Long-term ecosystem nutrient pool status for aspen forest harvest simulations on glacial till and sandy outwash soils\u3csup\u3e†\u3c/sup\u3e

Sandy outwash and glacial till soils compose large amounts of public forestland due to historically poor agricultural yields. The outwash soils have low fertility, poor nutrient retention and are restricted from whole-tree harvesting (WTH) in some states, whereas the glacial till has medium nutrient retention and fertility, and is unrestricted from WTH. To assess the long-term sustainability of harvesting, a nutrient budget was constructed from field measurements, the National Cooperative Soil Survey (NCSS) database, and literature values for stem-only harvesting (SOH) and WTH at a 45-year rotation length and 11 rotations were simulated. The budgets showed that SOH and WTH recovery years, or the time necessary for the inputs to match outputs through leaching and one harvest, exceeded common rotation lengths for both soil types under all weathering scenarios, and the average WTH reduced the total available rotations by one harvest. The large variation in soil nutrient pools and harvest removals complicated the ability to identify the difference between SOH and WTH early in the model, but differences became apparent with sequential harvests. The recovery years were 2–20 times the 45-year rotation length under all weathering rates. Taken together, models in this study bridge the gap between short-and long-term studies and bring into question the sustainability of WTH and SOH practices on nutrient-poor soils

The Northern White-Cedar Recruitment Bottleneck: Understanding the Effects of Substrate, Competition, and Deer Browsing

Research Highlights: Regenerating northern white-cedar (Thuja occidentalis L.) is challenging throughout much of its range. This study attempts to relate differences in natural regeneration to stand- and seedbed-level factors. Background and Objectives: Lack of regeneration of northern white-cedar is often attributed to overbrowsing by white-tailed deer (Odocoileus virginianus Zimmerman) because white-cedar is a preferred winter browse species. However, there are many other factors that may contribute to regeneration failure for white-cedar including its specific seedbed requirements and competition from other, often faster-growing trees and shrubs. Materials and Methods: We surveyed five mature white-cedar stands in Wisconsin, USA that have had little to no management in the past 50+ years to find stem densities of natural white-cedar regeneration in three height classes. We also collected data at each stand on potential predictor variables including overstory attributes, competitive environment, seedbed, and browsing by deer. We used model selection to create separate models to predict stem density of each white-cedar regeneration height class. Results: None of the measures of deer browsing used in this study were found to be associated with white-cedar regeneration. Soil pH, competition from other seedlings and saplings, and stem density of white-cedar in the overstory were found to be potentially associated with white-cedar regeneration. Conclusions: While browsing by deer is likely a factor affecting white-cedar regeneration in many areas, this study highlights the challenge of quantifying deer browse effects, as well as showing that other factors likely contribute to the difficulty of regenerating white-cedar

Plant communities and landform relationships delineate the components of soil complexes subject to whole-tree harvest restriction

Whole-tree harvesting practices are often restricted on soils with intrinsically low soil nutrients. Delineating restricted soil components from nonrestricted components presents a challenge for forest practitioners working with glaciofluvial parent materials. We investigated a suite of biological and edaphic factors to assess if restricted soil components are functionally distinct from nonrestricted soil components in partially restricted soil complexes across northern Wisconsin, USA. This study tested repeatable methods for soil delineation based on soil particle size, varying in spatial extent, and are independent of county boundaries or discrepancies across soil survey areas. Habitat type provided the most robust tool to distinguish the soil components and is the recommended approach. Habitat typing currently requires field data collection, since it does not exist as a geographic information system product in Wisconsin. Timber sale establishment often occurs during the off-season when habitat typing is not possible. In this case, an additional decision tree including habitat type and deviation from mean elevation (DEV) model is provided to estimate soil components remotely and verify the results when habitat typing is feasible. The 1,300-m radius DEV performed best with a 10-m digital elevation model, and reflects the differences between moraines and outwash plains landforms. These findings suggest that the utility of the DEV model and spatially explicit soil textural information in delineating restricted series from soil complexes has promise in light of improved remotely sensed information and geographic information system technologies that can be field-validated by habitat type

Evaluating Adaptive Management Options for Black Ash Forests in the Face of Emerald Ash Borer Invasion

The arrival and spread of emerald ash borer (EAB) across the western Great Lakes region has shifted considerable focus towards developing silvicultural strategies that minimize the impacts of this invasive insect on the structure and functioning of black ash (Fraxinus nigra) wetlands. Early experience with clearcutting in these forests highlighted the risks of losing ash to EAB from these ecosystems, with stands often retrogressing to marsh-like conditions with limited tree cover. Given these experiences and an urgency for increasing resilience to EAB, research efforts began in north-central Minnesota in 2009 followed by additional studies and trials in Michigan and Wisconsin to evaluate the potential for using regeneration harvests in conjunction with planting of replacement species to sustain forested wetland habitats after EAB infestations. Along with these more formal experiments, a number of field trials and demonstrations have been employed by managers across the region to determine effective ways for reducing the vulnerability of black ash forest types to EAB. This paper reviews the results from these recent experiences with managing black ash for resilience to EAB and describes the insights gained on the ecological functioning of these forests and the unique, foundational role played by black ash

The Case for Stand Management Guideliens as Dynamic as Global Change: Aspen Forest Stockings of the Western Great Lakes

Since the development of contemporary stocking techniques a century ago, the combination of climatic, atmospheric, financial, and social factors that determine forest management strategies have changed, altering aspen stand dynamics in the western Great Lakes, USA. Despite this, aspen management is still informed by 1970s management guides that are based on 1920s inventories; hence, a century exists between the data that underlie current management guidelines and current stand conditions. We hypothesized that current aspen stands may support higher stocking and height growth than nearly a century ago at relatively similar age and site indices, due to increased atmospheric CO2 concentrations and fertilization, intensive coppice harvests, and other factors. To explore this question, we compared historic aspen observations with comparable contemporary data from the USDA Forest Service\u27s Forest Inventory and Analysis program. The results show increased stand stocking levels as well as increased height growth of aspen throughout the region over the historic inventory data. Although other controlled experimental studies support the hypothesis of increased carbon fertilization altering aspen size-density relationships, our study is the first to examine an empirical application to forest management guides. Our results suggest a comprehensive reevaluation of aspen growth dynamics under contemporary environmental conditions is warranted. We highlight the need to assess the value of current stocking standards in an era of increasingly variable environmental conditions and to reimagine a more dynamic, responsive, and predictive approach to guide forest management for future application as global change may accelerate

Synthesis of plant-soil feedback effects on eastern North American tree species: implications for climate-adaptive forestry

Climate change represents an existential threat to many forest ecosystems because tree populations are often adapted to local climate means and variability. If tree populations cannot migrate or adapt, they risk becoming increasingly maladapted with climate change. This emerging mismatch underscores the need for climate adaptive management techniques, such as assisted migration of tree species, to help mitigate climate change impacts on forest ecosystems. Although biotic and abiotic factors are known to constrain tree establishment success, the extent to which they may determine the success of assisted migration plantings is poorly understood. Thus, defining the extent to which trees affect—and in turn are affected by local soil environments and microbial communities (i.e., plant-soil feedbacks; hereafter PSFs) remains important for guiding effective climate-adaptive forest management. Our objectives were to synthesize the current state of knowledge about the direction and magnitude of PSF effects on temperate tree species of eastern North America, and to identify key hypotheses important for guiding future research. To accomplish these goals, we conducted a meta-analysis of 26 peer-reviewed publications that addressed our criteria. Our compiled database included 61 tree species and was composed primarily of short-term greenhouse experiments that studied PSF effects by manipulating the soil biota in three ways: (1) soil was previously inoculated by a conspecific or heterospecific tree species (i.e., home vs. away), (2) soil was live or sterilized, or (3) soil was untreated or treated with fungicide. We found that PSF had significant effects on tree growth, with the direction and magnitude of PSF strongly dependent on tree mycorrhizal guild. Arbuscular mycorrhizal tree species grown in live or home soils grew 13–33% less than those in sterile or away soils, while ectomycorrhizal tree species grew 11–44% more in live or home than sterile or away soils. PSF effects were associated with several plant functional traits, including specific leaf area, tissue nitrogen, and specific root length. We provide suggestions on incorporating PSFs into assisted migration trials and outline key knowledge gaps for future research. Our synthesis of context-dependent effects of PSFs on tree performance will help inform management decisions involving assisted migration