Lightning, fire and longleaf pine: Using natural disturbance to guide management.

Abstract The importance of lightning as an ignition source for the fire adapted longleaf pine (Pinus palustris) ecosystem is widely recognized. Lightning also impacts this system on a smaller scale by causing individual tree mortality. The objective of this study was to determine mortality due to lightning and other agents in longleaf stands on the Ocala National Forest in central Florida and to quantify lightning ignited fire. Mortality from lightning was also tracked in longleaf stands on the Savannah River Site in South Carolina. Lightning killed more trees than any other agent with a mean mortality of nearly 1 tree/3 ha/yr in Florida and 1 tree/8 ha/yr in South Carolina. The probability of a tree being struck by lightning increased as a function of tree height at both sites, i.e. lightning preferentially removed the largest trees from the stand. In addition lightning strikes were clumped within stands, sometimes killed multiple trees with a single strike, and often hit trees on the edge of existing gaps. The combination of these processes means gaps suitable for regeneration within longleaf stands are created quite rapidly. This information provides guidelines for the development of selection harvest systems based on this natural disturbance. Although lightning activity was greatest during the summer months in Florida and most fires occurred in June, the probability of a strike causing a fire was highest in February to May. Published by Elsevier B.V

Influence of reproduction cutting methods on structure, growth and regeneration of longleaf pine forests in flatwoods and uplands

Though longleaf pine (Pinus palustris Mill.) forests have been primarily managed with even-aged methods, interest is increasing in uneven-aged systems, as a means of achieving a wider range of stewardship goals. Selection silviculture has been practiced on a limited scale in longleaf pine, but difficulty with using traditional approaches and absence of an evaluation across a range of site types has left managers in doubt concerning its suitability. This study was conducted to quantify the effects on stand dynamics of applying single-tree selection, group selection, irregular shelterwood and uniform shelterwood in longleaf pine forests on flatwoods and uplands of the southeastern United States. Selection treatments reduced stand basal area to approximately 11.5 m2 ha-1 and shelterwood treatments left a basal area of approximately 5.8 m2 ha-1. In spite of initial decreases in tree density and standing volume, growth rates were normal in all stands (1–5% per year), as were subsequent increases in basal area and tree density. Despite the continuing abundance of saw-palmetto (Serenoa repens W. Bartram) cover and absence of prescribed fire during the eight post-treatment years, significant increases in pine regeneration were observed in all treated stands in the flatwoods. Because of a multi-year drought in the uplands, pine seedling numbers dramatically declined, no matter which reproduction approach was employed. Although seedling numbers eventually began to recover, they were again precipitously depressed by a wildfire in 2013. Even with such losses, sufficient pine seedlings remained in each treatment to foster successful stand regeneration. Single-tree selection produced less overall change in the forest ecosystem than group selection, which caused less alteration than shelterwood treatment. Single-tree selection appears to be an effective way for achieving stand regeneration, while maintaining a continuous canopy cover that aids in the control of woody competitors and supports an array of resource values. Selection silviculture seems to be a lower risk approach for guiding forests along a trajectory of gradual improvement, with adjustments provided by frequent surface fires and periodic tree harvest. Long-term observation will be required to verify that selection can sustain forest ecosystems on sites characterized by differing environments

Influence of reproduction cutting methods on structure, growth and regeneration of longleaf pine forests in flatwoods and uplands

Though longleaf pine (Pinus palustris Mill.) forests have been primarily managed with even-aged methods, interest is increasing in uneven-aged systems, as a means of achieving a wider range of stewardship goals. Selection silviculture has been practiced on a limited scale in longleaf pine, but difficulty with using traditional approaches and absence of an evaluation across a range of site types has left managers in doubt concerning its suitability. This study was conducted to quantify the effects on stand dynamics of applying single-tree selection, group selection, irregular shelterwood and uniform shelterwood in longleaf pine forests on flatwoods and uplands of the southeastern United States. Selection treatments reduced stand basal area to approximately 11.5 m2 ha-1 and shelterwood treatments left a basal area of approximately 5.8 m2 ha-1. In spite of initial decreases in tree density and standing volume, growth rates were normal in all stands (1–5% per year), as were subsequent increases in basal area and tree density. Despite the continuing abundance of saw-palmetto (Serenoa repens W. Bartram) cover and absence of prescribed fire during the eight post-treatment years, significant increases in pine regeneration were observed in all treated stands in the flatwoods. Because of a multi-year drought in the uplands, pine seedling numbers dramatically declined, no matter which reproduction approach was employed. Although seedling numbers eventually began to recover, they were again precipitously depressed by a wildfire in 2013. Even with such losses, sufficient pine seedlings remained in each treatment to foster successful stand regeneration. Single-tree selection produced less overall change in the forest ecosystem than group selection, which caused less alteration than shelterwood treatment. Single-tree selection appears to be an effective way for achieving stand regeneration, while maintaining a continuous canopy cover that aids in the control of woody competitors and supports an array of resource values. Selection silviculture seems to be a lower risk approach for guiding forests along a trajectory of gradual improvement, with adjustments provided by frequent surface fires and periodic tree harvest. Long-term observation will be required to verify that selection can sustain forest ecosystems on sites characterized by differing environments

The National Fire and Fire Surrogate Study: Effects of Fuel Reduction Methods on Forest Vegetation Structure and Fuels

Changes in vegetation and fuels were evaluated from measurements taken before and after fuel reduction treatments (prescribed. re, mechanical treatments, and the combination of the two) at 12 Fire and Fire Surrogate (FFS) sites located in forests with a surface. re regime across the conterminous United States. To test the relative effectiveness of fuel reduction treatments and their effect on ecological parameters we used an information-theoretic approach on a suite of 12 variables representing the overstory (basal area and live tree, sapling, and snag density), the understory (seedling density, shrub cover, and native and alien herbaceous species richness), and the most relevant fuel parameters for wild. re damage (height to live crown, total fuel bed mass, forest floor mass, and woody fuel mass). In the short term (one year after treatment), mechanical treatments were more effective at reducing overstory tree density and basal area and at increasing quadratic mean tree diameter. Prescribed. re treatments were more effective at creating snags, killing seedlings, elevating height to live crown, and reducing surface woody fuels. Overall, the response to fuel reduction treatments of the ecological variables presented in this paper was generally maximized by the combined mechanical plus burning treatment. If the management goal is to quickly produce stands with fewer and larger diameter trees, less surface fuel mass, and greater herbaceous species richness, the combined treatment gave the most desirable results. However, because mechanical plus burning treatments also favored alien species invasion at some sites, monitoring and control need to be part of the prescription when using this treatment

Ecological effects of alternative fuel-reduction treatments: highlights of the National Fire and Fire Surrogate study (FFS)

The 12-site National Fire and Fire Surrogate study (FFS) was a multivariate experiment that evaluated ecological consequences of alternative fuel-reduction treatments in seasonally dry forests of the US. Each site was a replicated experiment with a common design that compared an un-manipulated control, prescribed fire, mechanical and mechanical + fire treatments. Variables within the vegetation, fuelbed, forest floor and soil, bark beetles, tree diseases and wildlife were measured in 10-ha stands, and ecological response was compared among treatments at the site level, and across sites, to better understand the influence of differential site conditions. For most sites, treated stands were predicted to be more resilient to wildfire if it occurred shortly after treatment, but for most ecological variables, short-term response to treatments was subtle and transient. Strong site-specificity was observed in the response of most ecosystem variables, suggesting that practitioners employ adaptive management at the local scale. Because ecosystem components were tightly linked, adaptive management would need to include monitoring of a carefully chosen set of key variables. Mechanical treatments did not serve as surrogates for fire for most variables, suggesting that fire be maintained whenever possible. Restoration to pre-settlement conditions will require repeated treatments over time, with eastern forests requiring more frequent applications.Keywords: Oak, Prescribed fire, Frequent fire regimes, Pine, Mechanical treatment, Forest thinning, Dry forest management, Seasonally dry forest

Effects of Prescribed Fire and Other Plant Community Restoration Treatments on Tree Mortality, Bark Beetles, and Other Saproxylic Coleoptera of Longleaf Pine, Pinus palustris Mill., on the Coastal Plain of Alabama

Treatments to restore understory plant communities of mature (50–80-year old) longleaf pine (Pinus palustris Mill.) and reduce risks of wildfire were applied to 10 ha plots that had a substantial shrub layer due to lack of fire. Plots were located in the Coastal Plain of Alabama and treatments consisted of: (1) untreated control, (2) growing season prescribed burn, (3) thin only, (4) thin plus growing season burn, and (5) herbicide plus growing season burn. Thin plus burn plots had significantly higher tree mortality compared to burn only and control plots and, overall, fire was the primary cause of tree death. Most tree mortality occurred within 1-year of treatment. From 2002 to 2004, we captured 75,598 Coleoptera in multiple funnel traps comprising 17 families and 130 species. Abundance of all Coleoptera combined was not different among treatments. Species richness was significantly higher on thin plus burn plots compared to thin only and control plots. Scolytinae (Coleoptera: Curculionidae) were more abundant on thin plus burn plots compared to control plots in fall 2002 but in fall 2003 they were more abundant on thin plus burn, thin only, and herbicide plus burn compared to controls. Among Scolytinae, Dendroctonus terebrans (Olivier), Xyleborinus saxeseni(Ratzeburg), Xyleborus sp. 3, and Hylastes tenuis (Eichhoff), showed varying responses to the treatments. Other Curculionidae were significantly more abundant on thin only and herbicide plus burn plots compared to all other treatments in spring 2003 and in spring 2004 they were more abundant on herbicide plus burn plots compared to thin plus burn treatments. Among Cerambycidae, Xylotrechus sagittatus (Germar) was higher in abundance in fall 2003 on thin plus burn plots compared to all other treatments except herbicide plus burn plots. Within the predator complex, Trogositidae were higher on thin plus burn plots compared to all other treatments except thin only plots in spring 2003, and Cleridae abundance was higher in spring 2004 on burn only plots compared to all other treatments. Linear regression analyses of dead trees per plot versus various Coleoptera showed captures of Buprestidae, Cerambycidae, Trogositidae, Acanthocinus nodosus (Fabricius), Temnochila virescens (Fabricius), and X. saxeseni increased with increasing number of dead trees. Our results show that the restoration treatments tested did not cause increased bark beetle-related tree mortality and they did not negatively affect populations of early successional saproxylic beetle fauna