Comments on the ophiuroid family Protasteridae and description of a new genus from the Lower Devonian of the Fox Bay Formation, Falkland Islands

Asterozoan fossils are comparatively rare in Gondwana compared with Laurentia, especially in the Devonian. We examined the only fossil ophiuroid yet known from the Falkland Islands and assess its significance for the evolution of the clade. This ophiuroid, herein distinguished as a new genus and species, Darwinaster coleenbiggsae, belongs to the same suprageneric group as Protaster, which was established on a series of Middle–Upper Ordovician taxa and persisted into the late Palaeozoic remarkably little changed in morphology. This single example is part of a much wider fauna that includes fossils from the Bokkeveld Group, South Africa and the Precordillera of Argentina. Existing palaeobiogeographic reconstructions confirm that these faunas once existed on contiguous terranes and characterized a distinct suite of similar palaeoenvironments within the Malvinokaffric Realm. This study reviews the existing record of Devonian asterozoans and revises Protasteridae

On the age of the Ballantrae Complex, SW Scotland

The Ballantrae Complex, SW Scotland, is an ophiolitic assemblage of mostly Early and early Middle Ordovician age (Tremadoc–Arenig in terms of the British Ordovician Series). Its varied components were generated and assembled in the Iapetus Ocean, then obducted on to the Laurentian continental margin by the earliest Llanvirn. The timing of obduction is constrained by biostratigraphic and radiometric data. It was most probably a polyphase process initiated at about the beginning of the Arenig, at around 478 Ma. However, parts of the Complex are significantly younger, with some recent evidence taken to suggest an earliest Llanvirn age of about 464 Ma for the emplacement of some of the volcanic and pelagic sedimentary rocks. The oldest strata in the succession that now unconformably overlies the Ballantrae Complex were deposited at about 463 Ma. Hence there may have been as little as one million years available for the final stages of the Complex's tectonic assembly, obduction, uplift, erosion and downfaulting. Obduction of the Complex has been invoked as a factor in the initiation of the Grampian Orogeny and, whilst there is a broad correlation in timing, the detail from Ballantrae militates against a causal relationship

Charles Darwin's discovery of Devonian fossils in the Falkland Islands, 1833, and its controversial consequences

In March 1833 Charles Darwin discovered Devonian fossils in the Falkland Islands. He was excited by his find but could have had little premonition of the long-running geological controversy that he was initiating. Darwin's fossils matched a coeval South African fauna, and as further collections were made the association was apparently strengthened. A particularly important contribution arose around 1910 through collaborations between a local collector, Constance Allardyce, and professional palaeontologists: Ernest Schwarz in South Africa and John Clarke in the USA. The accumulating evidence was seized upon by the early proponents of ‘displacement theory’ - continental drift - notably Alexander Du Toit, who relocated the Falkland Islands northward for his 1927 South Atlantic reconstruction. A more radical, but geologically sounder proposal arose in 1952 when Ray Adie suggested that the Falkland Islands, rotated through 180°, had originated as the eastward culmination of the Cape Fold Belt and Karoo Basin. In effect, Adie had presciently described a rotated microplate, perhaps the first on record. An opposing view saw the Falkland Islands as part of a fixed, South American promontory, and argument around these two contrasting interpretations of South Atlantic geology continues to the present day

A synopsis of the Ordovician System in its birthplace - Britain and Ireland

Rock successions in Britain and Ireland, and more especially those in North Wales, were instrumental in the founding and naming of the Ordovician System, and the Anglo-Welsh series established both initially and subsequently were used widely as a standard for Ordovician chronostratigraphy. Although now largely superseded in the global scheme of series and stages, they retain their local and regional importance. The Ordovician System in Britain and Ireland documents the history of a segment of the Earth's crust that incorporated opposing peri-Gondwanan and peri-Laurentian/Laurentian margins of the Iapetus Ocean during its closure, and is accordingly complex. The complexity arises from the volcanic and tectonic processes that accompanied oceanic closure coupled with the effects of eustatic sea-level changes, including the far-field effects of the Late Ordovician glaciation. For the past three decades, Ordovician successions in Britain and Ireland have been discussed in terms of terranes. Here we review Ordovician successions in each terrane, incorporating the results of recent research and correlating those successions via biostratigraphical schemes and radiometric dates to the global Ordovician series and stages

The Furongian (late Cambrian) Steptoean positive carbon isotope excursion (SPICE) in Avalonia

The Steptoean Positive Carbon Isotope Excursion (SPICE) of earliest late Cambrian (Furongian) age is identified in England. The excursion is found within a c. 145 m thick siliciclastic succession within the middle and higher part of the Outwoods Shale Formation of Warwickshire, and reaches a maximum 13Corg amplitude of 4.1 at values of –25.6. Biostratigraphical data show that the excursion occupies the greater part of the Olenus Biozone, an equivalent of the Glyptagnostus reticulatus Biozone that marks the base of the Furongian and coeval base of the Steptoean in North America. The amplitude of the excursion approaches that recorded in limestone-dominated Laurentian successions, and is greater than that recently documented for organic-rich mudstones of palaeocontinental Baltica in southern Sweden. A minor positive excursion above the SPICE may equate with a similar excursion recognized in Siberia. The SPICE in the Outwoods Shale Formation seems closely linked to the widely recognized early Furongian eustatic sea-level rise. There is no evidence in the English succession for slightly later regression, elsewhere considered coincident with the peak of the excursion and pivotal to some previous models explaining the SPICE. Supplementary material: Analytical results, including total organic carbon (TOC) values for each sample, are available at www.geolsoc.org.uk/SUP18455

The pedigree and influence of fossil collections from the Falkland Islands : from Charles Darwin to continental drift

The Falkland Islands are typical of remote territories in that their early geological exploration was piecemeal and opportunistic. Whilst the resulting fossil collections (dominantly a Devonian fauna of the Malvinokaffric realm) remain the basis for modern interpretations, published accounts misrepresent their extent and provenance. Charles Darwin first discovered fossils during his 1833 visit aboard HMS Beagle, with subsequent British collections acquired in 1842 and 1876, respectively, by the Erebus and Terror and Challenger expeditions and in 1903 by the Scotia expedition. Darwin's collection, and much of the other material, is now held by The Natural History Museum, London (NHM) but some Darwin specimens were assimilated into other collections whilst at least one NHM ‘Darwin’ specimen was not collected by him. There may also be some uncertainty as to the origin of the Scotia collection, now held in Edinburgh by National Museums Scotland, in relation to a contemporary Swedish collection now held in Stockholm. The NHM holdings were supplemented by a number of enigmatic donations from private individuals and then by fossils collected during the first ‘official’ geological survey of the islands in 1920–1922. Meanwhile a large collection was built up in New York through collaboration in 1909 with a local collector – the Governor's wife! The regional associations of the fossils established the African heritage of Falklands geology, and thereby contributed to an understanding of continental drift as the mechanism for the fragmentation of the Gondwana supercontinent. The Falkland Islands are now regarded as a rotated microplate created during the break-up

The Furongian (late Cambrian) Steptoean Positive Carbon Isotope Excursion (SPICE) in Avalonia

<p>The Steptoean Positive Carbon Isotope Excursion (SPICE) of earliest late Cambrian (Furongian) age is identified in England. The excursion is found within a <em>c</em>. 145 m thick siliciclastic succession within the middle and higher part of the Outwoods Shale Formation of Warwickshire, and reaches a maximum δ¹³C_org amplitude of 4.1‰ at values of −25.6‰. Biostratigraphical data show that the excursion occupies the greater part of the <em>Olenus</em> Biozone, an equivalent of the <em>Glyptagnostus reticulatus</em> Biozone that marks the base of the Furongian and coeval base of the Steptoean in North America. The amplitude of the excursion approaches that recorded in limestone-dominated Laurentian successions, and is greater than that recently documented for organic-rich mudstones of palaeocontinental Baltica in southern Sweden. A minor positive excursion above the SPICE may equate with a similar excursion recognized in Siberia. The SPICE in the Outwoods Shale Formation seems closely linked to the widely recognized early Furongian eustatic sea-level rise. There is no evidence in the English succession for slightly later regression, elsewhere considered coincident with the peak of the excursion and pivotal to some previous models explaining the SPICE. </p

A refined chronology for the Cambrian succession of southern Britain

three dated (U–Pb, zircon) ash beds from biostratigraphically constrained Avalonian successions of Shropshire (England) and Pembrokeshire (Wales) delimit the traditional ‘Lower'–‘Middle' Cambrian boundary and resolve a problematic regional correlation. In Shropshire, a date of 514.45 ± 0.36 [0.81 including tracer calibration and 238U decay constant errors] Ma from near the top of the Lower Comley Sandstone Formation provides a maximum age for the boundary between Cambrian Stages 3 and 4, and a date of 509.10 ± 0.22 [0.77 including tracer calibration and 238U decay constant errors] Ma from the basal Quarry Ridge Grits, Upper Comley Sandstone Formation, provides a minimum age for the boundary between Cambrian Stages 4 and 5 (and thus Series 2 and 3). These dates offer a calibration of early metazoan evolution by directly constraining the age of the intervening Comley Limestones, which contain diverse small shelly fossils in addition to trilobites, and also a key early occurrence of exceptional, three-dimensionally preserved arthropods. In Pembrokeshire, an ash bed from the Caerfai Bay Shales Formation dates to 519.30 ± 0.23 [0.77 including tracer calibration and 238U decay constant errors] Ma, equivalent to a horizon low in the Lower Comley Sandstone Formation of Shropshire, possibly around the level at which trilobites make their first local appearance

A refined chronology for the Cambrian succession of southern Britain

<p>Three dated (U–Pb, zircon) ash beds from biostratigraphically constrained Avalonian successions of Shropshire (England) and Pembrokeshire (Wales) delimit the traditional ‘Lower'–‘Middle' Cambrian boundary and resolve a problematic regional correlation. In Shropshire, a date of 514.45 ± 0.36 [0.81 including tracer calibration and ²³⁸U decay constant errors] Ma from near the top of the Lower Comley Sandstone Formation provides a maximum age for the boundary between Cambrian Stages 3 and 4, and a date of 509.10 ± 0.22 [0.77 including tracer calibration and ²³⁸U decay constant errors] Ma from the basal Quarry Ridge Grits, Upper Comley Sandstone Formation, provides a minimum age for the boundary between Cambrian Stages 4 and 5 (and thus Series 2 and 3). These dates offer a calibration of early metazoan evolution by directly constraining the age of the intervening Comley Limestones, which contain diverse small shelly fossils in addition to trilobites, and also a key early occurrence of exceptional, three-dimensionally preserved arthropods. In Pembrokeshire, an ash bed from the Caerfai Bay Shales Formation dates to 519.30 ± 0.23 [0.77 including tracer calibration and ²³⁸U decay constant errors] Ma, equivalent to a horizon low in the Lower Comley Sandstone Formation of Shropshire, possibly around the level at which trilobites make their first local appearance. </p