Where does the time go? Assessing the chronostratigraphic fidelity of sedimentary geological outcrops in the pliocene–pleistocene red crag formation, eastern england

It is widely understood that Earth’s stratigraphic record is an incomplete record of time, but the implications that this has for interpreting sedimentary outcrop has received little attention. Here we consider how time is preserved at outcrop using the Neogene-Quaternary Red Crag Formation, England. The Red Crag Formation hosts sedimentological and ichnological proxies that can be used to assess the time taken to accumulate outcrop expressions of strata, as ancient depositional environments fluctuated between states of deposition, erosion and stasis. We use these to estimate how much time is preserved at outcrop scale and find that every outcrop provides only a vanishingly small window onto unanchored weeks to months within the 600-800 ka of ‘Crag-time’. Much of the apparently missing time may be accounted for by the parts of the formation at subcrop, rather than outcrop: stratigraphic time has not been lost, but is hidden. The time-completeness of the Red Crag Formation at outcrop appears analogous to that recorded in much older rock units, implying that direct comparison between strata of all ages is valid and that perceived stratigraphic incompleteness is an inconsequential barrier to viewing the outcrop sedimentary-stratigraphic record as a truthful chronicle of Earth history

A graphic method for depicting horizontal direction data on vertical outcrop photographs

Outcrop photographs which show two-dimensional representations of three-dimensionally dipping surfaces (e.g., bedding planes, cross-bed foresets) are commonly utilized in the description of sedimentary strata. In many instances, accurate depiction of the dip direction of such features is paramount for understanding their interpretation, and for visualizing the true form of three-dimensional bodies (e.g., conceptualizing the form of an architectural element in a cliff-face, preserved as a vertical slice that has been cut oblique to paleocurrent direction). However, as an outcrop photograph often presents information on a vertical plane and directional data refers to a horizontal plane, the accurate co-depiction of both sets of information may be challenging. There is presently no universal method for illustrating such measurements on outcrop photographs: techniques in common usage are often imprecise, and the lack of uniformity hinders comparison between different images. Here we present a method for accurately depicting horizontal direction data on vertical outcrop photographs which permits instant visualization of dip relative to the illustrated outcrop geometry. The method is simple to apply, does not compromise primary data, and is unobtrusive to other visual information within images; thus having utility across a broad spectrum of geological investigations

EDIACARAN LIFE CLOSE to LAND: COASTAL and SHOREFACE HABITATS of the EDIACARAN MACROBIOTA, the CENTRAL FLINDERS RANGES, SOUTH AUSTRALIA

ABSTRACTThe Rawnsley Quartzite of South Australia hosts some of the world's most diverse Ediacaran macrofossil assemblages, with many of the constituent taxa interpreted as early representatives of metazoan clades. Globally, a link has been recognized between the taxonomic composition of individual Ediacaran bedding-plane assemblages and specific sedimentary facies. Thorough characterization of fossil-bearing facies is thus of fundamental importance for reconstructing the precise environments and ecosystems in which early animals thrived and radiated, and distinguishing between environmental and evolutionary controls on taxon distribution. This study refines the paleoenvironmental interpretations of the Rawnsley Quartzite (Ediacara Member and upper Rawnsley Quartzite). Our analysis suggests that previously inferred water depths for fossil-bearing facies are overestimations. In the central regions of the outcrop belt, rather than shelf and submarine canyon environments below maximum (storm-weather) wave base, and offshore environments between effective (fair-weather) and maximum wave base, the succession is interpreted to reflect the vertical superposition and lateral juxtaposition of unfossiliferous non-marine environments with fossil-bearing coastal and shoreface settings. Facies comprise: 1, 2) amalgamated channelized and cross-bedded sandstone (major and minor tidally influenced river and estuarine channels, respectively), 3) ripple cross-laminated heterolithic sandstone (intertidal mixed-flat), 4) silty-sandstone (possible lagoon), 5) planar-stratified sandstone (lower shoreface), 6) oscillation-ripple facies (middle shoreface), 7) multi-directed trough- and planar-cross-stratified sandstone (upper shoreface), 8) ripple cross-laminated, planar-stratified rippled sandstone (foreshore), 9) adhered sandstone (backshore), and 10) planar-stratified and cross-stratified sandstone with ripple cross-lamination (distributary channels). Surface trace fossils in the foreshore facies represent the earliest known evidence of mobile organisms in intermittently emergent environments. All facies containing fossils of the Ediacaran macrobiota remain definitively marine. Our revised shoreface and coastal framework creates greater overlap between this classic “White Sea” biotic assemblage and those of younger, relatively depauperate “Nama”-type biotic assemblages located in Namibia. Such overlap lends support to the possibility that the apparent biotic turnover between these assemblages may reflect a genuine evolutionary signal, rather than the environmental exclusion of particular taxa.NERC ERC Dr Schürmann Foundatio

High-energy flood events recorded in the Mesoproterozoic Meall Dearg Formation, NW Scotland; their recognition and implications for the study of pre-vegetation alluvium

The Mesoproterozoic-Neoproterozoic Torridonian Sandstone of NW Scotland has renewed sedimentological significance following recent advances in the understanding of pre-vegetation alluvium, as it is one of the most extensive and easily-accessible successions of such strata worldwide. This paper presents the first modern sedimentological analysis of the unit’s constituent Meall Dearg Formation (late Mesoproterozoic), recognizing a dominance of alluvial facies, with subordinate aeolian facies. Alluvial strata within the Meall Dearg Formation contain direct evidence for event deposition by high-energy floods, including: (1) widespread upper- and transitional-upper flow regime elements; (2) frequent stacking of successively-lower flow-regime elements; (3) common subcritical subaqueous dune fields with superimposed ripple marks; (4) occasional thin, desiccated mudstones; and (5) evidence that microbial mats colonized substrates during intervals of sedimentary stasis. Together these strands of primary sedimentary geological evidence indicate that the alluvial deposition of the Meall Dearg Formation was typified by supercritical flows during high-energy floods, punctuated by prolonged intervals of sedimentary stasis. The preservation potential of all of the features was boosted by highly aggradational sedimentary conditions. These primary observations have implications for the ‘norm’ in pre-vegetation alluvium and confirm that, despite increasing recognition of diversity within Precambrian fluvial systems, classical pre-vegetation motifs of high-energy alluvial flood deposits preserved as “sheet-braided” alluvium are still an archetypal sedimentary signature in some instances. Using supportive evidence from the Meall Dearg Formation, we recommend that the term “sheet-braided” be used in inverted commas in future studies; this emphasises the polygenetic depositional nature of the 20:1 width:thickness fluvial style, while maintaining the value of the term in the isolation of a key characteristic of pre-vegetation sedimentary architecture.Supported by Shell International Exploration and Production B.V under Research Framework agreement PT38181. Two anonymous reviewers and editor Adrian Hartley are thanked for comments which improved the manuscript

Evolutionary synchrony of Earth's biosphere and sedimentary-stratigraphic record

The landscapes and seascapes of Earth’s surface provide the theatre for life, but to what extent did the actors build the stage? The role of life in the long-term shaping of the planetary surface needs to be understood to ascertain whether Earth is singular among known rocky planets, and to frame predictions of future changes to the biosphere. Modern geomorphic observations and modelling have made strides in this respect, but an under-utilized lens through which to interrogate these questions resides in the most complete tangible record of our planetary history: the sedimentary-stratigraphic record (SSR). The characteristics of the SSR have been frequently explained with reference to changes in boundary conditions such as relative sea level, climate, and tectonics. Yet despite the fact that the long-term accrual of the SSR was contemporaneous with the evolution of almost all domains of life on Earth, causal explanations related to biological activity have often been overlooked, particularly within siliciclastic strata. This paper explores evidence for the ways in which organisms have influenced the SSR throughout Earth history and emphasizes that further investigation can help lead us towards a mechanistic understanding of how the planetary surface has co-evolved with life. The practicality of discerning life signatures in the SSR is discussed by: 1) distinguishing biologically-dependent versus biologically-influenced sedimentary signatures; 2) emphasizing the importance of determining relative time-length scales of processes and demonstrating how different focal lengths of observation (individual geological outcrops and the complete SSR) can reveal different insights; and 3) promoting an awareness of issues of equifinality and underdetermination that may hinder the recognition of life signatures. Multiple instances of life signatures and their historic range within the SSR are reviewed, with examples covering siliciclastic, biogenic and chemogenic strata, and trigger organisms from across the spectrum of Earth’s extant and ancient life. With this novel perspective, the SSR is recognised as a dynamic archive that expands and complements the fossil and geochemical records that it hosts, rather than simply being a passive repository for them. The SSR is shown to be both the record and the result of long-term evolutionary synchrony between life and planetary surface processes

Negligible microbial matground influence on pre-vegetation river functioning: Evidence from the Ediacaran-Lower Cambrian Series Rouge, France

The pre-Silurian alluvial rock record is dominated by accumulations of laterally-extensive, sheet-like sandstone strata with minimal mudrock; a depositional style frequently explained as representing fluvial processes particular to “pre-vegetation” Earth. While the sedimentological and geomorphological influence of Palaeozoic embryophytes and other higher vegetation has been commonly inferred, the influence of the non-marine microbial matgrounds that preceded them has been less well studied. The ?Ediacaran-Cambrian Series Rouge of northern France and the Channel Islands is a rare example of a predominantly alluvial succession which exhibits both pre-vegetation sedimentary motifs and evidence for the existence of terrestrial microbial mats. The latter include likely microbial sedimentary surface textures, the enigmatic matground “pseudofossils” Aristophycus and Arumberia, and probable mat fragments and mat-related microtextures preserved in argillaceous sediment. The sedimentological characteristics of the Series Rouge are described and analysed in order to assess the role of microbial influences on pre-vegetation alluvial systems. Near ubiquitously trough-cross bedded sheet-braided facies, with rarely preserved channel-forms, indicate that alluvial sedimentation was dominated by in-channel dune migration, and depositional-dip exposures reveal the periodic downstream migration of complex bar-forms. Lateral accretion elements and minor discontinuous lenses of more argillaceous material are locally present. Thus, despite the evidence for matgrounds, sedimentary architecture was essentially ‘abiotic’. Using this evidence from the Series Rouge, we argue that the surficial cohesion provided by matgrounds did not exceed thresholds for reworking by hydrodynamic processes thus having little or no effect on their preserved sedimentary architecture.Supported by Shell International Exploration and Production B.V under Research Framework agreement PT38181

Short-term evolution of primary sedimentary surface textures (microbial, abiotic, ichnological) on a dry stream bed: modern observations and ancient implications

A wide variety of sub-ripple-scale sedimentary surface textures are known from bedding planes in the sedimentary rock record. Many of these textures were traditionally ascribed an abiotic origin (e.g., due to rain drop impact, adhesion, etc.), but in recent decades the role of microbial mats and biofilms in sculpting and mediating some forms has become increasingly recognized. Microbial sedimentary textures are now well-described and understood from modern tidal environments and biological soil crusts, but descriptions from fluvial settings are less common, despite their known occurrence in ancient alluvium. This paper reports a suite of primary sedimentary surface textures which were observed forming in discrete bodies of standing water in the lower reaches of the ephemeral Murchison River, Western Australia. Microbial sedimentary signatures included bubble impressions (burst and intact) and roll-ups, in addition to reduced horizons. Many of these features exhibited rapid temporal evolution of their morphology in the dry days following an interval of heavy rain. Significantly, these microbial features were witnessed in close spatial proximity to other abiotic and biotic sedimentary surface textures including raindrop impressions, adhesion marks, desiccation cracks, and vertebrate and invertebrate traces. Such proximity of abiotic and microbial sedimentary surface textures is rarely reported from bedding planes in the rock record, but these modern observations emphasize the fact that, particularly in non-marine environments, such structures should not be expected to be mutually exclusive. An appreciation of the fact that primary sedimentary surface textures such as these develop during intervals of stasis in a sedimentation system is crucial to our understanding of their significance and diversity in the rock record.APS was supported by the Natural Environment Research Council [grant number NE/L002507/1]. WJM was supported by Shell International Exploration and Production B.V under Research Framework 604 agreement PT38181

Molecular Gas in Three z ∼ 7 Quasar Host Galaxies

We present ALMA band 3 observations of the CO(6-5), CO(7-6), and [C i] 369 μm emission lines in three of the highest-redshift quasar host galaxies at . These measurements constitute the highest-redshift CO detections to date. The target quasars have previously been detected in [C ii] 158 μm emission and the underlying FIR dust continuum. We detect (spatially unresolved, at a resolution of > 2″, or ≈14 kpc) CO emission in all three quasar hosts. In two sources, we detect the continuum emission around 400 μm (rest-frame), and in one source we detect [C i] at low significance. We derive molecular gas reservoirs of (1-3) T10 10 in the quasar hosts, i.e., approximately only 10 times the mass of their central supermassive black holes. The extrapolated [C ii]-to-CO(1-0) luminosity ratio is 2500-4200, consistent with measurement s in galaxies at lower redshift. The detection of the [C i] line in one quasar host galaxy and the limit on the [C i] emission in the other two hosts enables a first characterization of the physical properties of the interstellar medium in z ∼ 7 quasar hosts. In the sources, the derived global CO/[C ii] /[C i] line ratios are consistent with expectations from photodissociation regions, but not X-ray-dominated regions. This suggest that quantities derived from the molecular gas and dust emission are related to ongoing star-formation activity in the quasar hosts, providing further evidence that the quasar hosts studied here harbor intense starbursts in addition to their active nucleus

Specific involvement of atypical PKCζ/PKMζ in spinal persistent nociceptive processing following peripheral inflammation in rat.

BACKGROUND: Central sensitization requires the activation of various intracellular signalling pathways within spinal dorsal horn neurons, leading to a lowering of activation threshold and enhanced responsiveness of these cells. Such plasticity contributes to the manifestation of chronic pain states and displays a number of features of long-term potentiation (LTP), a ubiquitous neuronal mechanism of increased synaptic strength. Here we describe the role of a novel pathway involving atypical PKCζ/PKMζ in persistent spinal nociceptive processing, previously implicated in the maintenance of late-phase LTP. RESULTS: Using both behavioral tests and in vivo electrophysiology in rats, we show that inhibition of this pathway, via spinal delivery of a myristoylated protein kinase C-ζ pseudo-substrate inhibitor, reduces both pain-related behaviors and the activity of deep dorsal horn wide dynamic range neurons (WDRs) following formalin administration. In addition, Complete Freund's Adjuvant (CFA)-induced mechanical and thermal hypersensitivity was also reduced by inhibition of PKCζ/PKMζ activity. Importantly, this inhibition did not affect acute pain or locomotor behavior in normal rats and interestingly, did not inhibited mechanical allodynia and hyperalgesia in neuropathic rats. Pain-related behaviors in both inflammatory models coincided with increased phosphorylation of PKCζ/PKMζ in dorsal horn neurons, specifically PKMζ phosphorylation in formalin rats. Finally, inhibition of PKCζ/PKMζ activity decreased the expression of Fos in response to formalin and CFA in both superficial and deep laminae of the dorsal horn. CONCLUSIONS: These results suggest that PKCζ, especially PKMζ isoform, is a significant factor involved in spinal persistent nociceptive processing, specifically, the manifestation of chronic pain states following peripheral inflammation

Equation of state and high-pressure/high-temperature phase diagram of magnesium

The phase diagram of magnesium has been investigated to 211 GPa at 300 K, and to 105 GPa at 4500 K, by using a combination of x-ray diffraction and resistive and laser heating. The ambient pressure hcp structure is found to start transforming to the bcc structure at ∼45 GPa, with a large region of phase-coexistence that becomes smaller at higher temperatures. The bcc phase is stable to the highest pressures reached. The hcp-bcc phase boundary has been studied on both compression and decompression, and its slope is found to be negative and steeper than calculations have previously predicted. The laser-heating studies extend the melting curve of magnesium to 105 GPa and suggest that, at the highest pressures, the melting temperature increases more rapidly with pressure than previously reported. Finally, we observe some evidence of a new phase in the region of 10 GPa and 1200 K, where previous studies have reported a double-hexagonal-close-packed (dhcp) phase. However, the additional diffraction peaks we observe cannot be accounted for by the dhcp phase alone