1,302 research outputs found
Gain control from beyond the classical receptive field in primate primary visual cortex
Gain control is a salient feature of information processing throughout the visual system. Heeger (1991, 1992) described a mechanism that could underpin gain control in primary visual cortex (VI). According to this model, a neuron's response is normalized by dividing its output by the sum of a population of neurons, which are selective for orientations covering a broad range. Gain control in this scheme is manifested as a change in the semisaturation constant (contrast gain) of a VI neuron. Here we examine how flanking and annular gratings of the same or orthogonal orientation to that preferred by a neuron presented beyond the receptive field modulate gain in V1 neurons in anesthetized marmosets (Callithrix jacchus). To characterize how gain was modulated by surround stimuli, the Michaelis-Menten equation was fitted to response versus contrast functions obtained under each stimulus condition. The modulation of gain by surround stimuli was modelled best as a divisive reduction in response gain. Response gain varied with the orientation of surround stimuli, but was reduced most when the orientation of a large annular grating beyond the classical receptive field matched the preferred orientation of neurons. The strength of surround suppression did not vary significantly with retinal eccentricity or laminar distribution. In the mannoset, as in macaques (Angelucci et al., 2002a,b), gain control over the sort of distances reported here (up to 10 deg) may be mediated by feedback from extrastriate areas
Environmental constraints influencing survival of an African parasite in a north temperate habitat: effects of temperature on development within the host
The monogenean Protopolystoma xenopodis has been established in Wales for >40 years following introduction with Xenopus laevis from South Africa. This provides an experimental system for determining constraints affecting introduced species in novel environments. Parasite development post-infection was followed at 15, 20 and 25 °C for 15 weeks and at 10 °C for51 year and correlated with temperatures recorded inWales. Development was slowed/arrested at410 °C which reflects habitat conditions for >6 months/year. There was wide variation in growth at constant temperature (body size differing by >10 times) potentially attributable in part to genotype-specific host-parasite interactions. Parasite density had no effect on size but host sex did: worms in males were 1·8 times larger than in females. Minimum time to patency was 51 days at 25 °C and 73 days at 20 °C although some infections were still not patent at both temperatures by 105 days p.i. In Wales, fastest developing infections may mature within one summer (about 12 weeks), possibly accelerated by movements of hosts into warmer surface waters. Otherwise, development slows/stops in October–April, delaying patency to about 1 year p.i., while wide variation in developmental rates may impose delays of 2 years in some primary infections and even longer in secondary infections
Environmental constraints influencing survival of an African parasite in a north temperate habitat: effects of temperature on egg development
SUMMARYFactors affecting survival of parasites introduced to new geographical regions include changes in environmental temperature. Protopolystoma xenopodis is a monogenean introduced with the amphibian Xenopus laevis from South Africa to Wales (probably in the 1960s) where low water temperatures impose major constraints on life-cycle processes. Effects were quantified by maintenance of eggs from infections in Wales under controlled conditions at 10, 12, 15, 18, 20 and 25°C. The threshold for egg viability/ development was 15°C. Mean times to hatching were 22 days at 25°C, 32 days at 20°C, extending to 66 days at 15°C. Field temperature records provided calibration of transmission schedules. Although egg production continues year-round, all eggs produced during >8 months/ year die without hatching. Output contributing significantly to transmission is restricted to 10 weeks (May-mid-July). Host infection, beginning after a time lag of 8 weeks for egg development, is also restricted to 10 weeks (July-September). Habitat temperatures (mean 15·5°C in summer 2008) allow only a narrow margin for life-cycle progress: even small temperature increases, predicted with 'global warming', enhance infection. This system provides empirical data on the metrics of transmission permitting long-term persistence of isolated parasite populations in limiting environments
Metallicities of 0.3<z<1.0 Galaxies in the GOODS-North Field
We measure nebular oxygen abundances for 204 emission-line galaxies with
redshifts 0.3<z<1.0 in the Great Observatories Origins Deep Survey North
(GOODS-N) field using spectra from the Team Keck Redshift Survey (TKRS). We
also provide an updated analytic prescription for estimating oxygen abundances
using the traditional strong emission line ratio, R_{23}, based on the
photoionization models of Kewley & Dopita (2003). We include an analytic
formula for very crude metallicity estimates using the [NII]6584/Halpha ratio.
Oxygen abundances for GOODS-N galaxies range from 8.2< 12+log(O/H)< 9.1
corresponding to metallicities between 0.3 and 2.5 times the solar value. This
sample of galaxies exhibits a correlation between rest-frame blue luminosity
and gas-phase metallicity (i.e., an L-Z relation), consistent with L-Z
correlations of previously-studied intermediate-redshift samples. The zero
point of the L-Z relation evolves with redshift in the sense that galaxies of a
given luminosity become more metal poor at higher redshift. Galaxies in
luminosity bins -18.5<M_B<-21.5 exhibit a decrease in average oxygen abundance
by 0.14\pm0.05 dex from z=0 to z=1. This rate of metal enrichment means that
28\pm0.07% of metals in local galaxies have been synthesized since z=1, in
reasonable agreement with the predictions based on published star formation
rate densities which show that ~38% of stars in the universe have formed during
the same interval. (Abridged)Comment: AASTeX, 49 pages, 16 figures, accepted for publication in The
Astrophysical Journa
Consequences of a Change in the Galactic Environment of the Sun
The interaction of the heliosphere with interstellar clouds has attracted
interest since the late 1920's, both with a view to explaining apparent
quasi-periodic climate "catastrophes" as well as periodic mass extinctions.
Until recently, however, models describing the solar wind - local interstellar
medium (LISM) interaction self-consistently had not been developed. Here, we
describe the results of a two-dimensional (2D) simulation of the interaction
between the heliosphere and an interstellar cloud with the same properties as
currently, except that the neutral H density is increased from the present
value of n(H) ~ 0.2 cm^-3 to 10 cm^-3. The mutual interaction of interstellar
neutral hydrogen and plasma is included. The heliospheric cavity is reduced
considerably in size (approximately 10 - 14 AU to the termination shock in the
upstream direction) and is highly dynamical. The interplanetary environment at
the orbit of the Earth changes markedly, with the density of interstellar H
increasing to ~2 cm^-3. The termination shock itself experiences periods where
it disappears, reforms and disappears again. Considerable mixing of the shocked
solar wind and LISM occurs due to Rayleigh-Taylor-like instabilities at the
nose, driven by ion-neutral friction. Implications for two anomalously high
concentrations of 10Be found in Antarctic ice cores 33 kya and 60 kya, and the
absence of prior similar events, are discussed in terms of density enhancements
in the surrounding interstellar cloud. The calculation presented here supports
past speculation that the galactic environment of the Sun moderates the
interplanetary environment at the orbit of the Earth, and possibly also the
terrestrial climate.Comment: 23 pages, 2 color plates (jpg), 3 figures (eps
Why Are Radio-Galaxies Prolific Producers of Type Ia Supernovae?
An analysis of SNIa events in early type galaxies from the Cappellaro et al
(1999) database provides conclusive evidence that the rate of type Ia
Supernovae (SNe) in radio-loud galaxies is about 4 times higher than the rate
measured in radio-quiet galaxies, i.e. SNIa-rate SNu as compared to SNIa-rate SNu. The actual value of the
enhancement is likely to be in the range (P). This
finding puts on robust empirical grounds the results obtained by Della Valle &
Panagia (2003) on the basis of a smaller sample of SNe. We analyse the possible
causes of this result and conclude that the enhancement of SNIa explosion rate
in radio-loud galaxies has the same origin as their being strong radio sources,
but there is no causality link between the two phenomena. We argue that
repeated episodes of interaction and/or mergers of early type galaxies with
dwarf companions, on times-scale of about 1 Gyr, are responsible for inducing
both strong radio activity observed in 14% of early type galaxies and to
supply an adequate number of SNIa progenitors to the stellar population of
ellipticals.Comment: 26 pages+6 figures, ApJ, in pres
The Evolutionary Status of Isolated Dwarf Irregular Galaxies II. Star Formation Histories and Gas Depletion
The results of UBV and H alpha imaging of a large sample of isolated dwarf
irregular galaxies are interpreted in the context of composite stellar
population models. The observed optical colors are best fit by composite
stellar populations which have had approximately constant star formation rates
for at least 10 Gyr. The galaxies span a range of central surface brightness,
from 20.5 to 25.0 mag arcsec^{-2}; there is no correlation between surface
brightness and star formation history. Although the current star formation
rates are low, it is possible to reproduce the observed luminosities without a
major starburst episode. The derived gas depletion timescales are long,
typically ~20 Gyr. These results indicate that dwarf irregular galaxies will be
able to continue with their slow, but constant, star formation activity for at
least another Hubble time.
The sample of isolated dIs is compared to a sample of star bursting dwarf
galaxies taken from the literature. The star bursting dwarf galaxies have many
similar properties; the main difference between these two types of gas-rich
dwarf galaxies is that the current star formation is concentrated in the center
of the star bursting systems while it is much more distributed in the quiescent
dIs. This results in pronounced color gradients for the starbursting dwarf
galaxies, while the majority of the quiescent dwarf irregular galaxies have
minor or non-existent color gradients. Thus, the combination of low current
star formation rates, blue colors, and the lack of significant color gradients
indicates that star formation percolates slowly across the disk of normal dwarf
galaxies in a quasi-continuous manner.Comment: 16 pages, uses emulateapj, to appear in The Astronomical Journal
(April 2001
The Revival of Galactic Cosmic Ray Nucleosynthesis?
Because of the roughly linear correlation between Be/H and Fe/H in low
metallicity halo stars, it has been argued that a ``primary'' component in the
nucleosynthesis of Be must be present in addition to the ``secondary''
component from standard Galactic cosmic ray nucleosynthesis. In this paper we
critically re-evaluate the evidence for the primary versus secondary character
of Li, Be, and B evolution, analyzing both in the observations and in Galactic
chemical evolution models. While it appears that [Be/H] versus [Fe/H] has a
logarithmic slope near 1, it is rather the Be-O trend that directly arises from
the physics of spallation production. Using new abundances for oxygen in halo
stars based on UV OH lines, we find that the Be-O slope has a large uncertainty
due to systematic effects, rendering it difficult to distinguish from the data
between the secondary slope of 2 and the primary slope of 1. The possible
difference between the Be-Fe and Be-O slopes is a consequence of the variation
in O/Fe versus Fe: recent data suggests a negative slope rather than zero
(i.e., Fe O) as is often assumed. In addition to a phenomenological
analysis of Be and B evolution, we have also examined the predicted LiBeB, O,
and Fe trends in Galactic chemical evolution models which include outflow.
Based on our results, it is possible that a good fit to the LiBeB evolution
requires only traditional the Galactic cosmic ray spallation, and the (primary)
neutrino-process contribution to B11. We thus suggest that these two processes
might be sufficient to explain Li6, Be, and B evolution in the Galaxy, without
the need for an additional primary source of Be and B.Comment: 25 pages, latex, 8 ps figures, figure 1 correcte
Low Mass Stars and the He3 Problem
The prediction of standard chemical evolution models of higher abundances of
He3 at the solar and present-day epochs than are observed indicates a possible
problem with the yield of He3 for stars in the range of 1-3 solar masses.
Because He3 is one of the nuclei produced in Big Bang Nucleosynthesis (BBN), it
is noted that galactic and stellar evolution uncertainties necessarily relax
constraints based on He3. We incorporate into chemical evolution models which
include outflow, the new yields for He3 of Boothroyd & Malaney (1995) which
predict that low mass stars are net destroyers of He3. Since these yields do
not account for the high \he3/H ratio observed in some planetary nebulae, we
also consider the possibility that some fraction of stars in the 1 - 3 solar
mass range do not destroy their He3 in theirpost main-sequence phase. We also
consider the possibility that the gas expelled by stars in these mass ranges
does not mix with the ISM instantaneously thus delaying the He3 produced in
these stars, according to standard yields, from reaching the ISM. In general,
we find that the Galactic D and He3 abundances can be fit regardless of whether
the primordial D/H value is high (2 x 10^{-4}) or low (2.5 x 10^{-5}).Comment: 20 pages, latex, 9 ps figure
The Chemical Enrichment History of the Large Magellanic Cloud
Ca II triplet spectroscopy has been used to derive stellar metallicities for
individual stars in four LMC fields situated at galactocentric distances of
3\arcdeg, 5\arcdeg, 6\arcdeg\@ and 8\arcdeg\@ to the north of the Bar. Observed
metallicity distributions show a well defined peak, with a tail toward low
metallicities. The mean metallicity remains constant until 6\arcdeg\@
([Fe/H]-0.5 dex), while for the outermost field, at 8\arcdeg, the mean
metallicity is substantially lower than in the rest of the disk
([Fe/H]-0.8 dex). The combination of spectroscopy with deep CCD
photometry has allowed us to break the RGB age--metallicity degeneracy and
compute the ages for the objects observed spectroscopically. The obtained
age--metallicity relationships for our four fields are statistically
indistinguishable. We conclude that the lower mean metallicity in the outermost
field is a consequence of it having a lower fraction of intermediate-age stars,
which are more metal-rich than the older stars. The disk age--metallicity
relationship is similar to that for clusters. However, the lack of objects with
ages between 3 and 10 Gyr is not observed in the field population. Finally, we
used data from the literature to derive consistently the age--metallicity
relationship of the bar. Simple chemical evolution models have been used to
reproduce the observed age--metallicity relationships with the purpose of
investigating which mechanism has participated in the evolution of the disk and
bar. We find that while the disk age--metallicity relationship is well
reproduced by close-box models or models with a small degree of outflow, that
of the bar is only reproduced by models with combination of infall and outflow.Comment: 45 pages, 10 figures, accepted for publication in Astronomical
Journa
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