Spherical alterations of handles: embedding the manifold plus construction

A key tool in our earlier work on ends of manifolds high-dimensional manifolds was an ability to embed cobordisms provided by the Quillen Plus Construction into those ends. Here we develop a `spherical modification' trick which provides a constructive approach to obtaining such embeddings. More importantly, this approach allows for more general embedding results. In this paper we develop generalizations of the plus construction and show how the corresponding cobordisms can be embedded in manifolds satisfying appropriate fundamental group properties. Results obtained here play an important role in our ongoing study of noncompact manifolds.Comment: This final version will appear in Algebraic & Geometric Topology. Small corrections, including a fix to the statement of Theorem 5.3. 22 pages, 4 figure

Environmental constraints influencing survival of an African parasite in a north temperate habitat: effects of temperature on development within the host

The monogenean Protopolystoma xenopodis has been established in Wales for >40 years following introduction with Xenopus laevis from South Africa. This provides an experimental system for determining constraints affecting introduced species in novel environments. Parasite development post-infection was followed at 15, 20 and 25 °C for 15 weeks and at 10 °C for51 year and correlated with temperatures recorded inWales. Development was slowed/arrested at410 °C which reflects habitat conditions for >6 months/year. There was wide variation in growth at constant temperature (body size differing by >10 times) potentially attributable in part to genotype-specific host-parasite interactions. Parasite density had no effect on size but host sex did: worms in males were 1·8 times larger than in females. Minimum time to patency was 51 days at 25 °C and 73 days at 20 °C although some infections were still not patent at both temperatures by 105 days p.i. In Wales, fastest developing infections may mature within one summer (about 12 weeks), possibly accelerated by movements of hosts into warmer surface waters. Otherwise, development slows/stops in October–April, delaying patency to about 1 year p.i., while wide variation in developmental rates may impose delays of 2 years in some primary infections and even longer in secondary infections

Environmental constraints influencing survival of an African parasite in a north temperate habitat: effects of temperature on egg development

SUMMARYFactors affecting survival of parasites introduced to new geographical regions include changes in environmental temperature. Protopolystoma xenopodis is a monogenean introduced with the amphibian Xenopus laevis from South Africa to Wales (probably in the 1960s) where low water temperatures impose major constraints on life-cycle processes. Effects were quantified by maintenance of eggs from infections in Wales under controlled conditions at 10, 12, 15, 18, 20 and 25°C. The threshold for egg viability/ development was 15°C. Mean times to hatching were 22 days at 25°C, 32 days at 20°C, extending to 66 days at 15°C. Field temperature records provided calibration of transmission schedules. Although egg production continues year-round, all eggs produced during >8 months/ year die without hatching. Output contributing significantly to transmission is restricted to 10 weeks (May-mid-July). Host infection, beginning after a time lag of 8 weeks for egg development, is also restricted to 10 weeks (July-September). Habitat temperatures (mean 15·5°C in summer 2008) allow only a narrow margin for life-cycle progress: even small temperature increases, predicted with 'global warming', enhance infection. This system provides empirical data on the metrics of transmission permitting long-term persistence of isolated parasite populations in limiting environments

Manifolds with non-stable fundamental groups at infinity, II

In this paper we continue an earlier study of ends non-compact manifolds. The over-arching goal is to investigate and obtain generalizations of Siebenmann's famous collaring theorem that may be applied to manifolds having non-stable fundamental group systems at infinity. In this paper we show that, for manifolds with compact boundary, the condition of inward tameness has substatial implications for the algebraic topology at infinity. In particular, every inward tame manifold with compact boundary has stable homology (in all dimensions) and semistable fundamental group at each of its ends. In contrast, we also construct examples of this sort which fail to have perfectly semistable fundamental group at infinity. In doing so, we exhibit the first known examples of open manifolds that are inward tame and have vanishing Wall finiteness obstruction at infinity, but are not pseudo-collarable.Comment: Published by Geometry and Topology at http://www.maths.warwick.ac.uk/gt/GTVol7/paper7.abs.htm

Biologie des populations des Monogènes Polystomatidae

Les cycles des monogènes polystomatidae montrent une très grande diversité. Parmi ceux qui infestent des amphibiens anoures, on devrait s'attendre à ce que la taille des populations parasites montre des différences prononcées, selon que des réinfestations interviennent régulièrement chaque année, ou qu'il n'y en ait qu'une dans la vie de l'hôte. Toutefois, à quelques exceptions près, les niveaux d'infestation sont généralement bas, quelle que soit la durée de vie de l'hôte. Les facteurs susceptibles de réguler les populations de polystomatidae parasites d'amphibiens anoures sont récapitulés ici, et nous nous penchons plus particulièrement sur les mécanismes contrôlant l'infestation et, par voie de conséquence, la survie post-infestation. Les effets d'un éventail de facteurs sont envisagés, parmi lesquels les contraintes environnementales externes (en particulier, la température), les facteurs liés à l'hôte (dont le comportement et la durée de vie) et les facteurs propres au parasite (dont la compétition intraspécifique). Deux genres de Polystomatidae témoignent d'une régulation densité-dépendante des infrapopulations unique, contrôlée par la production de deux types de larves. Il existe des données de terrain et de laboratoire qui permettent de quantifier les effets de ces différents paramètres pour un certain nombre d'espèces de Polystomes. Les résultats obtenus pour Pseudodiplorchis americanus suggèrent que, même lorsqu'ils sont combinés, les effets de ces différents facteurs ne suffisent pas à rendre compte de la puissante régulation que l'on observe dans les populations naturelles où, malgré de massives infestations annuelles, les populations de parasites adultes sont faibles en effectif et remarquablement stables d'une année à l'autre. C'est la preuve indirecte qu'une importante régulation intervient par l'intermédiaire de l'immunité-hôte. Pour pouvoir pousser plus loin l'interprétation de la dynamique des populations parasites, il est donc maintenant nécessaire d'intégrer les composantes écologiques et immunologiques. Pour pouvoir pousser plus loin l'interprétation de la dynamique des populations parasites, il est donc maintenant nécessaire d'intégrer les composantes écologiques et immunologiques

A study of pulsed thermography for life assessment of thin EB-PVD TBCs undergoing oxidation ageing

This paper presents an assessment of ageing for thin Thermal Barrier Coatings (TBC) using active thermography. As TBCs undergo ageing during their service life, sintering changes the porosity, elements migrate from the substrate, and micro-cracks build up in the structure of the material, exhibiting a change in thermal conductivity and diffusion properties. As the material ages and these properties change over time, it is possible to exploit trends in this change for characterisation of coating ageing, which would provide a diagnostics tool to estimate remaining useful life. In this study, through-depth diffusivity measurement has been applied to thin EB-PVD coatings which are artificially aged via oxidation furnace cycles. In order to address the difficulties of capturing a fast thermal event in a thin coating, a novel parametric study approach has been carried out to optimise data capture and analysis, maximising available frames for the model fitting step. Through-depth diffusivities have been measured during ageing for six samples, yielding a repeatable trend in thermal diffusivity measurements, with three features, which can be exploited for ageing characterisation of thin EB-PVD TBCs, and used as an alarm of imminent failure

Consequences of a Change in the Galactic Environment of the Sun

The interaction of the heliosphere with interstellar clouds has attracted interest since the late 1920's, both with a view to explaining apparent quasi-periodic climate "catastrophes" as well as periodic mass extinctions. Until recently, however, models describing the solar wind - local interstellar medium (LISM) interaction self-consistently had not been developed. Here, we describe the results of a two-dimensional (2D) simulation of the interaction between the heliosphere and an interstellar cloud with the same properties as currently, except that the neutral H density is increased from the present value of n(H) ~ 0.2 cm^-3 to 10 cm^-3. The mutual interaction of interstellar neutral hydrogen and plasma is included. The heliospheric cavity is reduced considerably in size (approximately 10 - 14 AU to the termination shock in the upstream direction) and is highly dynamical. The interplanetary environment at the orbit of the Earth changes markedly, with the density of interstellar H increasing to ~2 cm^-3. The termination shock itself experiences periods where it disappears, reforms and disappears again. Considerable mixing of the shocked solar wind and LISM occurs due to Rayleigh-Taylor-like instabilities at the nose, driven by ion-neutral friction. Implications for two anomalously high concentrations of 10Be found in Antarctic ice cores 33 kya and 60 kya, and the absence of prior similar events, are discussed in terms of density enhancements in the surrounding interstellar cloud. The calculation presented here supports past speculation that the galactic environment of the Sun moderates the interplanetary environment at the orbit of the Earth, and possibly also the terrestrial climate.Comment: 23 pages, 2 color plates (jpg), 3 figures (eps

Bergmann's body size rule operates in facultatively endothermic insects: evidence from a complex of cryptic bumblebee species

According to Bergmann’s rule we expect species with larger body size to inhabit locations with a cooler climate, where they may be well adapted to conserve heat and resist starvation. This rule is generally applied to endotherms. In contrast, body size in ectothermic invertebrates has been suggested to follow the reverse ecogeographic trend: these converse Bergmann’s patterns may be driven by the ecological constraints of shorter season length and lower food availability in cooler high latitude locations. Such patterns are particularly common in large insects due to their longer development times. As large and facultatively endothermic insects, bumblebees could thus be expected to follow either trend. In this investigation, we studied body size of three bumblebee species over a large spatial area and investigated whether interspecific trends in body size correspond to differences in their distribution consistent with either Bergmann’s or a converse Bergmann’s rule. We examined the body size of queens, males and workers of the Bombus lucorum complex of cryptic bumblebee species from across the whole of Great Britain. We found interspecific differences in body size corresponding to Bergmann’s rule: queens and males of the more northerly distributed, cool-adapted, species were largest. In contrast, the mean body size of the worker caste did not vary between the three species. These differences in body size may have evolved under selection pressures for thermoregulation or starvation resistance. We suggest that this case study in facultatively endothermic insects may help clarify the selection pressures governing Bergmann rule trends more generally

Old and young bulges in late-type disk galaxies

ABRIDGED: We use HSTACS and NICMOS imaging to study the structure and colors of a sample of nine late-type spirals. We find: (1) A correlation between bulge and disks scale-lengths, and a correlation between the colors of the bulges and those of the inner disks. Our data show a trend for bulges to be more metal-enriched than their surrounding disks, but otherwise no simple age-metallicity connection between these systems; (2) A large range in bulge stellar population properties, and, in particular, in stellar ages. Specifically, in about a half of the late-type bulges in our sample the bulk of the stellar mass was produced recently. Thus, in a substantial fraction of the z=0 disk-dominated bulged galaxies, bulge formation occurs after the formation/accretion of the disk; (3) In about a half of the late-type bulges in our sample, however, the bulk of the stellar mass was produced at early epochs; (4) Even these "old" late-type bulges host a significant fraction of stellar mass in a young(er) c component; (5) A correlation for bulges between stellar age and stellar mass, in the sense that more massive late-type bulges are older than less massive late-type bulges. Since the overall galaxy luminosity (mass) also correlates with the bulge luminosity (mass), it appears that the galaxy mass regulates not only what fraction of itself ends up in the bulge component, but also "when" bulge formation takes place. We show that dynamical friction of massive clumps in gas-rich disks is a plausible disk-driven mode for the formation of "old" late-type bulges. If disk evolutionary processes are responsible for the formation of the entire family of late-type bulges, CDM simulations need to produce a similar number of initially bulgeless disks in addition to the disk galaxies that are observed to be bulgeless at z=0.Comment: ApJ in press; paper with high resolution figures available at http://www.exp-astro.phys.ethz.ch/carollo/carollo1_2006.pdf; B, I, and H surface brightness profiles published in electronic tabular for