-threshold distribution is constrained to the observed value, the ESS flowering function contains an age component, but underpredicts the mean flowering size. An analytical approximation is used to explore the effect of variation in the flowering strategy on the ESS predictions. Elasticity analysis is used to partition the agespecific contributions to the finite rate of increase (u) of the survival-growth and fecundity components of the model. We calculate the adaptive landscape that defines the ESS and generate a fitness landscape for invading phenotypes in the presence of the observed flowering strategy. The implications of these results for the patterns of genetic diversity in the flowering strategy and for testing evolutionary models are discussed. Results proving the existence of a dominant eigenvalue and its associated eigenvectors in general size- and age-dependent integral projection models are presented

    Demonstration of the temporal matter-wave Talbot effect for trapped matter waves

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    We demonstrate the temporal Talbot effect for trapped matter waves using ultracold atoms in an optical lattice. We investigate the phase evolution of an array of essentially non-interacting matter waves and observe matter-wave collapse and revival in the form of a Talbot interference pattern. By using long expansion times, we image momentum space with sub-recoil resolution, allowing us to observe fractional Talbot fringes up to 10th order.Comment: 17 pages, 7 figure

    Longitudinal double-spin asymmetry and cross section for inclusive neutral pion production at midrapidity in polarized proton collisions at sqrt(s) = 200 GeV

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    We report a measurement of the longitudinal double-spin asymmetry A_LL and the differential cross section for inclusive Pi0 production at midrapidity in polarized proton collisions at sqrt(s) = 200 GeV. The cross section was measured over a transverse momentum range of 1 < p_T < 17 GeV/c and found to be in good agreement with a next-to-leading order perturbative QCD calculation. The longitudinal double-spin asymmetry was measured in the range of 3.7 < p_T < 11 GeV/c and excludes a maximal positive gluon polarization in the proton. The mean transverse momentum fraction of Pi0's in their parent jets was found to be around 0.7 for electromagnetically triggered events.Comment: 6 pages, 3 figures, submitted to Phys. Rev. D (RC

    Kaon Production and Kaon to Pion Ratio in Au+Au Collisions at \snn=130 GeV

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    Mid-rapidity transverse mass spectra and multiplicity densities of charged and neutral kaons are reported for Au+Au collisions at \snn=130 GeV at RHIC. The spectra are exponential in transverse mass, with an inverse slope of about 280 MeV in central collisions. The multiplicity densities for these particles scale with the negative hadron pseudo-rapidity density. The charged kaon to pion ratios are K+/π=0.161±0.002(stat)±0.024(syst)K^+/\pi^- = 0.161 \pm 0.002 {\rm (stat)} \pm 0.024 {\rm (syst)} and K/π=0.146±0.002(stat)±0.022(syst)K^-/\pi^- = 0.146 \pm 0.002 {\rm (stat)} \pm 0.022 {\rm (syst)} for the most central collisions. The K+/πK^+/\pi^- ratio is lower than the same ratio observed at the SPS while the K/πK^-/\pi^- is higher than the SPS result. Both ratios are enhanced by about 50% relative to p+p and pˉ\bar{\rm p}+p collision data at similar energies.Comment: 6 pages, 3 figures, 1 tabl

    Azimuthal anisotropy and correlations in p+p, d+Au and Au+Au collisions at 200 GeV

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    We present the first measurement of directed flow (v1v_1) at RHIC. v1v_1 is found to be consistent with zero at pseudorapidities η\eta from -1.2 to 1.2, then rises to the level of a couple of percent over the range 2.4<η<42.4 < |\eta| < 4. The latter observation is similar to data from NA49 if the SPS rapidities are shifted by the difference in beam rapidity between RHIC and SPS. Back-to-back jets emitted out-of-plane are found to be suppressed more if compared to those emitted in-plane, which is consistent with {\it jet quenching}. Using the scalar product method, we systematically compared azimuthal correlations from p+p, d+Au and Au+Au collisions. Flow and non-flow from these three different collision systems are discussed.Comment: Quark Matter 2004 proceeding, 4 pages, 3 figure

    Measurement of the parity-violating longitudinal single-spin asymmetry for W±W^{\pm} boson production in polarized proton-proton collisions at s=500\sqrt{s} = 500 GeV

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    We report the first measurement of the parity violating single-spin asymmetries for midrapidity decay positrons and electrons from W+W^{+} and WW^{-} boson production in longitudinally polarized proton-proton collisions at s=500\sqrt{s}=500 GeV by the STAR experiment at RHIC. The measured asymmetries, ALW+=0.27±0.10  (stat.)±0.02  (syst.)±0.03  (norm.)A^{W^+}_{L}=-0.27\pm 0.10\;({\rm stat.})\pm 0.02\;({\rm syst.}) \pm 0.03\;({\rm norm.}) and ALW=0.14±0.19  (stat.)±0.02  (syst.)±0.01  (norm.)A^{W^-}_{L}=0.14\pm 0.19\;({\rm stat.})\pm 0.02 \;({\rm syst.})\pm 0.01\;({\rm norm.}), are consistent with theory predictions, which are large and of opposite sign. These predictions are based on polarized quark and antiquark distribution functions constrained by polarized DIS measurements.Comment: 6 pages, 4 figures, submitted to Physics Review Letter

    Azimuthal anisotropy: the higher harmonics

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    We report the first observations of the fourth harmonic (v_4) in the azimuthal distribution of particles at RHIC. The measurement was done taking advantage of the large elliptic flow generated at RHIC. The integrated v_4 is about a factor of 10 smaller than v_2. For the sixth (v_6) and eighth (v_8) harmonics upper limits on the magnitudes are reported.Comment: 4 pages, 6 figures, contribution to the Quark Matter 2004 proceeding
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