Tertiary tectono-metamorphic evolution of the European margin during Alpine collison: example of the Leventina Nappe (Central Alps, Switzerland)

The Leventina Nappe represents one of the lowermost exposed units in the Alpine nappe stack and corresponds to a slice of the European margin that was entrained into the Alpine continental accretionary prism during the Tertiary tectonic event. This study yields details regarding the tectonic and metamorphic history of the Leventina Nappe, through detailed analysis of structures and shear zone patterns, and the examination of the Si-content of white mica along a north-south profile. The Leventina Nappe underwent three phases of ductile deformation. Foliation S1 is mostly sub-parallel to the regionally dominant structural fabric (the S2 foliation). S2 foliation is penetratively developed in the structurally higher portions of the Leventina Nappe toward the Simano Nappe, while it is only weakly developed in the core of the Leventina Nappe. A 50 to 200 m wide mylonite zone, with a D2 top-to-NW sense of shear marks the boundary to the Simano Nappe. Throughout the Leventina Nappe only small-scale D2 shear bands (mm to cm wide) are observed, showing a top-to-NW sense of shear. Deformation phase D3 locally generated a vertical axial plane foliation (S3) associated with the large-scale D3 Leventina antiform. Microtextural evidence and phengite geobarometry were used to constrain the temperature and pressure conditions of equilibration of the Leventina Gneisses. Highest Si (pfu) values are preserved in the core of phengitic micas and reflect pressure and temperature conditions of around 8 kbar at 550°C and 10 kbar at 650°C in the northern and southern parts of the Leventina Nappe, respectively. Lower Si (pfu) values from the rims of white micas correspond to a metamorphic pressure of ca. 5 kbar during the exhumation of the unit. These metamorphic conditions are related to the underthrusting of the thinned European margin into the continental accretionary prism during late Eocene time. These new data allow us to propose a kinematic model for the Leventina Nappe during the Tertiary Alpine tectonic

Chemical mass transfer in shear zones and metacarbonate xenoliths: a comparison of four mass balance approaches

International audienceMass balance calculations have been performed through a comparison of published graphical and statistical approaches applied to two contrasted geological settings: (i) the development of a greenschist-facies ductile shear-zone that recorded a weak volume change but significant mass transfers, and (ii) the formation of exoskarns in metacarbonate xenoliths that recorded a large volume decrease related to huge mass transfers. The comparison of the four mass-balance approaches shows that, if uncertainties are ignored, (1) they yield similar results concerning the mobile vs immobile behaviour of many components; (2) they yield similar mass-change values on bulk rock and on individual chemical elements (bulk-rock mass-change values differ by a maximum of ca. 15 % between graphical and statistical treatments of the metacarbonate xenolith evolution). The main difference concerns the uncertainties on mass changes (for bulk rocks and individual elements), which are much larger with the graphical than with the statistical approaches when uncertainties on chemical elements are taken into account, as they should be.The main advantage of the graphical methods is their rapid implementation and the clarity of the diagrams. Their main disadvantages are that uncertainties on each chemical element and bulk compositions are not taken into account and the difficulty in choosing an accurate immobility field to precisely define errors. Graphical methods need to be completed by a statistical treatment that gives absolute mass transfer results. The statistical approaches have the advantage of taking into account the chemical heterogeneities of the compared populations, in conjunction to a precise data treatment. The statistical treatment is an important and necessary step to decipher and to be pertinent in interpreting mobility/immobility of chemical elements, and, thus, in the absolute quantification of mass and volume changes

Mechanical decoupling of high-pressure crustal units during continental subduction.

International audienceThe mechanics of the transition from continental subduction toward upper crustal nappe stacking is still poorly understood and is studied here through a 2D thermal and strength numerical modelling of a subducted passive margin. Geological observations in the core of most mountain belts show the piling up of several HP-LT upper crustal units that are most likely related to the detachment of upper crustal units from the subducted continental margin and to the subsequent stacking of the detached units at depths. The Adula unit (Lepontine dome, Central Alps, Switzerland) is a long and thin upper crustal unit and is used here as a natural case-study as it provides a well-documented example of these units. 2D thermal modeling shows that two steps, successive in time, characterized the burial history of the passive margin undergoing continental subduction: 1-an increase in the margin strength due to an increase in the confining pressure during the first million years of the margin subduction and 2-the progressive heating of the subducted margin from the overlying lithosphere leads to a decrease in the margin strength due to thermal weakening, which progressively counter-balances the increase in confining pressure. Two strength gradients develop within the subducted lithosphere: 1-along the slab, the strength decreases with increasing burial depth and 2-perpendicular to the slab, the strength increases with depth due to an inverse temperature distribution. The detachment of HP-LT continental units from the subducted margin could occur when the slab strength becomes lower than the applied net stress. This allows the detachment of ductile weakened thin and long upper-crustal units. The thickness and length of the detached crustal units are controlled by the following parameters, in order of their importance: subduction dip angle, crustal rheology, mantle heat flux and subduction velocity. The comparison of our model results with the geometry and PT conditions of the Adula unit yields an estimate of the Alpine subduction dip angle at the time of deformation and metamorphism

Alteration of pathogenicity-linked life-history traits by resistance of its host Solanum tuberosum impacts sexual reproduction of the plant pathogenic oomycete Phytophthora infestans.

International audienceAlthough sexual reproduction implies a cost, it represents an evolutionary advantage for the adaptation and survival of facultative sexual pathogens. Understanding the maintenance of sex in pathogens requires to analyse how host resistance will impact their sexual reproduction through the alteration of their life-history traits. We explored this experimentally using potato (Solanum tuberosum) and one of its pathogens, the heterothallic oomycete Phytophthora infestans. Sexual reproduction was highest on hosts favouring asexual multiplication of the pathogen, suggesting similar nutritional requirements for both sexual and asexual sporulation. Sexual reproduction was also highest on hosts decreasing the latent period, probably because of a trade-off between growth and reproduction. Distinguishing host effects on each pathogenic trait remains however uneasy, as most life-history traits linked to pathogenicity were not independent of each other. We argue that sexual reproduction of P. infestans is an adaptation to survive when the host is susceptible and rapidly destroyed

Large calcite and bulk-rock volume loss in metacarbonate xenoliths from the Quérigut massif (French Pyrenees)

Chemical mass transfer was quantified in a metacarbonate xenolith enclosed within the granodiorite of the Quérigut massif (Pyrenees, France). Mass balance calculations suggest a strong decrease of CaO, SrO and CO2 contents (up to −90%), correlated with a decrease of modal calcite content as the contact is approached. Most other chemical elements behave immobile during metasomatism. They are therefore passively enriched. Only a small increase of SiO2, Al2O3 and Fe2O3 contents occurs in the immediate vicinity of the contact. Hence, in this study, skarn formation is characterized by the lack of large chemical element influx from the granitoid protolith. A large decrease of the initial carbonate volume (up to −86%) resulted from a combination of decarbonation reactions and loss of CaO and CO2. The resulting volume change has potentially important consequences for the interpretation of stable isotope profiles: the isotope alteration could have occured over greater distances than those observed toda

Reconstructing the total shortening history of the NW Himalaya.

The onset of India-Asia contact can be dated with both biostratigraphic analysis of syn-collisional sedimentary successions deposited on each side of the Indus Suture zone, and by radiometric dating of Indian crustal rocks which have undergone subduction to great depths in the earliest subduction-collision stages. These data, together with paleomagnetic data show that the initial contact of the Indian and Asian continental margins occurred at the Paleocene/Eocene boundary, corresponding to 55 ± 2 Ma. Such dating, which is consistent with all available geological evidence, including the record of magnetic anomalies in the Indian ocean and decrease of magmatic activity related to oceanic subduction can thus be considered as accurate and robust. The sedimentary record of the Tethys Himalaya rules out obduction of oceanic allochtons directly onto the Indian continental margin during the Late Cretaceous. The commonly inferred Late Cretaceous ophiolite obduction events may have thus occurred in intra-oceanic setting close to the Asian margin before its final emplacement onto the India margin during the Eocene. Granitoid and sedimentary rocks of the Indian crust, deformed during Permo-Carboniferous rifting, reached a depth of some 100 km about 1 Myr after the final closure of the Neo-Tethys, and began to be exhumed between 50 and 45 Ma. At this stage, the foreland basin sediments from Pakistan to India show significant supply from volcanic arcs and ophiolites of the Indus Suture Zone, indicating the absence of significant relief along the proto-Himalayan belt. Inversion of motion may have occurred within only 5 to 10 Myr after the collision onset, as soon as thicker and buoyant Indian crust chocked the subduction zone. The arrival of thick Indian crust within the convergent zone 50-45 Myr ago led to progressive stabilization of the India/Asia convergent rate and rapid stabilization of the Himalayan shortening rate of about 2 cm.yr-1. This first period also corresponds to the onset of terrestrial detrital sedimentation within the Indus Suture zone and to the Barrovian metamorphism on the Indian side of the collision zone. Equilibrium of the Himalayan thrust belt in terms of amount of shortening vs amount of erosion and thermal stabilization less than 10 Myr after the initial India/Asia contact is defined as the collisional regime. In contrast, the first 5 to 10 Myr corresponds to the transition from oceanic subduction to continental collision, characterized by a marked decrease of the shortening rate, onset of aerial topography, and progressive heating of the convergent zone. This period is defined as the continental subduction phase, accommodating more than 30% of the total Himalayan shortening

A Trade-Off Between Sporangia Size and Number Exists in the Potato Late Blight Pathogen Phytophthora infestans, and Is Not Altered by Biotic and Abiotic Factors

The negative relationship between offspring size and number is a classic example of trade-off between life-history traits, reported many times in animal and plant species. Here, we wanted to ascertain whether such a trade-off occurred in the oomycete Phytophthora infestans, and whether it was impacted by biotic and abiotic factors. We thus conducted three infection experiments under controlled conditions and measured the number and the size of sporangia (asexual propagules) produced on potato by different P. infestans isolates. In all experiments, we observed a negative relationship between sporangia size and number, demonstrating the existence of a trade-off. Moreover, although the potato host cultivar, temperature and host of origin (tomato or potato) all affected sporangia number, sporangia size or both, none of these biotic and abiotic factors did change the trade-off. Therefore, the trade-off between sporangia size and number could maintain the polyphenism for these traits in P. infestans populations, and favors the coexistence of distinct reproductive strategies within this species. Our results emphasize the relevance to focus on the relationship between offspring size and number in other fungal plant pathogens, as well as to study the impact of offspring size on fitness-linked traits (virulence and disease lesion development) in these organisms

Impact regimes and post-formation sequestration processes: implications for the origin of heavy noble gases in terrestrial planets

The difference between the measured atmospheric abundances of neon, argon, krypton, and xenon for Venus, Earth, and Mars is striking. Because these abundances drop by at least 2 orders of magnitude as one moves outward from Venus to Mars, the study of the origin of this discrepancy is a key issue that must be explained if we are to fully understand the different delivery mechanisms of the volatiles accreted by the terrestrial planets. In this work, we aim to investigate whether it is possible to quantitatively explain the variation of the heavy noble gas abundances measured on Venus, Earth, and Mars, assuming that cometary bombardment was the main delivery mechanism of these noble gases to the terrestrial planets. To do so, we use recent dynamical simulations that allow the study of the impact fluxes of comets upon the terrestrial planets during the course of their formation and evolution. Assuming that the mass of noble gases delivered by comets is proportional to the rate at which they collide with the terrestrial planets, we show that the krypton and xenon abundances in Venus and Earth can be explained in a manner consistent with the hypothesis of cometary bombardment. In order to explain the krypton and xenon abundance differences between Earth and Mars, we need to invoke the presence of large amounts of CO2-dominated clathrates in the Martian soil that would have efficiently sequestered these noble gases. Two different scenarios based on our model can also be used to explain the differences between the neon and argon abundances of the terrestrial planets. In the first scenario, cometary bombardment of these planets would have occurred at epochs contemporary with the existence of their primary atmospheres. Comets would have been the carriers of argon, krypton, and xenon, while neon would have been gravitationally captured by the terrestrial planets. In the second scenario, we consider impacting comets that contained significantly smaller amounts of argon, an idea supported by predictions of noble gas abundances in these bodies, provided that they formed from clathrates in the solar nebula. In this scenario, neon and argon would have been supplied to the terrestrial planets via the gravitational capture of their primary atmospheres whereas the bulk of their krypton and xenon would have been delivered by comets