Graph models of habitat mosaics

Graph theory is a body of mathematics dealing with problems of connectivity, flow, and routing in networks ranging from social groups to computer networks. Recently, network applications have erupted in many fields, and graph models are now being applied in landscape ecology and conservation biology, particularly for applications couched in metapopulation theory. In these applications, graph nodes represent habitat patches or local populations and links indicate functional connections among populations (i.e. via dispersal). Graphs are models of more complicated real systems, and so it is appropriate to review these applications from the perspective of modelling in general. Here we review recent applications of network theory to habitat patches in landscape mosaics. We consider (1) the conceptual model underlying these applications; (2) formalization and implementation of the graph model; (3) model parameterization; (4) model testing, insights, and predictions available through graph analyses; and (5) potential implications for conservation biology and related applications. In general, and for a variety of ecological systems, we find the graph model a remarkably robust framework for applications concerned with habitat connectivity. We close with suggestions for further work on the parameterization and validation of graph models, and point to some promising analytic insights. © 2009 Blackwell Publishing Ltd/CNRS

Conservation and ecology of wetland birds in Africa

Conservation managers worldwide are increasingly faced with the challenges of managing and protecting fragmented landscapes, largely as a consequence of human activities. Over recent decades, ecological theory has made a significant contribution to the development of landscape-scale conservation and practice. However, recommendations accounting for what is practically achievable in the modern-day landscape are currently lacking, while criteria for conservation planning and prioritisation continue to neglect the role of habitat networks at the required spatial scale for the long-term persistence of biodiversity. In this thesis, I test and apply ideas surrounding the complexities of managing and conserving species in a landscape context, using a suite of bird species endemic to papyrus (Cyperus papyrus) swamps in East and Central Africa as a model system. In the face of large-scale habitat loss and degradation, practical measures that account for the fragmented nature of this system, the needs of multiple specialist species, and the reliance on this habitat by local people, are urgently required. I first review the concepts originating from reserve design theory to provide a decision-making framework for those involved in landscape-scale conservation amid 21st century challenges to biodiversity, highlighting the key principles to be considered for informed choices to be made. Second, I show that the needs of local people can be compatible with conservation planning in the tropics, and may play an important part in maintaining habitat quality for species residing in historically disturbed landscapes. Third, I develop a novel framework to make an explicit link between metapopulation dynamics and conservation planning. Despite differences in the patch-level dynamics of individual species, areas of habitat where populations of multiple species are resistant to extinction, and resilient because of high chances of (re)colonization can be identified, highlighting where resources could be invested to ensure species have the capacity to respond to future change. Finally, I simulate the metapopulation dynamics of the papyrus-endemic birds to demonstrate that the optimal conservation strategy for the long-term persistence of all species residing in a network depends on the characteristics of individual species, and the total area that can be protected. Overall, this thesis develops and tests the ecological theory used in spatial conservation planning, emphasising the importance of habitat disturbance and interspecific ecological differences for the effective management of habitat networks. The results increase the evidence base for the conservation of wetland birds in Africa, as well as for species residing in fragmented landscapes more generally.Natural Environment Research Council (NERC) CASE studentship with the RSPB (Royal Society for the Protection of Birds

An Optimization Approach to Improve Equitable Access to Local Parks

Local parks are public resources that promote human and environmental welfare. Unfortunately, park inequities are commonplace as historically marginalized groups may have insufficient access. Platforms exist to identify the geographical areas that would benefit from future park improvements. However, these platforms do not include budget, infrastructure, and environmental considerations that are relevant to park location decisions. To support recreational and government agencies in addressing inequities in the distribution and quality of parks, we propose a mixed-integer program that minimizes insufficient access, defined as weighted deviations across multiple categories. We consider an equity-focused min-max objective and an overall objective to minimize total weighted deviations. We apply the model to a case study of Asheville, North Carolina. We conduct extensive data collection to parameterize the model. In policy analyses, we consider the effects of available budget, planning horizons, strategic demographic priorities, and thresholds of access. The model reflects user-defined criteria and goals, and the results suggest that the framework may be generalizable to other cities. This study serves as the first step in the development and incorporation of mathematical modeling to achieve social goals within the recreational setting

Methodical basis for landscape structure analysis and monitoring: inclusion of ecotones and small landscape elements

Habitat variation is considered as an expression of biodiversity at landscape level in addition to genetic variation and species variation. Thus, effective methods for measuring habitat pattern at landscape level can be used to evaluate the status of biological conservation. However, the commonly used model (i.e. patch-corridor-matrix) for spatial pattern analysis has deficiencies. This model assumes discrete structures within the landscape without explicit consideration of “transitional zones” or “gradients” between patches. The transitional zones, often called “ecotones”, are dynamic and have a profound influence on adjacent ecosystems. Besides, this model takes landscape as a flat surface without consideration of the third spatial dimension (elevation). This will underestimate the patches’ size and perimeter as well as distances between patches especially in mountainous regions. Thus, the mosaic model needs to be adapted for more realistic and more precise representation of habitat pattern regarding to biodiversity assessment. Another part of information that has often been ignored is “small biotopes” inside patches (e.g. hedgerows, tree rows, copse, and scattered trees), which leads to within-patch heterogeneity being underestimated. The present work originates from the integration of the third spatial dimension in land-cover classification and landscape structure analysis. From the aspect of data processing, an integrated approach of Object-Based Image Analysis (OBIA) and Pixel-Based Image Analysis (PBIA) is developed and applied on multi-source data set (RapidEye images and Lidar data). At first, a general OBIA procedure is developed according to spectral object features based on RapidEye images for producing land-cover maps. Then, based on the classified maps, pixel-based algorithms are designed for detection of the small biotopes and ecotones using a Normalized Digital Surface Model (NDSM) which is derived from Lidar data. For describing habitat pattern under three-dimensional condition, several 3D-metrics (measuring e.g. landscape diversity, fragmentation/connectivity, and contrast) are proposed with spatial consideration of the ecological functions of small biotopes and ecotones. The proposed methodology is applied in two real-world examples in Germany and China. The results are twofold. First, it shows that the integrated approach of object-based and pixel-based image processing is effective for land-cover classification on different spatial scales. The overall classification accuracies of the main land-cover maps are 92 % in the German test site and 87 % in the Chinese test site. The developed Red Edge Vegetation Index (REVI) which is calculated from RapidEye images has been proved more efficient than the traditionally used Normalized Differenced Vegetation Index (NDVI) for vegetation classification, especially for the extraction of the forest mask. Using NDSM data, the third dimension is helpful for the identification of small biotopes and height gradient on forest boundary. The pixel-based algorithm so-called “buffering and shrinking” is developed for the detection of tree rows and ecotones on forest/field boundary. As a result the accuracy of detecting small biotopes is 80 % and four different types of ecotones are detected in the test site. Second, applications of 3D-metrics in two varied test sites show the frequently-used landscape diversity indices (i.e. Shannon’s diversity (SHDI) and Simpson’s diversity (SIDI)) are not sufficient for describing the habitats diversity, as they quantify only the habitats composition without consideration on habitats spatial distribution. The modified 3D-version of Effective Mesh Size (MESH) that takes ecotones into account leads to a realistic quantification of habitat fragmentation. In addition, two elevation-based contrast indices (i.e. Area-Weighted Edge Contrast (AWEC) and Total Edge Contrast Index (TECI)) are used as supplement to fragmentation metrics. Both ecotones and small biotopes are incorporated into the contrast metrics to take into account their edge effect in habitat pattern. This can be considered as a further step after fragmentation analysis with additional consideration of the edge permeability in the landscape structure analysis. Furthermore, a vector-based algorithm called “multi-buffer” approach is suggested for analyzing ecological networks based on land-cover maps. It considers small biotopes as stepping stones to establish connections between patches. Then, corresponding metrics (e.g. Effective Connected Mesh Size (ECMS)) are proposed based on the ecological networks. The network analysis shows the response of habitat connectivity to different dispersal distances in a simple way. Those connections through stepping stones act as ecological indicators of the “health” of the system, indicating the interpatch communications among habitats. In summary, it can be stated that habitat diversity is an essential level of biodiversity and methods for quantifying habitat pattern need to be improved and adapted to meet the demands for landscape monitoring and biodiversity conservation. The approaches presented in this work serve as possible methodical solution for fine-scale landscape structure analysis and function as “stepping stones” for further methodical developments to gain more insights into the habitat pattern.Die Lebensraumvielfalt ist neben der genetischen Vielfalt und der Artenvielfalt eine wesentliche Ebene der Biodiversität. Da diese Ebenen miteinander verknüpft sind, können Methoden zur Messung der Muster von Lebensräumen auf Landschaftsebene erfolgreich angewandt werden, um den Zustand der Biodiversität zu bewerten. Das zur räumlichen Musteranalyse auf Landschaftsebene häufig verwendete Patch-Korridor-Matrix-Modell weist allerdings einige Defizite auf. Dieses Modell geht von diskreten Strukturen in der Landschaft aus, ohne explizite Berücksichtigung von „Übergangszonen“ oder „Gradienten“ zwischen den einzelnen Landschaftselementen („Patches“). Diese Übergangszonen, welche auch als „Ökotone“ bezeichnet werden, sind dynamisch und haben einen starken Einfluss auf benachbarte Ökosysteme. Außerdem wird die Landschaft in diesem Modell als ebene Fläche ohne Berücksichtigung der dritten räumlichen Dimension (Höhe) betrachtet. Das führt dazu, dass die Flächengrößen und Umfänge der Patches sowie Distanzen zwischen den Patches besonders in reliefreichen Regionen unterschätzt werden. Daher muss das Patch-Korridor-Matrix-Modell für eine realistische und präzise Darstellung der Lebensraummuster für die Bewertung der biologischen Vielfalt angepasst werden. Ein weiterer Teil der Informationen, die häufig in Untersuchungen ignoriert werden, sind „Kleinbiotope“ innerhalb größerer Patches (z. B. Feldhecken, Baumreihen, Feldgehölze oder Einzelbäume). Dadurch wird die Heterogenität innerhalb von Patches unterschätzt. Die vorliegende Arbeit basiert auf der Integration der dritten räumlichen Dimension in die Landbedeckungsklassifikation und die Landschaftsstrukturanalyse. Mit Methoden der räumlichen Datenverarbeitung wurde ein integrierter Ansatz von objektbasierter Bildanalyse (OBIA) und pixelbasierter Bildanalyse (PBIA) entwickelt und auf einen Datensatz aus verschiedenen Quellen (RapidEye-Satellitenbilder und Lidar-Daten) angewendet. Dazu wird zunächst ein OBIA-Verfahren für die Ableitung von Hauptlandbedeckungsklassen entsprechend spektraler Objekteigenschaften basierend auf RapidEye-Bilddaten angewandt. Anschließend wurde basierend auf den klassifizierten Karten, ein pixelbasierter Algorithmus für die Erkennung von kleinen Biotopen und Ökotonen mit Hilfe eines normalisierten digitalen Oberflächenmodells (NDSM), welches das aus LIDAR-Daten abgeleitet wurde, entwickelt. Zur Beschreibung der dreidimensionalen Charakteristika der Lebensraummuster unter der räumlichen Betrachtung der ökologischen Funktionen von kleinen Biotopen und Ökotonen, werden mehrere 3D-Maße (z. B. Maße zur landschaftlichen Vielfalt, zur Fragmentierung bzw. Konnektivität und zum Kontrast) vorgeschlagen. Die vorgeschlagene Methodik wird an zwei realen Beispielen in Deutschland und China angewandt. Die Ergebnisse zeigen zweierlei. Erstens zeigt es sich, dass der integrierte Ansatz der objektbasierten und pixelbasierten Bildverarbeitung effektiv für die Landbedeckungsklassifikation auf unterschiedlichen räumlichen Skalen ist. Die Klassifikationsgüte insgesamt für die Hauptlandbedeckungstypen beträgt 92 % im deutschen und 87 % im chinesischen Testgebiet. Der eigens entwickelte Red Edge-Vegetationsindex (REVI), der sich aus RapidEye-Bilddaten berechnen lässt, erwies sich für die Vegetationsklassifizierung als effizienter verglichen mit dem traditionell verwendeten Normalized Differenced Vegetation Index (NDVI), insbesondere für die Gewinnung der Waldmaske. Im Rahmen der Verwendung von NDSM-Daten erwies sich die dritte Dimension als hilfreich für die Identifizierung von kleinen Biotopen und dem Höhengradienten, beispielsweise an der Wald/Feld-Grenze. Für den Nachweis von Baumreihen und Ökotonen an der Wald/Feld-Grenze wurde der sogenannte pixelbasierte Algorithmus „Pufferung und Schrumpfung“ entwickelt. Im Ergebnis konnten kleine Biotope mit einer Genauigkeit von 80 % und vier verschiedene Ökotontypen im Testgebiet detektiert werden. Zweitens zeigen die Ergebnisse der Anwendung der 3D-Maße in den zwei unterschiedlichen Testgebieten, dass die häufig genutzten Landschaftsstrukturmaße Shannon-Diversität (SHDI) und Simpson-Diversität (SIDI) nicht ausreichend für die Beschreibung der Lebensraumvielfalt sind. Sie quantifizieren lediglich die Zusammensetzung der Lebensräume, ohne Berücksichtigung der räumlichen Verteilung und Anordnung. Eine modifizierte 3D-Version der Effektiven Maschenweite (MESH), welche die Ökotone integriert, führt zu einer realistischen Quantifizierung der Fragmentierung von Lebensräumen. Darüber hinaus wurden zwei höhenbasierte Kontrastindizes, der flächengewichtete Kantenkontrast (AWEC) und der Gesamt-Kantenkontrast Index (TECI), als Ergänzung der Fragmentierungsmaße entwickelt. Sowohl Ökotone als auch Kleinbiotope wurden in den Berechnungen der Kontrastmaße integriert, um deren Randeffekte im Lebensraummuster zu berücksichtigen. Damit kann als ein weiterer Schritt nach der Fragmentierungsanalyse die Randdurchlässigkeit zusätzlich in die Landschaftsstrukturanalyse einbezogen werden. Außerdem wird ein vektorbasierter Algorithmus namens „Multi-Puffer“-Ansatz für die Analyse von ökologischen Netzwerken auf Basis von Landbedeckungskarten vorgeschlagen. Er berücksichtigt Kleinbiotope als Trittsteine, um Verbindungen zwischen Patches herzustellen. Weiterhin werden entsprechende Maße, z. B. die Effective Connected Mesh Size (ECMS), für die Analyse der ökologischen Netzwerke vorgeschlagen. Diese zeigen die Auswirkungen unterschiedlicher angenommener Ausbreitungsdistanzen von Organismen bei der Ableitung von Biotopverbundnetzen in einfacher Weise. Diese Verbindungen zwischen Lebensräumen über Trittsteine hinweg dienen als ökologische Indikatoren für den „gesunden Zustand“ des Systems und zeigen die gegenseitigen Verbindungen zwischen den Lebensräumen. Zusammenfassend kann gesagt werden, dass die Vielfalt der Lebensräume eine wesentliche Ebene der Biodiversität ist. Die Methoden zur Quantifizierung der Lebensraummuster müssen verbessert und angepasst werden, um den Anforderungen an ein Landschaftsmonitoring und die Erhaltung der biologischen Vielfalt gerecht zu werden. Die in dieser Arbeit vorgestellten Ansätze dienen als mögliche methodische Lösung für eine feinteilige Landschaftsstrukturanalyse und fungieren als ein „Trittsteine” auf dem Weg zu weiteren methodischen Entwicklungen für einen tieferen Einblick in die Muster von Lebensräumen

Micro-topography associated to forest edges

Forest edges are often defined as the discontinuity between the forest habitat and an adjacent open habitat, thus they are based on a clear difference in the structure of the dominant vegetation. However, beside this very general definition, in the field we can observe a large diversity of edges, with often different kinds of micro-topography features: bank, ditch, stone wall, path, etc. As these elements are rather common in many temperate forest edges, it seems important to start to characterize them more clearly and with consistency. From a set of observations in south-western France, we build a first typology of the micro-topographic elements associated to forest edges. For each of them we describe the process, natural or human induced, at their origin, and according to the literature available, we identify some of their key ecological roles. Banks, generated by the differential erosion between forest and crops along slopes, are especially analyzed since they are the most common micro-topographic element in our region. It offers many micro-habitat conditions in the soil used by a wide range of species, notably by several bee species. More research is required to study in details the importance of such micro-topographic elements

Ecological Status and Importance of Grasslands (Vidis) in Conservation of Avian

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Linking and Harmonizing Forest Spatial Pattern Analyses at European, National and Regional Scales for a Better Characterization of Forest Vulnerability and Resilience

The project ¿Linking and harmonizing the forests spatial pattern analyses at European, national and regional scales for a better characterization of the forests vulnerability and resilience¿ (JRC contract 382391 F1SC) covers one of the seven topics to be studied in the frame of the Regulation (EC) 2152/2003 on the monitoring of forest and environmental interactions, the so-called "Forest Focus" Regulation. It first demonstrates the application of the mathematical morphology based forest spatial pattern analysis tool developed at the Joint Research Centre (GUIDOS); it then develops one index related to potential forest vulnerability, resistance and resilience (FVRR index) on the basis of multi-criteria fuzzy modelling technics. Its final aim is to address linkages between forest spatial pattern and forest ecological functionality with emphasis on forest vulnerability, resistance, resilience. This study was conducted by a European consortium coordinated by the University of Molise (Italy) and included partners from the University of Hamburg (Germany), the European Forest Institute (Finland) and the Forest Research (United Kingdom). The overall supervision of the project and the processing of forest spatial pattern were done by the Joint Research Centre.JRC.DDG.H.7-Land management and natural hazard