1,132 research outputs found

    Magnetohydrodynamic waves in solar partially ionized plasmas: two-fluid approach

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    We derive the dynamics of magnetohydrodynamic waves in two-fluid partially ionized plasmas and to compare the results with those obtained under single-fluid description. Two-fluid magnetohydrodynamic equations are used, where ion-electron plasma and neutral particles are considered as separate fluids. Dispersion relations of linear magnetohydrodynamic waves are derived for simplest case of homogeneous medium. Frequencies and damping rates of waves are obtained for different parameters of background plasma. We found that two- and single-fluid descriptions give similar results for low frequency waves. However, the dynamics of MHD waves in two-fluid approach is significantly changed when the wave frequency becomes comparable or higher than ion-neutral collision frequency. Alfven and fast magneto-acoustic waves attain their maximum damping rate at particular frequencies (for example, the peak frequency equals 2.5 ion-neutral collision frequency for 50 % of neutral Hydrogen) in wave spectrum. The damping rates are reduced for higher frequency waves. The new mode of slow magneto-acoustic wave appears for higher frequency branch, which is connected to neutral hydrogen fluid. The single-fluid approach perfectly deals with slow processes in partially ionized plasmas, but fails for time-scales smaller than ion-neutral collision time. Therefore, two-fluid approximation should be used for the description of relatively fast processes. Some results of single-fluid description, for example the damping of high-frequency Alfven waves in the solar chromosphere due to ion-neutral collisions, should be revised in future.Comment: 8 pages, 7 figures, accepted in A&

    Collisional dissipation of Alfvén waves in a partially ionised solar chromosphere

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    Certain regions of the solar atmosphere are at sufficiently low temperatures to be only partially ionised. The lower chromosphere contains neutral atoms, the existence of which greatly increases the efficiency of the damping of waves due to collisional friction momentum transfer. More specifically the Cowling conductivity can be up to 12 orders of magnitude smaller than the Spitzer value, so that the main damping mechanism in this region is due to the collisions between neutrals and positive ions (Khodachenko et al. 2004, A&A, 422, 1073). Using values for the gas density and temperature as functions of height taken from the VAL C model of the quiet Sun (Vernazza et al. 1981, ApJS, 45, 635), an estimate is made for the dependance of the Cowling conductivity on height and strength of magnetic field. Using both analytic and numerical approaches the passage of Alfvén waves over a wide spectrum through this partially ionised region is investigated. Estimates of the efficiency of this region in the damping of Alfvén waves are made and compared for both approaches. We find that Alfvén waves with frequencies above 0.6 Hz are completely damped and frequencies below 0.01 Hz unaffected

    Damping of Alfven waves in solar partially ionized plasmas: effect of neutral helium in multi-fluid approach

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    Chromospheric and prominence plasmas contain neutral atoms, which may change the plasma dynamics through collision with ions. Most of the atoms are neutral hydrogen, but a significant amount of neutral helium may also be present in the plasma with a particular temperature. Damping of MHD waves due to ion collision with neutral hydrogen is well studied, but the effects of neutral helium are largely unknown. We aim to study the effect of neutral helium in the damping of Alfven waves in solar partially ionized plasmas. We consider three-fluid magnetohydrodynamic (MHD) approximation, where one component is electron-proton-singly ionized helium and other two components are the neutral hydrogen and neutral helium atoms. We derive the dispersion relation of linear Alfven waves in isothermal and homogeneous plasma. Then we solve the dispersion relation and derive the damping rates of Alfven waves for different plasma parameters. The presence of neutral helium significantly enhances the damping of Alfven waves compared to the damping due to neutral hydrogen at certain values of plasma temperature (10000-40000 K) and ionization. Damping rates have a peak near the ion-neutral collision frequency, but decrease for the higher part of wave spectrum. Collision of ions with neutral helium atoms can be of importance for the damping of Alfven waves in chromospheric spicules and in prominence-corona transition regions.Comment: 7 pages, 4 figures, accepted in A&

    Escherichia coli TatA and TatB Proteins Have N-out, C-in Topology in Intact Cells

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    The twin arginine protein transport (Tat) system translocates folded proteins across the cytoplasmic membrane of prokaryotes and the thylakoid membrane of chloroplasts. In Escherichia coli, TatA, TatB, and TatC are essential components of the machinery. A complex of TatB and TatC acts as the substrate receptor, whereas TatA is proposed to form the Tat transport channel. TatA and TatB are related proteins that comprise an N-terminal transmembrane helix and an adjacent amphipathic helix. Previous studies addressing the topological organization of TatA have given conflicting results. In this study, we have addressed the topological arrangement of TatA and TatB in intact cells by labeling of engineered cysteine residues with the membrane-impermeable thiol reagent methoxypolyethylene glycol maleimide. Our results show that TatA and TatB share an N-out, C-in topology, with no evidence that the amphipathic helices of either protein are exposed at the periplasmic side of the membrane. We further show that the N-out, C-in topology of TatA is fixed and is not affected by the absence of other Tat components or by the overproduction of a Tat substrate. These data indicate that topological reorganization of TatA is unlikely to accompany Tat-dependent protein transport

    Staphylococcus aureus toxin LukSF dissociates from its membrane receptor target to enable renewed ligand sequestration

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    Staphylococcus aureus Panton-Valentine leukocidin is a pore-forming toxin targeting the human C5a receptor (hC5aR), enabling this pathogen to battle the immune response by destroying phagocytes through targeted lysis. The mechanisms that contribute to rapid cell lysis are largely unexplored. Here, we show that cell lysis may be enabled by a process of toxins targeting receptor clusters and present indirect evidence for receptor recycling that allows multiple toxin pores to be formed close together. With the use of live cell single-molecule super-resolution imaging, Forster resonance energy transfer and nanoscale total internal reflection fluorescence colocalization microscopy, we visualized toxin pore formation in the presence of its natural docking ligand. We demonstrate disassociation of hC5aR from toxin complexes and simultaneous binding of new ligands. This effect may free mobile receptors to amplify hyperinflammatory reactions in early stages of microbial infections and have implications for several other similar bicomponent toxins and the design of new antibiotics.Haapasalo, K., Wollman, A. J. M., de Haas, C. J. C., van Kessel, K. P. M., van Strijp, J. A. G., Leake, M. C. Staphylococcus aureus toxin LukSF dissociates from its membrane receptor target to enable renewed ligand sequestration.Peer reviewe
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