Occurrence and management of oak in southern Swedish forests

This article describes the current proportions of forest types with oak (Quercus robur and Q. petraea) in southern Sweden, provides an overview of oak distribution over time and reviews literature about oak regeneration relevant for the region. Further we discuss silvicultural possibilities to maintain and promote oak in Scandinavia. In Götaland pure oak forest covers 1% of the forest area and mixed forest types with > 10% oak proportion cover approximately 10% of the area. Common types of mixture are spruce-oak and pine-oak forest. Both mixtures are frequent in mature forest, especially pine-oak. Additionally, about one third of spruce-oak mixtures can be found in medium-aged forest. Intensive management would be necessary to promote single oak trees in old pine stands or spruce plantations, but the proportion of oak in coniferous forest provides some potential to maintain additional oak trees. The distribution of acorns by Jays, enhanced measures against browsing, and the release of single oak trees from competing tree species could help to maintain more oak trees for nature conservation. However, regarding management of oak for timber production, conventional methods are recommended. Planting after clear cutting of coniferous forest, or short shelter periods after mast years in oak stands, are established methods to regenerate pure oak stands. Another possibility to develop mature oak forest are mixed oak-spruce plantations, as traditionally practised in a small region in southern Sweden. The different approaches of oak management in Sweden were presented in April 2012 on the annual meeting of the section silviculture of DVFFA (German Union of Forest Research Organizations) in Wermsdorf near Leipzig to give an overview and access to recent forest research in Sweden

Heterogen skog

Svensk skogsforskning är inriktad på skötsel av trädslagsrena bestånd. I södra Sverige utgörs dock skogarna i relativt hög grad av blandskogar med olika sammansättningar. Kunskaperna om hur sådana skogar sköts är inte tillräcklig. Det finns många andra kombinationer av trädslag och beståndsstrukturer som är mindre typiska än tall–gran eller gran–björk och även mindre utforskade. Andra exempel på kunskapsbrist att sköta bestånd är skog med någon form av självföryngring. Om den tillämpade skogsforskningen inte har samma resurser för att utforska blandskog eller flerskiktad skog som bestånd med bara ett trädslag, då finns ett behov av ett mer generellt koncept för att också sköta heterogena bestånd. Praktiska råd i skogen för de som ibland arbetar med mer heterogena bestånd, av mindre vanlig typ: Börja med att kartlägga diameterfördelningen av beståndet, använd Heureka Beståndsvis och ha koll på självföryngring ifall du vill uppskatta framtidens beståndsutveckling och skötsel! Sedan kan man göra jämförelser med fallstudier beskrivna i litteraturen

Mixing effects on Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies (L.) Karst.) productivity along a climatic gradient across Europe

Mixed-species stands have been found to be more productive than would be expected from the performance of their component species in monocultures due to facilitation and complementarity between species, although these interactions depend on the combination of species present. Our study focuses on monospecific and mixed species stands of Scots pine and Norway spruce using 20 triplets established in nine countries along a climatic gradient across Europe. Differences in mean tree and stand characteristics, productivity and stand structure were assessed. Basal area increment in mixed stands was 8% higher than expected while volume increment was only 2% greater. Scots pine trees growing in mixed-species stands showed 11% larger quadratic mean diameter, 7% larger dominant diameter, 17% higher basal area and 25% higher stand volume than trees growing in mono specific stands. Norway spruce showed only a non-significant tendency to lower mean values of diameters, heights, basal area, as well standing volume in mixtures than monocultures. Stand structure indices differed between mixed stands and monocultures of Scots pine showing a greater stratification in mixed-species stands. Furthermore, the studied morphological traits showed little variability for trees growing in monospecific stands, except for diameter at breast height, crown length and crown length ratio. For trees growing in mixed stands, all the morphological traits of the trees were identified as different. Some of these morphological traits were associated with relative productivity. Nevertheless, relative productivity in mixed-species stands was not related to site conditions

With increasing site quality asymmetric competition and mortality reduces Scots pine (Pinus sylvestris L.) stand structuring across Europe

Heterogeneity of structure can increase mechanical stability, stress resistance and resilience, biodiversity and many other functions and services of forest stands. That is why many silvicultural measures aim at enhancing structural diversity. However, the effectiveness and potential of structuring may depend on the site conditions. Here, we revealed how the stand structure is determined by site quality and results from site-dependent partitioning of growth and mortality among the trees. We based our study on 90 mature, even-aged, fully stocked monocultures of Scots pine (Pines sylvestris L.) sampled in 21 countries along a productivity gradient across Europe. A mini-simulation study further analyzed the site-dependency of the interplay between growth and mortality and the resulting stand structure. The overarching hypothesis was that the stand structure changes with site quality and results from the site-dependent asymmetry of competition and mortality.First, we show that Scots pine stands structure across Europe become more homogeneous with increasing site quality. The coefficient of variation and Gini coefficient of stem diameter and tree height continuously decreased, whereas Stand Density Index and stand basal area increased with site index.Second, we reveal a site-dependency of the growth distribution among the trees and the mortality. With increasing site index, the asymmetry of both competition and growth distribution increased and suggested, at first glance, an increase in stand heterogeneity. However, with increasing site index, mortality eliminates mainly small instead of all-sized trees, cancels the size variation and reduces the structural heterogeneity.Third, we modelled the site-dependent interplay between growth partitioning and mortality. By scenario runs for different site conditions, we can show how the site-dependent structure at the stand level emerges from the asymmetric competition and mortality at the tree level and how the interplay changes with increasing site quality across Europe.Our most interesting finding was that the growth partitioning became more asymmetric and structuring with increasing site quality, but that the mortality eliminated predominantly small trees, reduced their size variation and thus reversed the impact of site quality on the structure. Finally, the reverse effects of mode of growth partitioning and mortality on the stand structure resulted in the highest size variation on poor sites and decreased structural heterogeneity with increasing site quality. Since our results indicate where heterogeneous structures need silviculture interventions and where they emerge naturally, we conclude that these findings may improve system understanding and modelling and guide forest management aiming at structurally rich forests

Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effect

The increasing disturbances in monocultures around the world are testimony to their instability under global change. Many studies have claimed that temporal stability of productivity increases with species richness, although the ecological fundamentals have mainly been investigated through diversity experiments. To adequately manage forest ecosystems, it is necessary to have a comprehensive understanding of the effect of mixing species on the temporal stability of productivity and the way in which it is influenced by climate conditions across large geographical areas. Here, we used a unique dataset of 261 stands combining pure and two-species mixtures of four relevant tree species over a wide range of climate conditions in Europe to examine the effect of species mixing on the level and temporal stability of productivity. Structural equation modelling was employed to further explore the direct and indirect influence of climate, overyielding, species asynchrony and additive effect (i.e. temporal stability expected from the species growth in monospecific stands) on temporal stability in mixed forests. We showed that by adding only one tree species to monocultures, the level (overyielding: +6%) and stability (temporal stability: +12%) of stand growth increased significantly. We identified the key effect of temperature on destabilizing stand growth, which may be mitigated by mixing species. We further confirmed asynchrony as the main driver of temporal stability in mixed stands, through both the additive effect and species interactions, which modify between-species asynchrony in mixtures in comparison to monocultures. Synthesis and applications. This study highlights the emergent properties associated with mixing two species, which result in resource efficient and temporally stable production systems. We reveal the negative impact of mean temperature on temporal stability of forest productivity and how the stabilizing effect of mixing two species can counterbalance this impact. The overyielding and temporal stability of growth addressed in this paper are essential for ecosystem services closely linked with the level and rhythm of forest growth. Our results underline that mixing two species can be a realistic and effective nature-based climate solution, which could contribute towards meeting EU climate target policies.Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effectpublishedVersio

Replacing monocultures with mixed-species stands: Ecosystem service implications of two production forest alternatives in Sweden

Whereas there is evidence that mixed-species approaches to production forestry in general can provide positive outcomes relative to monocultures, it is less clear to what extent multiple benefits can be derived from specific mixed-species alternatives. To provide such insights requires evaluations of an encompassing suite of ecosystem services, biodiversity, and forest management considerations provided by specific mixtures and monocultures within a region. Here, we conduct such an assessment in Sweden by contrasting even-aged Norway spruce (Picea abies)-dominated stands, with mixed-species stands of spruce and birch (Betula pendula or B. pubescens), or spruce and Scots pine (Pinus sylvestris). By synthesizing the available evidence, we identify positive outcomes from mixtures including increased biodiversity, water quality, esthetic and recreational values, as well as reduced stand vulnerability to pest and pathogen damage. However, some uncertainties and risks were projected to increase, highlighting the importance of conducting comprehensive interdisciplinary evaluations when assessing the pros and cons of mixtures

Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effect

The increasing disturbances in monocultures around the world are testimony to their instability under global change. Many studies have claimed that temporal stability of productivity increases with species richness, although the ecological fundamentals have mainly been investigated through diversity experiments. To adequately manage forest ecosystems, it is necessary to have a comprehensive understanding of the effect of mixing species on the temporal stability of productivity and the way in which it is influenced by climate conditions across large geographical areas. Here, we used a unique dataset of 261 stands combining pure and two-species mixtures of four relevant tree species over a wide range of climate conditions in Europe to examine the effect of species mixing on the level and temporal stability of productivity. Structural equation modelling was employed to further explore the direct and indirect influence of climate, overyielding, species asynchrony and additive effect (i.e. temporal stability expected from the species growth in monospecific stands) on temporal stability in mixed forests. We showed that by adding only one tree species to monocultures, the level (overyielding: +6%) and stability (temporal stability: +12%) of stand growth increased significantly. We identified the key effect of temperature on destabilizing stand growth, which may be mitigated by mixing species. We further confirmed asynchrony as the main driver of temporal stability in mixed stands, through both the additive effect and species interactions, which modify between-species asynchrony in mixtures in comparison to monocultures. Synthesis and applications. This study highlights the emergent properties associated with mixing two species, which result in resource efficient and temporally stable production systems. We reveal the negative impact of mean temperature on temporal stability of forest productivity and how the stabilizing effect of mixing two species can counterbalance this impact. The overyielding and temporal stability of growth addressed in this paper are essential for ecosystem services closely linked with the level and rhythm of forest growth. Our results underline that mixing two species can be a realistic and effective nature-based climate solution, which could contribute towards meeting EU climate target policies

Natural Regeneration in a Multi-Layered Pinus sylvestris-Picea abies Forest after Target Diameter Harvest and Soil Scarification

Forest management in Sweden can be characterized by even-aged silviculture heavily relying on three established harvest regimes: clearcutting, the seed-tree method, and the shelterwood system. Less intense, small-scale retention harvest systems such as single tree and group selection harvest are rarely used. In addition, natural regeneration dynamics without enrichment planting have barely been studied. Consequently, this study examined natural regeneration establishment in a multi-layered Pinus sylvestris-Picea abies forest stand in southwest Sweden after target diameter harvesting and soil scarification. The creation of forest canopy gaps had a positive effect on total seedling density five years after harvest, mainly due to a significantly higher number of Betula pendula individuals. Seedling density of more desirable tree species suitable for continuous cover forestry such as Fagus sylvatica, Quercus petraea and Picea abies also increased substantially in gaps when compared to pre-harvest conditions or the unharvested plots. In contrast, soil scarification did not increase the number of seedlings of desired tree species due to a significant decrease in Picea abies abundance. Soil moisture and gap size significantly improved Betula pendula seedling establishment while a larger number of Quercus petraea seedlings were observed in Vaccinium myrtillus patches. We conclude that canopy gaps are beneficial under the encountered stand conditions to initiate forest regeneration, and that soil scarification without the timely occurrence of a mast year of desired tree species is not effective in the type of forest studied

Examination of classification and deep learning procedures for the camera basedrecognition of natural objects.

In dieser Arbeit wird eine Auswahl an Verfahren vorgestellt und auf ihre Eignung zur Klassifizierung natürlicher Objekte untersucht. Hierzu werden eine Reihe von Experimenten basierend auf publizierten Verfahren durchgeführt, in denen überprüft wird, wie gut die entsprechenden Verfahren natürliche Objekte in Fotos erkennen können. Als natürliche Objekte wurden für diese Testreihe verschiedene Gemüse- und Obstsorten gewählt. Diese Arbeit ist dazu ausgelegt, als Entscheidungshilfe für die Erstellung von Anwendungen zu dienen, in welchen es notwendig ist natürliche Objekte korrekt zu klassifizieren.This paper presents a selection of methods and examines them for their suitability to recognize natural objects. For this purpose, a series of experiments based on published procedures are conducted. These experiments test how said procedures perform and if they are used to recognize natural objects in images. The natural objects used in these experiments are different kinds of vegetables and fruits. This paper is intended to serve as a decision-making aid for the implementation of an application which should, amongst other things, recognize natural objects