The fossil phase in the life of a galaxy group

We investigate the origin and evolution of fossil groups in a concordance ΛCDM cosmological simulation. We consider haloes with masses between 1 × 1013 and 5 × 1013h−1M⊙, and study the physical mechanisms that lead to the formation of the large gap in magnitude between the brightest and the second most bright group member, which is typical for these fossil systems. Fossil groups are found to have high dark matter concentrations, which we can relate to their early formation time. The large magnitude gaps arise after the groups have built up half of their final mass, due to merging of massive group members. We show that the existence of fossil systems is primarily driven by the relatively early infall of massive satellites, and that we do not find a strong environmental dependence for these systems. In addition, we find tentative evidence for fossil group satellites falling in on orbits with typically lower angular momentum, which might lead to a more efficient merger on to the host. We find a population of groups at higher redshifts that go through a ‘fossil phase': a stage where they show a large magnitude gap, which is terminated by renewed infall from their environmen

The Fossil Phase in the Life of a Galaxy Group

We investigate the origin and evolution of fossil groups in a concordance LCDM cosmological simulation. We consider haloes with masses between (1-5)\times10^{13} \hMsun and study the physical mechanisms that lead to the formation of the large gap in magnitude between the brightest and the second most bright group member, which is typical for these fossil systems. Fossil groups are found to have high dark matter concentrations, which we can relate to their early formation time. The large magnitude-gaps arise after the groups have build up half of their final mass, due to merging of massive group members. We show that the existence of fossil systems is primarily driven by the relatively early infall of massive satellites, and that we do not find a strong environmental dependence for these systems. In addition, we find tentative evidence for fossil group satellites falling in on orbits with typically lower angular momentum, which might lead to a more efficient merger onto the host. We find a population of groups at higher redshifts that go through a ``fossil phase'': a stage where they show a large magnitude-gap, which is terminated by renewed infall from their environment.Comment: 9 pages and 8 figures, submitted to MNRA

Separating underwater ambient noise from flow noise recorded on stereo acoustic tags attached to marine mammals

A.M.v.B.B. and P.B. were funded by The Netherlands Ministry of Defence. Fieldwork efforts and support for P.M. and F.S. was provided by the US Office of Naval Research [award numbers N00014-08-1-0984 and N00014-10-1-0355]. P.W. received a PhD studentship with matched funding from The Netherlands Ministry of Defence (administered by The Netherlands Organisation for Applied Scientific Research, TNO) and UK Natural Environment Research Council [NE/J500276/1].Sound-recording acoustic tags attached to marine animals are commonly used in behavioural studies. Measuring ambient noise is of interest to efforts to understand responses of marine mammals to anthropogenic underwater sound, or to assess their communication space. Noise of water flowing around the tag reflects the speed of the animal, but hinders ambient noise measurement. Here, we describe a correlation-based method for stereo acoustic tags to separate the relative contributions of flow and ambient noise. The uncorrelated part of the noise measured in digital acoustic recording tag (DTAG) recordings related well to swim speed of a humpback whale (Megaptera novaeangliae), thus providing a robust measure of flow noise over a wide frequency bandwidth. By removing measurements affected by flow noise, consistent ambient noise estimates were made for two killer whales (Orcinus orca) with DTAGs attached simultaneously. The method is applicable to any multi-channel acoustic tag, enabling application to a wide range of marine species.Publisher PDFPeer reviewe

How effectively do horizontal and vertical response strategies of long-finned pilot whales reduce sound exposure from naval sonar?

PJW was supported with studentships of The Netherlands Ministry of Defence (grant number 032.30370/01.02) and the VSB Foundation (grant number VSB.08/228-E) and Ren e Dekeling is acknowledged for making funding possible. The 3S project was supported by the US Office of Naval Research, The Netherlands Ministry of Defence, Royal Norwegian Navy and Norwegian Ministry of Defence, and by World Wildlife Fund Norway. PLT received funding from the MASTS pooling initiative (The Marine Alliance for Science and Technology for Scotland) and their support is gratefully acknowledged.The behaviour of a marine mammal near a noise source can modulate the sound exposure it receives. We demonstrate that two long-finned pilot whales both surfaced in synchrony with consecutive arrivals of multiple sonar pulses. We then assess the effect of surfacing and other behavioural response strategies on the received cumulative sound exposure levels and maximum sound pressure levels (SPLs) by modelling realistic spatiotemporal interactions of a pilot whale with an approaching source. Under the propagation conditions of our model, some response strategies observed in the wild were effective in reducing received levels (e.g. movement perpendicular to the source's line of approach), but others were not (e.g. switching from deep to shallow diving; synchronous surfacing after maximum SPLs). Our study exemplifies how simulations of source-whale interactions guided by detailed observational data can improve our understanding about motivations behind behaviour responses observed in the wild (e.g., reducing sound exposure, prey movement).PostprintPeer reviewe

First indications that northern bottlenose whales are sensitive to behavioural disturbance from anthropogenic noise

-Although northern bottlenose whales were the most heavily hunted beaked whale, we have little information about this species in its remote habitat of the North Atlantic Ocean. Underwater anthropogenic noise and disruption of their natural habitat may be major threats, given the sensitivity of other beaked whales to such noise disturbance. We attached dataloggers to 13 northern bottlenose whales and compared their natural sounds and movements to those of one individual exposed to escalating levels of 1–2 kHz upsweep naval sonar signals. At a received sound pressure level (SPL) of 98 dB re 1 μPa, the whale turned to approach the sound source, but at a received SPL of 107 dB re 1 μPa, the whale began moving in an unusually straight course and then made a near 180° turn away from the source, and performed the longest and deepest dive (94 min, 2339 m) recorded for this species. Animal movement parameters differed significantly from baseline for more than 7 h until the tag fell off 33–36 km away. No clicks were emitted during the response period, indicating cessation of normal echolocation-based foraging. A sharp decline in both acoustic and visual detections of conspecifics after exposure suggests other whales in the area responded similarly. Though more data are needed, our results indicate high sensitivity of this species to acoustic disturbance, with consequent risk from marine industrialization and naval activity

Northern bottlenose whales in a pristine environment respond strongly to close and distant navy sonar signals

The research described in this paper was supported by US Office of Naval Research (ONR grants N00014-15-1-2533 and N00014-16-1-3059), US Strategic Environmental Research and Development Program (SERDP award RC-2337), the French Ministry of Defence (DGA) and the Netherlands Ministry of Defence. P.L.T. acknowledges support from the MASTS pooling initiative (Marine Alliance for Science and Technology for Scotland; supported by the Scottish Funding Council, grant reference HR09011, and contributing institutions). All relevant data and scripts are within the paper and its electronic supplementary material, or available from the Dryad Digital Repository: https://doi.org/10.5061/dryad.fc5c0s4Impact assessments for sonar operations typically use received sound levels to predict behavioural disturbance in marine mammals. However, there are indications that cetaceans may learn to associate exposures from distant sound sources with lower perceived risk. To investigate the roles of source distance and received level in an area without frequent sonar activity, we conducted multi-scale controlled exposure experiments (n = 3) with 12 northern bottlenose whales near Jan Mayen, Norway. Animals were tagged with high-resolution archival tags (n = 1 per experiment) or medium-resolution satellite tags (n = 9 in total) and subsequently exposed to sonar. We also deployed bottom-moored recorders to acoustically monitor for whales in the exposed area. Tagged whales initiated avoidance of the sound source over a wide range of distances (0.8–28 km), with responses characteristic of beaked whales. Both onset and intensity of response were better predicted by received sound pressure level (SPL) than by source distance. Avoidance threshold SPLs estimated for each whale ranged from 117–126 dB re 1 µPa, comparable to those of other tagged beaked whales. In this pristine underwater acoustic environment, we found no indication that the source distances tested in our experiments modulated the behavioural effects of sonar, as has been suggested for locations where whales are frequently exposed to sonar.Publisher PDFPeer reviewe

Modelling the broadband propagation of marine mammal echolocation clicks for click-based population density estimates

Funding: U.S. Office of Naval Research (ONR Grant No. N00014-14-1-0409); P.L.T. acknowledges funding received from the MASTS pooling initiative (The Marine Alliance for Science and Technology for Scotland). MASTS is funded by the Scottish Funding Council (Grant No. HR09011) and contributing institutions.Passive acoustic monitoring with widely-dispersed hydrophones has been suggested as a cost-effective method to monitor population densities of echolocating marine mammals. This requires an estimate of the area around each receiver over which vocalizations are detected—the “effective detection area” (EDA). In the absence of auxiliary measurements enabling estimation of the EDA, it can be modelled instead. Common simplifying model assumptions include approximating the spectrum of clicks by flat energy spectra, and neglecting the frequency-dependence of sound absorption within the click bandwidth (narrowband assumption), rendering the problem amenable to solution using the sonar equation. Here, it is investigated how these approximations affect the estimated EDA and their potential for biasing the estimated density. EDA was estimated using the passive sonar equation, and by applying detectors to simulated clicks injected into measurements of background noise. By comparing model predictions made using these two approaches for different spectral energy distributions of echolocation clicks, but identical click source energy level and detector settings, EDA differed by up to a factor of 2 for Blainville's beaked whales. Both methods predicted relative density bias due to narrowband assumptions ranged from 5% to more than 100%, depending on the species, detector settings, and noise conditions.Publisher PDFPeer reviewe

Modelling the broadband propagation of marine mammal echolocation clicks for click-based population density estimates

Passive acoustic monitoring with widely-dispersed hydrophones has been suggested as a cost-effective method to monitor population densities of echolocating marine mammals. This requires an estimate of the area around each receiver over which vocalizations are detected—the “effective detection area” (EDA). In the absence of auxiliary measurements enabling estimation of the EDA, it can be modelled instead. Common simplifying model assumptions include approximating the spectrum of clicks by flat energy spectra, and neglecting the frequency-dependence of sound absorption within the click bandwidth (narrowband assumption), rendering the problem amenable to solution using the sonar equation. Here, it is investigated how these approximations affect the estimated EDA and their potential for biasing the estimated density. EDA was estimated using the passive sonar equation, and by applying detectors to simulated clicks injected into measurements of background noise. By comparing model predictions made using these two approaches for different spectral energy distributions of echolocation clicks, but identical click source energy level and detector settings, EDA differed by up to a factor of 2 for Blainville's beaked whales. Both methods predicted relative density bias due to narrowband assumptions ranged from 5% to more than 100%, depending on the species, detector settings, and noise conditions