Assessing the protective role of alpine forests against rockfall at regional scale

Worldwide, mountain forests represent a significant factor in reducing rockfall risk over long periods of time on large potential disposition areas. While the economic value of technical protection measures against rockfall can be clearly determined and their benefits indicated, there is no general consensus on the quantification of the protective effect of forests. Experience shows that wherever there is forest, the implementation of technical measures to reduce risk of rockfall might often be dispensable or cheaper, and large deforestations (e.g. after windthrows, forest fires, clear-cuts) often show an increased incidence of rockfall events. This study focussed on how the protective effect of a forest against rockfall can be quantified on an alpine transregional scale. We therefore estimated the runout length, in terms of the angle of reach, of 700 individual rockfall trajectories from 39 release areas from Austria, Germany, Italy and Slovenia. All recorded rockfall events passed through forests which were classified either as coppice forests or, according to the CORINE classification of land cover, as mixed, coniferous or broadleaved dominated high forest stands. For each individual rockfall trajectory, we measured the forest structural parameters stem number, basal area, top height, ratio of shrub to high forest and share of coniferous trees. To quantify the protective effect of forests on rockfall, a hazard reduction factor is introduced, defined as the ratio between an expected angle of reach without forest and the back-calculated forest-influenced angles of reach. The results show that forests significantly reduce the runout length of rockfall. The highest reduction was observed for mixed high forest stands, while the lowest hazard reduction was observed for high forest stands dominated either by coniferous or broadleaved tree species. This implies that as soon as one tree species dominates, the risk reduction factor becomes lower. Coppice forests showed the lowest variability in hazard reduction. Hazard reduction due to forests increases, on average, by 7% for an increase in the stem number by 100 stems per hectare. The proposed concept allows a global view of the effectiveness of protective forests against rockfall processes and thus enable to value forest ecosystem services for future transregional assessments on a European level. Based on our results, general cost%benefit considerations of nature-based solutions against rockfall, such as protective forests as well as first-order evaluations of rockfall hazard reduction effects of silvicultural measures within the different forest types, can be supported

With increasing site quality asymmetric competition and mortality reduces Scots pine (Pinus sylvestris L.) stand structuring across Europe

Heterogeneity of structure can increase mechanical stability, stress resistance and resilience, biodiversity and many other functions and services of forest stands. That is why many silvicultural measures aim at enhancing structural diversity. However, the effectiveness and potential of structuring may depend on the site conditions. Here, we revealed how the stand structure is determined by site quality and results from site-dependent partitioning of growth and mortality among the trees. We based our study on 90 mature, even-aged, fully stocked monocultures of Scots pine (Pines sylvestris L.) sampled in 21 countries along a productivity gradient across Europe. A mini-simulation study further analyzed the site-dependency of the interplay between growth and mortality and the resulting stand structure. The overarching hypothesis was that the stand structure changes with site quality and results from the site-dependent asymmetry of competition and mortality.First, we show that Scots pine stands structure across Europe become more homogeneous with increasing site quality. The coefficient of variation and Gini coefficient of stem diameter and tree height continuously decreased, whereas Stand Density Index and stand basal area increased with site index.Second, we reveal a site-dependency of the growth distribution among the trees and the mortality. With increasing site index, the asymmetry of both competition and growth distribution increased and suggested, at first glance, an increase in stand heterogeneity. However, with increasing site index, mortality eliminates mainly small instead of all-sized trees, cancels the size variation and reduces the structural heterogeneity.Third, we modelled the site-dependent interplay between growth partitioning and mortality. By scenario runs for different site conditions, we can show how the site-dependent structure at the stand level emerges from the asymmetric competition and mortality at the tree level and how the interplay changes with increasing site quality across Europe.Our most interesting finding was that the growth partitioning became more asymmetric and structuring with increasing site quality, but that the mortality eliminated predominantly small trees, reduced their size variation and thus reversed the impact of site quality on the structure. Finally, the reverse effects of mode of growth partitioning and mortality on the stand structure resulted in the highest size variation on poor sites and decreased structural heterogeneity with increasing site quality. Since our results indicate where heterogeneous structures need silviculture interventions and where they emerge naturally, we conclude that these findings may improve system understanding and modelling and guide forest management aiming at structurally rich forests

Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effect

The increasing disturbances in monocultures around the world are testimony to their instability under global change. Many studies have claimed that temporal stability of productivity increases with species richness, although the ecological fundamentals have mainly been investigated through diversity experiments. To adequately manage forest ecosystems, it is necessary to have a comprehensive understanding of the effect of mixing species on the temporal stability of productivity and the way in which it is influenced by climate conditions across large geographical areas. Here, we used a unique dataset of 261 stands combining pure and two-species mixtures of four relevant tree species over a wide range of climate conditions in Europe to examine the effect of species mixing on the level and temporal stability of productivity. Structural equation modelling was employed to further explore the direct and indirect influence of climate, overyielding, species asynchrony and additive effect (i.e. temporal stability expected from the species growth in monospecific stands) on temporal stability in mixed forests. We showed that by adding only one tree species to monocultures, the level (overyielding: +6%) and stability (temporal stability: +12%) of stand growth increased significantly. We identified the key effect of temperature on destabilizing stand growth, which may be mitigated by mixing species. We further confirmed asynchrony as the main driver of temporal stability in mixed stands, through both the additive effect and species interactions, which modify between-species asynchrony in mixtures in comparison to monocultures. Synthesis and applications. This study highlights the emergent properties associated with mixing two species, which result in resource efficient and temporally stable production systems. We reveal the negative impact of mean temperature on temporal stability of forest productivity and how the stabilizing effect of mixing two species can counterbalance this impact. The overyielding and temporal stability of growth addressed in this paper are essential for ecosystem services closely linked with the level and rhythm of forest growth. Our results underline that mixing two species can be a realistic and effective nature-based climate solution, which could contribute towards meeting EU climate target policies

Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effect

The increasing disturbances in monocultures around the world are testimony to their instability under global change. Many studies have claimed that temporal stability of productivity increases with species richness, although the ecological fundamentals have mainly been investigated through diversity experiments. To adequately manage forest ecosystems, it is necessary to have a comprehensive understanding of the effect of mixing species on the temporal stability of productivity and the way in which it is influenced by climate conditions across large geographical areas. Here, we used a unique dataset of 261 stands combining pure and two-species mixtures of four relevant tree species over a wide range of climate conditions in Europe to examine the effect of species mixing on the level and temporal stability of productivity. Structural equation modelling was employed to further explore the direct and indirect influence of climate, overyielding, species asynchrony and additive effect (i.e. temporal stability expected from the species growth in monospecific stands) on temporal stability in mixed forests. We showed that by adding only one tree species to monocultures, the level (overyielding: +6%) and stability (temporal stability: +12%) of stand growth increased significantly. We identified the key effect of temperature on destabilizing stand growth, which may be mitigated by mixing species. We further confirmed asynchrony as the main driver of temporal stability in mixed stands, through both the additive effect and species interactions, which modify between-species asynchrony in mixtures in comparison to monocultures. Synthesis and applications. This study highlights the emergent properties associated with mixing two species, which result in resource efficient and temporally stable production systems. We reveal the negative impact of mean temperature on temporal stability of forest productivity and how the stabilizing effect of mixing two species can counterbalance this impact. The overyielding and temporal stability of growth addressed in this paper are essential for ecosystem services closely linked with the level and rhythm of forest growth. Our results underline that mixing two species can be a realistic and effective nature-based climate solution, which could contribute towards meeting EU climate target policies.Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effectpublishedVersio

Emerging stability of forest productivity by mixing two species buffers temperature destabilizing effect

The increasing disturbances in monocultures around the world are testimony to their instability under global change. Many studies have claimed that temporal stability of productivity increases with species richness, although the ecological fundamentals have mainly been investigated through diversity experiments. To adequately manage forest ecosystems, it is necessary to have a comprehensive understanding of the effect of mixing species on the temporal stability of productivity and the way in which it is influenced by climate conditions across large geographical areas. Here, we used a unique dataset of 261 stands combining pure and two-species mixtures of four relevant tree species over a wide range of climate conditions in Europe to examine the effect of species mixing on the level and temporal stability of productivity. Structural equation modelling was employed to further explore the direct and indirect influence of climate, overyielding, species asynchrony and additive effect (i.e. temporal stability expected from the species growth in monospecific stands) on temporal stability in mixed forests. We showed that by adding only one tree species to monocultures, the level (overyielding: +6%) and stability (temporal stability: +12%) of stand growth increased significantly. We identified the key effect of temperature on destabilizing stand growth, which may be mitigated by mixing species. We further confirmed asynchrony as the main driver of temporal stability in mixed stands, through both the additive effect and species interactions, which modify between-species asynchrony in mixtures in comparison to monocultures. Synthesis and applications. This study highlights the emergent properties associated with mixing two species, which result in resource efficient and temporally stable production systems. We reveal the negative impact of mean temperature on temporal stability of forest productivity and how the stabilizing effect of mixing two species can counterbalance this impact. The overyielding and temporal stability of growth addressed in this paper are essential for ecosystem services closely linked with the level and rhythm of forest growth. Our results underline that mixing two species can be a realistic and effective nature-based climate solution, which could contribute towards meeting EU climate target policies

Can trees at high elevations compensate for growth reductions at low elevations due to climate warming?

Radial tree stem growth of Picea abies, Fagus sylvatica, Pinus sylvestris and Pinus cembra was monitored from 2012 until 2015 across sites in Austria with high resolution dendrometers. Seasonal cumulative diameter increment was modeled using a hierarchical nonlinear mixed-effects model framework based on a logistic growth curve. In the dry and warm year 2015 the average annual diameter increment of 0.30 cm decreased by 50% on lower elevation sites and by 10% on higher elevation sites. In the cool and moist year 2014, Norway spruce achieved a higher annual diameter increment than European beech, whereas the opposite occurred in the dry and warm years 2013 and 2015. In the mixed beech-spruce stand, beech's consumptive water-use strategy has obviously caused intensified stress for spruce in these drought periods. On higher elevation sites Norway spruce reacted more sensitively to climate fluctuation compared with stone pine, but overall reactions were only weak. Productivity varied strongly dependent on the social tree status with dominant and intermediate trees suffering more from drought. As warming and drought lowers increment rates on lower elevation sites and as trees on higher elevation sites react less flexible, productivity losses are expected for Austrian forests due to climate warming.The accepted manuscript in pdf format is listed with the files at the bottom of this page. The presentation of the authors' names and (or) special characters in the title of the manuscript may differ slightly between what is listed on this page and what is listed in the pdf file of the accepted manuscript; that in the pdf file of the accepted manuscript is what was submitted by the author

Do competition-density rule and self-thinning rule agree?

International audienceKey message Competition-density relationships and self-thinning are major principles in forest growth. They are combined, describing self-thinning as a marginal case of the competition-density relationship. Robust estimation techniques allow parameterizing of both from national forest inventory data even for minor species. ContextThe competition-density principle, which can mathematically be described by the competition-density rule, is an important principle in plant ecology. The border line relationship is the self-thinning rule. Despite the importance of both rules for forest management, they have been fit for few species. AimsThe aim of this study is to compare competition-density rule and self-thinning rule in particular with respect to potential density for 15 species from the Austrian National Forest Inventory (ANFI). MethodsThe self-thinning line was estimated using quantile regression. The competition-density rule was fit as four- and as three-parameter model, where the fourth parameter was substituted (a) with a specific slope from the self-thinning line estimated from the ANFI and (b) Reineke’s slope (−1.605). ResultsPotential density was highest for Austrian pine and Norway spruce, followed by silver fir and Scots pine; it was considerably lower for European larch, stone pine and broadleaf species. Species-specific slopes of the self-thinning line ranged between −1.5 and −2.0 and were significantly different from Reineke’s slope for Norway spruce, European larch and European beech. ConclusionsUsing robust estimation techniques, both competition-density rule and self-thinning line can also be fit for minor species, providing an important guide for practical forest management

Climate-sensitive radial increment model of Norway spruce in Tyrol based on a distributed lag model with penalized splines for year-ring time series

A novel methodological framework is presented for climate-sensitive modeling of annual radial stem increment using year-ring width time series. The approach is based on a generalized additive model with penalized regression splines together with a distributed time lag model taking into account smooth nonlinear effects of a series of monthly temperature and precipitation values as well as interactions thereof. Climate effects are also assumed to vary smoothly with time lag. The model framework enables that both the detrending of the individual time series and the regression modeling can be performed simultaneously in a single model step. The approach is applied to year-ring width time series of Norway spruce (Picea abies (L.) H. Karst.) trees in Tyrol, Austria. The marginal response curves show that tree growth is mainly promoted by high temperatures in late spring and early summer and by precipitation in fall and winter. Summer drought does not have a negative influence on the current year's radial increment; however, when it is associated with high temperatures, it lowers the increment in the subsequent growth period. Higher winter precipitation in conjunction with lower temperatures have a positive effect. A significant non-climate related long-term growth trend is demonstrated, probably reflecting NOx and SO2 emission trends in Austria.The accepted manuscript in pdf format is listed with the files at the bottom of this page. The presentation of the authors' names and (or) special characters in the title of the manuscript may differ slightly between what is listed on this page and what is listed in the pdf file of the accepted manuscript; that in the pdf file of the accepted manuscript is what was submitted by the author