Epitypification of the name Colchicum montanum L. (Colchicaceae)

[EN] The identity of the illustration previously selected as the lectotype of Colchicum montanum L. is shown to be demonstrably ambiguous. An epitype is designated that affixes the application of the name to the common “quitameriendas” that is widely distributed in the Iberian Peninsula

Proposal to conserve the name Inula (Asteraceae) with a conserved type

[EN] The genus Inula L. as traditionally circumscribed comprises ca. 90–100 species widely distributed in Europe, Asia and Africa; ca. 65 species are Eurasian and North-African perennial herbs and ca. 25 species are Central and South-African perennial herbs and shrubs (modified from Anderberg in Pl. Syst. Evol. 176: 75–123. 1991 and Flann (ed.), 2009+ Global Compositae Checklist Ac-cessed: 11 Nov. 2011). The delimitation of the genus is controversial and has been a matter of frequent discussion throughout botanical history (e.g., Beck, Europ. Inula-Art.: 1–59. 1882; Anderberg, l.c. 1991). The original Linnaean (Sp. Pl.: 881–884. 1753) circumscrip-tion of Inula has notably changed to include further Linnaean gen-era such as Conyza L. (l.c.: 861–863, nom. rej. vs. Conyza Less., Syn. Gen. Compos.: 203–204. 1832, nom. cons.) (i.e., I. bifrons L., I. candida (L.) Cass. and I. conyzae (Griess.) Meikle). Also Inulahas been split into several genera and many Linnaean species origi-nally placed in Inula were subsequently transferred to them, e.g., Pulicaria Gaertn. (P. dysenterica (L.) Bernh., P. odora (L.) Rchb. and P. vulgaris Gaertn.) and Limbarda Adans. (L. crithmoides ( L .) Dumort.)

Chorological study of the genus "Filago" L. ("Asteraceae, Gnaphalieae") in the Iberian Peninsula and Balearic Islands

La complejidad taxonómica del género Filago ha dado lugar a numerosos errores de determinación a lo largo de los años, lo que ha impedido un conocimiento preciso del área de distribución de las especies incluidas en el género. En este trabajo tratamos de recopilar y revisar críticamente todos los datos disponibles acerca de la distribución de los 20 taxones presentes en la Península Ibérica e Islas Baleares. Como resultado más destacable, aportamos 13 novedades provinciales. Además, se circunscriben de manera precisa las áreas de distribución de F. gaditana, F. germanica, F. lutescens subsp. atlantica y F. micropodioides cuyas áreas de distribución eran mal conocidas por problemas taxonómicos y nomenclaturales. Además, se excluye la presencia de F. asterisciflora en el territorio objeto de Flora iberica.The taxonomical complexity of the genus Filago has traditionally led to many determination mistakes which have hampered a precise knowledge of the exact distribution areas of the species included in the genus. We have tried to compile all the available data on the distribution in the Iberian Peninsula and Balearic Islands of 20 taxa from the genus Filago and have critically revised them. As a result, in 13 cases we report the presence of a particular species of Filago for the first time at the provincial level. The distribution ranges of F. gaditana, F. germanica, F. lutescens subsp. atlantica and F. micropodioides traditionally hindered by taxonomic and nomenclatural problems are reliably circumscribed. Last and remarkably, according to our results, F. asterisciflora is not present in the Iberian Peninsula and in the Balearic Islands

Exploring the intricate evolutionary history of the diploid-polyploid complexVeronica subsection Pentasepalae(Plantaginaceae)

[EN]Veronica subsection Pentasepalae is a diploid–polyploid complex of c. 20 species distributed in Eurasia and North Africa, in which species boundaries are difficult to determine. Here, we present the first comprehensive phylogenetic analysis of V. subsection Pentasepalae based on nucleotide sequences [internal transcribed spacer (ITS) and the plastid trnH-psbA and ycf6-psbM spacers] combined with ploidy estimations. Our results support the monophyly of the subsection. Five well-supported clades are recovered in the ITS sequence analyses, corresponding to broad geographical areas. The causes of the extensive incongruence found between the ITS and plastid DNA datasets, namely incomplete lineage sorting and/or hybridization and polyploidization, are discussed. Most of the diploids traditionally recognized based on morphological characters and one tetraploid are each recovered as monophyletic by the ITS sequence analyses. The Balkan species V. kindlii is resurrected. DNA ploidy level for V. teucrioides is reported here for the first time (2x). Diploid populations have been found for V. orbiculata, which was previously thought to be only tetraploid. Past contact in the amphi-Adriatic area between V. orsiniana and V. orbiculata is suggested. Finally, molecular analyses show that diploid V. jacquinii and diploid V. orbiculata are unrelated. This study contributes to our understanding of the evolutionary history of polyploid complexes, especially those in southern Europe, and highlights the importance of using multiple lines of evidence to investigate species boundaries in such actively diversifying groups

A nomenclatural treatment for Veronica subsect. Pentasepalae (Plantaginaceae sensu APG III) and typification of several names

[EN] The large genus Veronica has a complex evolutionary history. Within it, Veronica subsect. Pentasepalae represents an ideal system to study important evolutionary processes, such as hybridization and polyploidization. Delimitation of species boundaries within the subsection is a difficult task and the highly complicated taxonomy of the group has led to an accumulation of approximately 230 names for 22 accepted taxa. Many names have been used in different taxonomic senses even in recent Floras, scientific works, catalogues, Red Lists and internet resources. The lack of an updated taxonomic and nomenclatural framework represents a significant problem to develop further studies in several fields such as ecology, conservation, plant and evolutionary biology. A complete nomenclatural treatment for the whole subsection is provided here, that relies on results derived from the first phylogenetic analysis of V. subsect. Pentasepalae based on DNA sequence data, morphology and ploidy‐ level information. Nomenclatural types for 22 validly published names are designated. In total 21 lectotypes, 2 epitypes and 1 neotype are chosen. A new combination, Veronica linearis, is proposed

Characterization of 12 Polymorphic SSR Markers in Veronica Subsect.Pentasepalae (Plantaginaceae) and Cross-Amplification in 10 Other Subgenera

[EN] • Premise of the study: Microsatellite primers were developed in the perennial herbs of the diploid-polyploid complex Veronica subsect. Pentasepalae (Plantaginaceae) to investigate the role that hybridization has played in the evolution of the group, which includes several endangered species. • Methods and Results: Twelve pairs of primers leading to polymorphic and readable markers were identifi ed and optimized from V. jacquinii and V. orbiculata using a microsatellite-enriched library method and 454 GS-FLX technique. The set of primers amplifi ed dinucleotide to pentanucleotide repeats, and the number of alleles per locus ranged from one to six, one to 11, and one to nine for V. orsiniana , V. javalambrensis , and V. rosea , respectively. Transferability analyses were performed in 20 species corresponding to 10 different subgenera. • Conclusions: These results indicate the utility of the newly developed microsatellites across Veronica subsect. Pentasepalae , which will help in the study of gene fl ow patterns and genetic structure

A contribution toward clarifying the nomenclature of Veronica L. (Plantaginaceae)

[EN] During a taxonomic revision of the genus Veronica L. for the Flora iberica project, an attempt was made to fix the correct status of the names cited in the study area. With this aim, here we have designated lectotypes, neotypes and epitypes for 18 previously untypified names, mainly belonging to annual species, some of them widely distributed such as V. persica and V. praecox. These newly proposed types support the current use of the names concerned and therefore contribute to establishing the correct nomenclature of the genus

Overlooked singularity and tiny plants: the Filago desertorum clade (Gnaphalieae, Asteraceae)

[EN] Filago desertorum, as traditionally circumscribed, is a species that shows high levels of morphological variation. Previous authors have even suggested that this taxon should represent a heterogeneous assembly of true biological species. A taxonomic revision of the species included in the Filago desertorum clade was performed, and amplified fragment length polymorphism (AFLP) markers were used to explore the phylogenetic relationships among the members of the clade. Three species are recognized in the group, one of which is newly described. A full description of F. castroviejoi sp. nov., a complete nomenclatural treatment and a key to the species included in the clade are provided

Aportaciones al conocimiento de la flora de Valladolid

New floristic records for Valladolid province.Palabras clave. Flora. Corología. Valladolid. España.Key words. Flora. Chorology. Valladolid. Spain