High N, dry: Experimental nitrogen deposition exacerbates native shrub loss and nonnative plant invasion during extreme drought.

Hotter, longer, and more frequent global change-type drought events may profoundly impact terrestrial ecosystems by triggering widespread vegetation mortality. However, severe drought is only one component of global change, and ecological effects of drought may be compounded by other drivers, such as anthropogenic nitrogen (N) deposition and nonnative plant invasion. Elevated N deposition, for example, may reduce drought tolerance through increased plant productivity, thereby contributing to drought-induced mortality. High N availability also often favors invasive, nonnative plant species, and the loss of woody vegetation due to drought may create a window of opportunity for these invaders. We investigated the effects of multiple levels of simulated N deposition on a Mediterranean-type shrubland plant community in southern California from 2011 to 2016, a period coinciding with an extreme, multiyear drought in the region. We hypothesized that N addition would increase native shrub productivity, but that this would increase susceptibility to drought and result in increased shrub loss over time. We also predicted that N addition would favor nonnatives, especially annual grasses, leading to higher biomass and cover of these species. Consistent with these hypotheses, we found that high N availability increased native shrub canopy loss and mortality, likely due to the higher productivity and leaf area and reduced water-use efficiency we observed in shrubs subject to N addition. As native shrub cover declined, we also observed a concomitant increase in cover and biomass of nonnative annuals, particularly under high levels of experimental N deposition. Together, these results suggest that the impacts of extended drought on shrubland ecosystems may be more severe under elevated N deposition, potentially contributing to the widespread loss of native woody species and vegetation-type conversion

Branden als EGM-maatregel

Periodiek branden is een traditionele gebruiks- of beheermethode die in het verleden op meerdere plaatsen werd toegepast. Dit gebeurde met name in heideterreinen en soms in duingraslanden, om de vegetatie te verjongen en de opslag van struiken en bomen tegen te gaan. geleidelijk is de beheersmethode uit het zicht verdwenen. In dit rapport zijn de ervaringen met periodiek branden en de kennis die er in binnenland en buitenland is over de effecten op de beschikbaarheid van nutriënten, vegetatie en fauna op een rij gezet. Het rapport geeft aan in welke situaties branden een aantrekkelijke effectgerichte maatregel kan zijn. In dit onderzoek (in het kader van OBN) werkten samen: Stichting Bargerveen, Bware en Alterr

Nitrogen enrichment lowers Betula pendula green and yellow leaf stoichiometry irrespective of effects of elevated carbon dioxide.

Elevated carbon dioxide (C

Antichresis: a comparative study of classical Roman law and the contractual praxis from Roman Egypt

The authors examine how papyrological sources from Roman Egypt written in Greek on antichresis relate to classical Roman law. Antichresis attested in papyrological antichretic contracts had a lot in common with antichresis emerging from Roman dispute resolutions. There was only one substantive difference: in classical Roman law, protection of the debtor was emphasized, whereas in the Greek papyrological antichretic contracts the position of the creditor was favoured. Given the similarities found, the authors conclude that antichretic loan both as an independent legal institution and as a pactum antichreticum was a pan-Mediterranean legal concept.Coherent privaatrech

Global assessment of nitrogen deposition effects on terrestrial plant diversity : a synthesis

Atmospheric nitrogen (N) deposition is it recognized threat to plant diversity ill temperate and northern parts of Europe and North America. This paper assesses evidence from field experiments for N deposition effects and thresholds for terrestrial plant diversity protection across a latitudinal range of main categories of ecosystems. from arctic and boreal systems to tropical forests. Current thinking on the mechanisms of N deposition effects on plant diversity, the global distribution of G200 ecoregions, and current and future (2030) estimates of atmospheric N-deposition rates are then used to identify the risks to plant diversity in all major ecosystem types now and in the future. This synthesis paper clearly shows that N accumulation is the main driver of changes to species composition across the whole range of different ecosystem types by driving the competitive interactions that lead to composition change and/or making conditions unfavorable for some species. Other effects such its direct toxicity of nitrogen gases and aerosols long-term negative effects of increased ammonium and ammonia availability, soil-mediated effects of acidification, and secondary stress and disturbance are more ecosystem, and site-specific and often play a supporting role. N deposition effects in mediterranean ecosystems have now been identified, leading to a first estimate of an effect threshold. Importantly, ecosystems thought of as not N limited, such as tropical and subtropical systems, may be more vulnerable in the regeneration phase. in situations where heterogeneity in N availability is reduced by atmospheric N deposition, on sandy soils, or in montane areas. Critical loads are effect thresholds for N deposition. and the critical load concept has helped European governments make progress toward reducing N loads on sensitive ecosystems. More needs to be done in Europe and North America. especially for the more sensitive ecosystem types. including several ecosystems of high conservation importance. The results of this assessment Show that the Vulnerable regions outside Europe and North America which have not received enough attention are ecoregions in eastern and Southern Asia (China, India), an important part of the mediterranean ecoregion (California, southern Europe). and in the coming decades several subtropical and tropical parts of Latin America and Africa. Reductions in plant diversity by increased atmospheric N deposition may be more widespread than first thought, and more targeted Studies are required in low background areas, especially in the G200 ecoregions

Onderzoek naar de relatie van eikensterfte met droogte en bodemchemie

In verband met verhevigde eikensterfte is onderzocht of er verband is tussen de eikensterfte en droogte en/of bodemchemie. Hiervoor is op tien locaties in een opstand met veel sterfte en een opstand met weinig sterfte de jaarringbreedte van de laatste 50 jaar vergeleken met weersgegevens van de dichtstbijzijnde weerstations, de hoeveelheid fijne wortels op 0-25 cm en 25-50 cm diepte bepaald en aan de hand van grondmonsters de bodemchemie gekarakteriseerd. Uit dit onderzoek is gebleken dat er een duidelijk verband is tussen de van tijd tot tijd optredende sterke groeivermindering van eiken en droogteperioden. Niet alle sterke groeiverminderingen zijn hiermee te verklaren. Er zijn aanwijzingen dat het aantal vorstdagen in de lente ook een rol speelt. In opstanden met veel sterfte zitten gemiddeld bovenin het bodemprofiel meer wortels en onderin minder dan in opstanden met weinig sterfte. In opstanden met veel sterfte heeft de bodem in de meeste gevallen lagere gehaltes aan uitwisselbaar Ca en/of K en/of Mg (dus een lagere buffercapaciteit). Het gehele proces van primaire oorzaken en secundaire factoren, die verantwoordelijk zijn voor de sterfte, is nog niet helder. Hiervoor is een breder en fundamenteler onderzoek noodzakelijk. Voor het beheer van eikenbossen worden voorlopige adviezen gegeven

Eikensterfte: een serieus en complex probleem

Steeds meer beheerders zien met lede ogen aan hoe de eiken in hun bossen langzaam maar zeker afsterven. Over de precieze oorzaak is nog niet zo heel veel bekend. Wel maakt onderzoek duidelijk dat er meerdere oorzaken zijn die elkaar lijken te versterken. Droogte en aantasting door insecten zijn zeker een deel van het probleem, maar ook de snelle uitspoeling van basische kationen zijn waarschijnlijk een heel belangrijke facto

Environmental differences between sites control the diet and nutrition of the carnivorous plant Drosera rotundifolia

Background and aims: Carnivorous plants are sensitive to small changes in resource availability, but few previous studies have examined how differences in nutrient and prey availability affect investment in and the benefit of carnivory. We studied the impact of site-level differences in resource availability on ecophysiological traits of carnivory for Drosera rotundifolia L. Methods: We measured prey availability, investment in carnivory (leaf stickiness), prey capture and diet of plants growing in two bogs with differences in N deposition and plant available N: Cors Fochno (0.62 g m−2 yr.−1, 353 μg l−1), Whixall Moss (1.37 g m−2 yr.−1, 1505 μg l−1). The total N amount per plant and the contributions of prey/root N to the plants’ N budget were calculated using a single isotope natural abundance method. Results: Plants at Whixall Moss invested less in carnivory, were less likely to capture prey, and were less reliant on prey-derived N (25.5% compared with 49.4%). Actual prey capture did not differ between sites. Diet composition differed – Cors Fochno plants captured 62% greater proportions of Diptera. Conclusions: Our results show site-level differences in plant diet and nutrition consistent with differences in resource availability. Similarity in actual prey capture may be explained by differences in leaf stickiness and prey abundance

Evidence for differential effects of reduced and oxidised nitrogen deposition on vegetation independent of nitrogen load

Nitrogen (N) deposition impacts natural and semi-natural ecosystems globally. The responses of vegetation to N deposition may, however, differ strongly between habitats and may be mediated by the form of N. Although much attention has been focused on the impact of total N deposition, the effects of reduced and oxidised N, independent of the total N deposition, have received less attention. In this paper, we present new analyses of national monitoring data in the UK to provide an extensive evaluation of whether there are differences in the effects of reduced and oxidised N deposition across eight habitat types (acid, calcareous and mesotrophic grasslands, upland and lowland heaths, bogs and mires, base-rich mires, woodlands). We analysed data from 6860 plots in the British Countryside Survey 2007 for effects of total N deposition and N form on species richness, Ellenberg N values and grass:forb ratio. Our results provide clear evidence that that N deposition affects species richness in all habitats except base-rich mires, after factoring out correlated explanatory variables (climate and sulphur deposition). In addition, the form of N in deposition appears important for the biodiversity of grasslands and woodlands but not mires and heaths. Ellenberg N increased more in relation to NHx deposition than NOy deposition in all but one habitat type. Relationships between species richness and N form were habitat-specific: acid and mesotrophic grasslands appear more sensitive to NHx deposition while calcareous grasslands and woodlands appeared more responsive to NOy deposition. These relationships are likely driven by the preferences of the component plant species for oxidised or reduced forms of N, rather than by soil acidification