1,773 research outputs found

    Use of Remote Sensing for the Assessment of Farmland Biodiversity

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    In many countries farmland represent the most relevant type of land use. The conservation and the enhancement of biodiversity within these areas is primary objective for the achievement of sustainable management of agro-ecosystems and the level of biodiversity can be used as agri-environmental indicator. The measure of biodiversity over larger territories however, can be very expensive and time consuming. Earth observations have a rich history in characterizing and monitoring biodiversity and ecosystems. The possibility to use remotely sensed data for monitoring biodiversity was identified at the dawn of Earth observations and has been widely used for decades. Remotely sensed data can cost-effectively track changes in ecosystem distribution and status. The new generation of high-spatial and spectral resolution offers more hope for more direct monitoring of habitats and species. Recent developments in satellite remote sensing and in geographic information system (GIS) combined with user oriented computer programs allow us to use ecological principles for biodiversity characterization at landscape level more efficiently. This paper presents an approach for biodiversity characterization at landscape level using geospatial techniques. The present study hypothesizes that the complexity of the landscape is related to biological phytodiversity, and in turn phytodiversity can be considered a proxy of the overall biodiversity. The aim of the study is to estimate the farmland landscape complexity by using a remote sensing based approach. The approach has been validated in a Northern Italian agricultural area (Emilia-Romagna region). The proposed approach can provide policy makers and land use planners valuable, updated and low cost information on farmland landscape complexity and an estimate of its biodiversity.JRC.DDG.H.7 - Land management and natural hazard

    FIDELIO 2: Overview and theoretical foundations of the second version of the Fully Interregional Dynamic Econometric Long-term Input-Output model for the EU-27

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    This report serves as an update of "FIDELIO 1: Fully Interregional Dynamic Econometric Long-term Input-Output model for the EU27" by Kratena et al. (2013), i.e. the manual of the first version of the FIDELIO model. FIDELIO fits into the generation of macroeconomic multi-sectoral input-output (IO) models whose earliest contributions include the Cambridge MDM (Barker, 1976) and the INFORUM (Almon et al., 1974) models for the UK and the US, respectively. Such econometric IO models have grown over time in terms of complexity and scope and are used for macroeconomic modelling purposes alongside other types of general equilibrium models (including DSGE ones). This report explores the theoretical foundations of the latest version of the model, FIDELIO 2 (which has being developed between 2014 and 2016), and contains a description of its main features. With respect to its initial version, the model has been extended in a number of ways. For instance, and without entering into detail at this stage, seven non-EU countries are now included in the model (Brazil, China, India, Japan, Russia, Turkey and the USA) in addition to the 27 EU countries already included in the first version; both trade and household final demand are now modelled in a considerably more complex way than before; there is an environmental block dealing with greenhouse gas (GHG) emissions; and the base year is 2007 rather than 2005. Thus, it was deemed necessary to present all the new model characteristics in an organic way via the present technical report. The remainder of this report is organised as follows: Section 1 provides a concise macro-overview of FIDELIO 2 which relies very much on the first section of the FIDELIO 1 manual by Kratena et al. (2013). Section 2 presents the economic theories underlying the core blocks of FIDELIO 2. This report serves two main purposes. First, it is an adequate resource for the readers who are interested in the model's main features. Second, it facilitates the process of understanding all the details of FIDELIO 2 for those who want to learn the logic and the theory behind its construction. Such readers are expected to grasp the general structure of the model by reading Section 1, helped by the overview of the model's main economic flows contained in Figure 1. Then, Section 2 goes through the theoretical foundations of the various model blocks.JRC.B.5-Circular Economy and Industrial Leadershi

    Simple Screened Hydrogen Model of Excitons in Two-Dimensional Materials

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    We present a generalized hydrogen model for the binding energies (EBE_B) of excitons in two-dimensional (2D) materials that sheds light on the fundamental differences between excitons in two and three dimensions. In contrast to the well-known hydrogen model of three-dimensional (3D) excitons, the description of 2D excitons is complicated by the fact that the screening cannot be assumed to be local. We show that one can consistently define an effective 2D dielectric constant by averaging the screening over the extend of the exciton. For an ideal 2D semiconductor this leads to a simple expression for EBE_B that only depends on the excitonic mass and the 2D polarizability α\alpha. The model is shown to produce accurate results for 51 transition metal dichalcogenides. Remarkably, over a wide range of polarizabilities the expression becomes independent of the mass and we obtain EB2D3/(4πα)E_B^{2D}\approx3/(4\pi\alpha), which explains the recently observed linear scaling of exciton binding energies with band gap. It is also shown that the model accurately reproduces the non-hydrogenic Rydberg series in WS2_2 and can account for screening from the environment.Comment: 5 page

    The importance of immune gene variability (MHC) in evolutionary ecology and conservation

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    Genetic studies have typically inferred the effects of human impact by documenting patterns of genetic differentiation and levels of genetic diversity among potentially isolated populations using selective neutral markers such as mitochondrial control region sequences, microsatellites or single nucleotide polymorphism (SNPs). However, evolutionary relevant and adaptive processes within and between populations can only be reflected by coding genes. In vertebrates, growing evidence suggests that genetic diversity is particularly important at the level of the major histocompatibility complex (MHC). MHC variants influence many important biological traits, including immune recognition, susceptibility to infectious and autoimmune diseases, individual odours, mating preferences, kin recognition, cooperation and pregnancy outcome. These diverse functions and characteristics place genes of the MHC among the best candidates for studies of mechanisms and significance of molecular adaptation in vertebrates. MHC variability is believed to be maintained by pathogen-driven selection, mediated either through heterozygote advantage or frequency-dependent selection. Up to now, most of our knowledge has derived from studies in humans or from model organisms under experimental, laboratory conditions. Empirical support for selective mechanisms in free-ranging animal populations in their natural environment is rare. In this review, I first introduce general information about the structure and function of MHC genes, as well as current hypotheses and concepts concerning the role of selection in the maintenance of MHC polymorphism. The evolutionary forces acting on the genetic diversity in coding and non-coding markers are compared. Then, I summarise empirical support for the functional importance of MHC variability in parasite resistance with emphasis on the evidence derived from free-ranging animal populations investigated in their natural habitat. Finally, I discuss the importance of adaptive genetic variability with respect to human impact and conservation, and implications for future studies

    On initial conditions for the Hot Big Bang

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    We analyse the process of reheating the Universe in the electroweak theory where the Higgs field plays a role of the inflaton. We estimate the maximal temperature of the Universe and fix the initial conditions for radiation-dominated phase of the Universe expansion in the framework of the Standard Model (SM) and of the nuMSM -- the minimal extension of the SM by three right-handed singlet fermions. We show that the inflationary epoch is followed by a matter dominated stage related to the Higgs field oscillations. We investigate the energy transfer from Higgs-inflaton to the SM particles and show that the radiation dominated phase of the Universe expansion starts at temperature T_r~(3-15)*10^{13} GeV, where the upper bound depends on the Higgs boson mass. We estimate the production rate of singlet fermions at preheating and find that their concentrations at T_r are negligibly small. This suggests that the sterile neutrino Dark Matter (DM) production and baryogenesis in the nuMSM with Higgs-driven inflation are low energy phenomena, having nothing to do with inflation. We study then a modification of the nuMSM, adding to its Lagrangian higher dimensional operators suppressed by the Planck scale. The role of these operators in Higgs-driven inflation is clarified. We find that these operators do not contribute to the production of Warm Dark Matter (WDM) and to baryogenesis. We also demonstrate that the sterile neutrino with mass exceeding 100 keV (a Cold Dark Matter (CDM) candidate) can be created during the reheating stage of the Universe in necessary amounts. We argue that the mass of DM sterile neutrino should not exceed few MeV in order not to overclose the Universe.Comment: 41 pages, 5 figures. Journal version accepted in JCA

    Validation of internal reference genes for quantitative real-time PCR in a non-model organism, the yellow-necked mouse, Apodemus flavicollis

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    <p>Abstract</p> <p>Background</p> <p>Reference genes are used as internal standards to normalize mRNA abundance in quantitative real-time PCR and thereby allow a direct comparison between samples. So far most of these expression studies used human or classical laboratory model species whereas studies on non-model organism under in-situ conditions are quite rare. However, only studies in free-ranging populations can reveal the effects of natural selection on the expression levels of functional important genes. In order to test the feasibility of gene expression studies in wildlife samples we transferred and validated potential reference genes that were developed for lab mice (<it>Mus musculus</it>) to samples of wild yellow-necked mice, <it>Apodemus flavicollis</it>. The stability and suitability of eight potential reference genes was accessed by the programs BestKeeper, NormFinder and geNorm.</p> <p>Findings</p> <p>Although the three programs used different algorithms the ranking order of reference genes was significantly concordant and geNorm differed in only one, NormFinder in two positions compared to BestKeeper. The genes ordered by their mean rank from the most to the least stable gene were: <it>Rps18</it>, <it>Sdha</it>, <it>Canx</it>, <it>Actg1</it>, <it>Pgk1</it>, <it>Ubc</it>, <it>Rpl13a </it>and <it>Actb</it>. Analyses of the normalization factor revealed best results when the five most stable genes were included for normalization.</p> <p>Discussion</p> <p>We established a SYBR green qPCR assay for liver samples of wild <it>A. flavicollis </it>and conclude that five genes should be used for appropriate normalization. Our study provides the basis to investigate differential expression of genes under selection under natural selection conditions in liver samples of <it>A. flavicollis</it>. This approach might also be applicable to other non-model organisms.</p

    TRPV1-expressing primary afferents generate behavioral responses to pruritogens via multiple mechanisms

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    The mechanisms that generate itch are poorly understood at both the molecular and cellular levels despite its clinical importance. To explore the peripheral neuronal mechanisms underlying itch, we assessed the behavioral responses (scratching) produced by s.c. injection of various pruritogens in PLCβ3- or TRPV1-deficient mice. We provide evidence that at least 3 different molecular pathways contribute to the transduction of itch responses to different pruritogens: 1) histamine requires the function of both PLCβ3 and the TRPV1 channel; 2) serotonin, or a selective agonist, α-methyl-serotonin (α-Me-5-HT), requires the presence of PLCβ3 but not TRPV1, and 3) endothelin-1 (ET-1) does not require either PLCβ3 or TRPV1. To determine whether the activity of these molecules is represented in a particular subpopulation of sensory neurons, we examined the behavioral consequences of selectively eliminating 2 nonoverlapping subsets of nociceptors. The genetic ablation of MrgprD^+ neurons that represent ≈90% of cutaneous nonpeptidergic neurons did not affect the scratching responses to a number of pruritogens. In contrast, chemical ablation of the central branch of TRPV1+ nociceptors led to a significant behavioral deficit for pruritogens, including α-Me-5-HT and ET-1, that is, the TRPV1-expressing nociceptor was required, whether or not TRPV1 itself was essential. Thus, TRPV1 neurons are equipped with multiple signaling mechanisms that respond to different pruritogens. Some of these require TRPV1 function; others use alternate signal transduction pathways

    Re-visiting the relations: Galactic thin disc age-velocity dispersion relation

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    The velocity dispersion of stars in the solar neighbourhood thin disc increases with time after star formation. Nordstrom et al. (2004) is the most recent observational attempt to constrain the age-velocity dispersion relation. They fitted the age-velocity dispersion relations of each Galactic cardinal direction space velocity component, U (towards the Galactic centre), V (in the direction of Galactic rotation) and W (towards the North Galactic Pole), with power laws and interpreted these as evidence for continuous heating of the disc in all directions throughout its lifetime. We re-visit these relations with their data and use Famaey et al. (2005) to show that structure in the local velocity distribution function distorts the in-plane (U and V) velocity distributions away from Gaussian so that a dispersion is not an adequate parametrization of their functions. The age-sigma(W) relation can however be constrained because the sample is well phase-mixed vertically. We do not find any local signature of the stellar warp in the Galactic disc. Vertical disc heating does not saturate at an early stage. Our new result is that a power law is not required by the data: disc heating models that saturate after ~ 4.5 Gyr are equally consistent with observations.Comment: Accepted for publication in MNRAS, 24 pages, 20 figure
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