The Tiffanian Land-Mammal Age (Middle and Late Paleocene) In The Northern Bighorn Basin, Wyoming

http://deepblue.lib.umich.edu/bitstream/2027.42/61362/1/SecordPapers35DeepBlue.pd

INTERACTIONS OF A PALEOCENE RIVER, A RISING FOLD, AND EARLY-DIAGENETIC CONCRETIONS

The relative rates of sediment accumulation, erosion, and structural uplift determine whether a growing fold develops positive topographic relief, is beveled by antecedent streams, or is buried under thick growth strata. When folds rise in subsiding basins, upward, convergent flow of groundwater through the permeable growth strata that underlie antecedent streams enhances the flux of ions required for concretion growth. Early diagenetic concretions that grow in such alluvial strata may constitute the only clasts larger than sand size available for transport when antecedent streams become erosive. The first reworked concretions deposited by these streams should accurately mark the transition from aggradation to erosion as folds rise into the paths of streams. In this situation, the ability to differentiate reworked from in situ concretions is crucial. The west-vergent Simpson Ridge anticline, a N–S-trending, thick-skinned Laramide structure in east-central Wyoming, separates the larger Hanna basin from the Carbon basin. Near the north nose of this anticline, in situ ironoxide- rich concretions are abundant in folded Paleocene strata (Ferris Formation) and, just to the east, large, reworked, iron-rich concretions are abundant in younger, more gently dipping conglomerates in the basal Hanna Formation of the backlimb. Smaller reworked concretions are also present near the base of the Hanna Formation at least 7 km south of the anticlinal nose and just east of the fold’s axis. At the anticlinal nose, in situ (non-reworked) concretions up to 3 m X 1 m X 1 m are abundant at the top of an ~ 7- km-thick sequence of sandstones and siltstones that constitute the Late Cretaceous–early Paleocene Ferris Formation. Reworked concretions are absent in the strata hosting these in situ concretions, but reworked concretionary clasts up to 2 m in diameter are present in exposures of conglomerates in the lowermost Hanna Formation that lie just above the in situ Ferris concretions and southeast of the anticlinal nose. These early-diagenetic concretions were originally cemented by siderite (FeCO3). Oxidation of some small, rinded siderite-cemented clasts took place after their fluvial transport into the Hanna Formation, but abundant angular, un-rinded, iron-oxide-cemented clasts indicate that many large, in situ siderite concretions had resided in the vadose zone before they were entrained. The distribution of reworked concretions and the orientations of crossbeds show that antecedent Hanna streams eroded a swath at least 5 km wide across the rising structure. These streams transported Ferris Formation concretions southeastward into the Carbon basin, and deposited them in a conglomeratic sandstone body in the Hanna Formation. Large calcitecemented concretions, many with a pipe-like morphology, then grew within Hanna crossbeds. In many cases, these in situ concretions enclose transported, iron-rich concretions, but there is no evidence any calcite-cemented concretions were reworked. The NW–SE alignment of the pipes record southeastward flow of groundwater and thus also provide evidence (together with the orientation of the crossbeds) of the original continuity of the Hanna Formation across the anticline. Reworked concretions reveal the interplay of deposition, diagenesis, and erosion. Due to convergent groundwater flow over growing anticlines, early diagenetic concretions, both in situ and as reworked clasts, are especially likely to be found in growth strata

\u3ci\u3eMylanodon rosei\u3c/i\u3e, a New Metacheiromyid (Mammalia: Palaeanodonta) from the Late Tiffanian (Late Paleocene) of Northwestern Wyoming

Mylanodon rosei is a new genus and species of late Paleocene metacheiromyid palaeanodont from a new late Tiffanian locality, Y2K Quarry, in the Clarks Fork Basin, Wyoming. The type is an adult dentary with P4 and a molariform double-rooted M1. This provides the first evidence that molariform teeth were retained in early Metacheiromyidae. A second specimen is a juvenile dentary with a partial P3 and an unerupted P4. This is the first juvenile dentition known for a Paleocene metacheiromyid. The new specimens enable determination of dental homologies. Reduction of teeth in early metacheiromyids took place from back to front, opening the characteristic posterior diastema. Both Mylanodon and Propalaeanodon, a slightly older metacheiromyid, are intermediate morphologically and temporally between the older Tiffanian epoicotheriid Amelotabes and the younger Clarkforkian and Wasatchian metacheiromyid Palaeanodon. Propalaeanodon has a single-rooted M1, a derived characteristic not found in Mylanodon, suggesting that two lineages are involved and Propalaeanodon was not ancestral to Mylanodon

A new taeniolabidoid multituberculate (Mammalia) from the middle Puercan of the Nacimiento Formation, New Mexico, and a revision of taeniolabidoid systematics and phylogeny

Multituberculates were amongst the most abundant and taxonomically diverse mammals of the late Mesozoic and the Paleocene, reaching their zenith in diversity and body size in the Paleocene. Taeniolabidoidea, the topic of this paper, includes the largest known multituberculates, which possess highly complex cheek teeth adapted for herbivory. A new specimen from the early Paleocene (middle Puercan; biochron Pu2) of the Nacimiento Formation, New Mexico represents a new large-bodied taeniolabidoid genus and species, Kimbetopsalis simmonsae. A phylogenetic analysis to examine the relationships within Taeniolabidoidea that includes new information from Kimbetopsalis gen. et sp. nov. and gen. nov. and from new specimens of Catopsalis fissidens, first described here, and data from all other described North American and Asian taeniolabidoids. This analysis indicates that Catopsalis is nonmonophyletic and justifies our transfer of the basal-most taeniolabidoid ‘Catopsalis’ joyneri to a new genus, Valenopsalis. Kimbetopsalis and Taeniolabis form a clade (Taeniolabididae), as do the Asian Lambdopsalis, Sphenopsalis, and possibly also Prionessus (Lambdopsalidae). Taeniolabidoids underwent a modest taxonomic radiation during the early Paleocene of North America and underwent a dramatic increase in body size, with Taeniolabis taoensis possibly exceeding 100 kg. Taeniolabidoids appear to have gone extinct in North America by the late Paleocene but the appearance of lambdopsalids in the late Paleocene of Asia suggests that they dispersed from North America in the early to middle Paleocene

Male mastodon landscape use changed with maturation (late Pleistocene, North America)

Under harsh Pleistocene climates, migration and other forms of seasonally patterned landscape use were likely critical for reproductive success of mastodons (Mammut americanum) and other megafauna. However, little is known about how their geographic ranges and mobility fluctuated seasonally or changed with sexual maturity. We used a spatially explicit movement model that coupled strontium and oxygen isotopes from two serially sampled intervals (5+ adolescent years and 3+ adult years) in a male mastodon tusk to test for changes in landscape use associated with maturation and reproductive phenology. The mastodon’s early adolescent home range was geographically restricted, with no evidence of seasonal preferences. Following inferred separation from the matriarchal herd (starting age 12 y), the adolescent male’s mobility increased as landscape use expanded away from his natal home range (likely central Indiana). As an adult, the mastodon’s monthly movements increased further. Landscape use also became seasonally structured, with some areas, including northeast Indiana, used only during the inferred mastodon mating season (spring/summer). The mastodon died in this area (\u3e150 km from his core, nonsummer range) after sustaining a craniofacial injury consistent with a fatal blow from a competing male’s tusk during a battle over access to mates. Northeast Indiana was likely a preferred mating area for this individual and may have been regionally significant for late Pleistocene mastodons. Similarities between mammutids and elephantids in herd structure, tusk dimorphism, tusk function, and the geographic component of male maturation indicate that these traits were likely inherited from a common ancestor

Mylanodon rosei, a new metacheiromyid (Mammalia, Palaeanodonta) from the late Tiffinian (late Paleocene) of northwestern Wyoming

385-399http://deepblue.lib.umich.edu/bitstream/2027.42/48666/2/ID533.pd

Terrestrial carbon isotope stratigraphy and mammal turnover during post-PETM hyperthermals in the Bighorn Basin, Wyoming, USA

Paleogene hyperthermals, including the Paleocene–Eocene Thermal Maximum (PETM) and several other smaller events, represent global perturbations to Earth’s climate system and are characterized by warmer temperatures, changes in floral and faunal communities, and hydrologic changes. These events are identified in the geologic record globally by negative carbon isotope excursions (CIEs), resulting from the input of isotopically light carbon into Earth’s atmosphere. Much about the causes and effects of hyperthermals remains uncertain, including whether all hyperthermals were caused by the same underlying processes, how biotic effects scale with the magnitude of hyperthermals, and why CIEs are larger in paleosol carbonates relative to marine records. Resolving these questions is crucial for a full understanding of the causes of hyperthermals and their application to future climate scenarios. The primary purpose of this study was to identify early Eocene hyperthermals in the Fifteenmile Creek area of the south-central Bighorn Basin, Wyoming, USA. This area preserves a sequence of fluvial floodplain sedimentary rocks containing paleosol carbonates and an extensive record of fossil mammals. Previous analysis of faunal assemblages in this area revealed two pulses of mammal turnover and changes in diversity interpreted to correlate with the ETM2 and H2 hyperthermals that follow the PETM. This was, however, based on long-distance correlation of the fossil record in this area with chemostratigraphic records from elsewhere in the basin. We present new carbon isotope stratigraphies using micrite δ13C values from paleosol carbonate nodules preserved in and between richly fossiliferous mammal localities at Fifteenmile Creek to identify the stratigraphic positions of ETM2 and H2. Carbon isotope results show that the ETM2 and H2 hyperthermals, and possibly the subsequent I1 hyperthermal, are recorded at Fifteenmile Creek. ETM2 and H2 overlap with the two previously recognized pulses of mammal turnover. The CIEs for these hyperthermals are also somewhat smaller in magnitude than in more northerly Bighorn Basin records. We suggest that basin-wide differences in soil moisture and/or vegetation could contribute to variable CIE amplitudes in this and other terrestrial records

Systematics and Phylogeny of Late Paleocene and Early Eocene Palaeoryctinae (Mammalia, Insectivora) from the Clarks Fork and Bighorn Basins, Wyoming

p. 119-154http://deepblue.lib.umich.edu/bitstream/2027.42/41252/1/C31-5.pd

Scientific publishing and the reading of science in nineteenth-century Britain: a historiographical survey and guide to sources

[FIRST PARAGRAPH] It is now generally accepted that both the conception and practices of natural enquiry in the Western tradition underwent a series of profound developments in the late eighteenth and early nineteenth century—developments which have been variously characterized as a ‘second scientific revolution’ and, much more tellingly, as the ‘invention of science’. As several authors have argued, moreover, a crucial aspect of this change consisted in the distinctive audience relations of the new sciences. While eighteenth-century natural philosophy was distinguished by an audience relation in which, as William Whewell put it, ‘a large and popular circle of spectators and amateurs [felt] themselves nearly upon a level, in the value of their trials and speculations, with more profound thinkers’, the science which was invented in the late eighteenth and early nineteenth century was, as Simon Schaffer has argued, marked by the ‘emergence of disciplined, trained cadres of research scientists’ clearly distinguished from a wider, exoteric public. Similarly, Jan Golinski argues that the ‘emergence of new instrumentation and a more consolidated social structure for the specialist community’ for early nineteenth-century chemistry was intimately connected with the transformation in the role of its public audience to a condition of relative passivity. These moves were underpinned by crucial epistemological and rhetorical shifts—from a logic of discovery, theoretically open to all, to a more restrictive notion of discovery as the preserve of scientific ‘genius’, and from an open-ended philosophy of ‘experience’ to a far more restrictive notion of disciplined ‘expertise’. Both of these moves were intended to do boundary work, restricting the community active in creating and validating scientific knowledge, and producing a passive public

Pan-Cancer Analysis of lncRNA Regulation Supports Their Targeting of Cancer Genes in Each Tumor Context

Long noncoding RNAs (lncRNAs) are commonly dys-regulated in tumors, but only a handful are known toplay pathophysiological roles in cancer. We inferredlncRNAs that dysregulate cancer pathways, onco-genes, and tumor suppressors (cancer genes) bymodeling their effects on the activity of transcriptionfactors, RNA-binding proteins, and microRNAs in5,185 TCGA tumors and 1,019 ENCODE assays.Our predictions included hundreds of candidateonco- and tumor-suppressor lncRNAs (cancerlncRNAs) whose somatic alterations account for thedysregulation of dozens of cancer genes and path-ways in each of 14 tumor contexts. To demonstrateproof of concept, we showed that perturbations tar-geting OIP5-AS1 (an inferred tumor suppressor) andTUG1 and WT1-AS (inferred onco-lncRNAs) dysre-gulated cancer genes and altered proliferation ofbreast and gynecologic cancer cells. Our analysis in-dicates that, although most lncRNAs are dysregu-lated in a tumor-specific manner, some, includingOIP5-AS1, TUG1, NEAT1, MEG3, and TSIX, synergis-tically dysregulate cancer pathways in multiple tumorcontexts