67,000 years of coastal engagement at Panga ya Saidi, eastern Africa

The antiquity and nature of coastal resource procurement is central to understanding human evolution and adaptations to complex environments. It has become increasingly apparent in global archaeological studies that the timing, characteristics, and trajectories of coastal resource use are highly variable. Within Africa, discussions of these issues have largely been based on the archaeological record from the south and northeast of the continent, with little evidence from eastern coastal areas leaving significant spatial and temporal gaps in our knowledge. Here, we present data from Panga ya Saidi, a limestone cave complex located 15 km from the modern Kenyan coast, which represents the first long-term sequence of coastal engagement from eastern Africa. Rather than attempting to distinguish between coastal resource use and coastal adaptations, we focus on coastal engagement as a means of characterising human relationships with marine environments and resources from this inland location. We use aquatic mollusc data spanning the past 67,000 years to document shifts in the acquisition, transportation, and discard of these materials, to better understand long-term trends in coastal engagement. Our results show pulses of coastal engagement beginning with low-intensity symbolism, and culminating in the consistent low-level transport of marine and freshwater food resources, emphasising a diverse relationship through time. Panga ya Saidi has the oldest stratified evidence of marine engagement in eastern Africa, and is the only site in Africa which documents coastal resources from the Late Pleistocene through the Holocene, highlighting the potential archaeological importance of peri-coastal sites to debates about marine resource relationships.Introduction Site location and description Materials and methods Results - PYS mollusc assemblage characteristics - Comparative trends in mollusc discard Discussion Conclusio

Iron Age hunting and herding in coastal eastern Africa: ZooMS identification of domesticates and wild bovids at Panga ya Saidi, Kenya

The morphological differentiation of African bovids in highly fragmented zooarchaeological assemblages is a major hindrance to reconstructing the nature and spread of pastoralism in sub-Saharan Africa. Here we employ collagen peptide mass fingerprinting, known as Zooarchaeology by Mass Spectrometry (ZooMS), coupled with recently published African ZooMS reference datasets, to identify domesticates and wild bovids in Iron Age assemblages at the cave site of Panga ya Saidi in southeast Kenya. Through ZooMS we have identified all three major African livestock—sheep (Ovis aries), goat (Capra hircus) and cattle (Bos taurus)—at the site for the first time. The results provide critical evidence for the use of domesticates by resident foraging populations during the Iron Age, the period associated with the arrival of food production in coastal Kenya. ZooMS results show that livestock at Panga ya Saidi form a minor component of the assemblage compared to wild bovids, demonstrating the persistence of hunting and the secondary role of acquiring livestock in hunter-gatherer foodways during the introduction of agro-pastoralism. This study sheds new light on the establishment of food production in coastal eastern Africa, particularly the role of interactions between hunter-gatherers and neighbouring agro-pastoral groups in what was a protracted regional transition to farming.1. Introduction 2. Background 2.1. Problems with differentiating domesticates in African zooarchaeology 2.2. ZooMS in Africa 2.3. Panga ya Saidi, southeastern Kenya 2.3.1. Ecological setting 2.3.2. Archaeological background 3. Materials and methods 3.1. Sample Selection 3.2. ZooMS protocol 4. Results 5. Discussion 5.1. The impact of domestic livestock on hunting economies at PYS 5.2. Livestock trading or herding? 5.3. The role of domesticates in forager-farmer interactions 5.4. Behavioural and ecological implications of klipspringers at PYS 6. Conclusio

Do clinicians adhere to practice guidelines? A descriptive study at a referral hospital in Kenya

Background: Clinical guidelines when implemented correctly have shown to improve disease outcomes. This study describes utilization of Kenya National guidelines in managing ante partum haemorrhage (APH) in 3rd trimester.Objective: To describe adherence to clinical guideline in management of antepartum haemorrhage at Garissa Provincial General Hospital Design: Crossectional mixed methods studySetting: Garissa Provincial General HospitalSubjects: Medical records of patients managed for APH between 2002 and 2012 and Key Informant Interviews (KIIs) of Health workers.Results: 36.1% of the cases assessed were managed with strict adherence to guidelines. 90% of health care workers had high levels of awareness of the existence of guidelines and sited utilization challenges attributed to resource inadequacies.Conclusion: Clinicians are skilled on APH guidelines, but adherence levels are still low. Therefore, continuous appraisal of clinical practices, availing equipment, facilities and supplies to reinforce adherence is recommended

Distinguishing African bovids using Zooarchaeology by Mass Spectrometry (ZooMS): new peptide markers and insights into Iron Age economies in Zambia

Assessing past foodways, subsistence strategies, and environments depends on the accurate identification of animals in the archaeological record. The high rates of fragmentation and often poor preservation of animal bones at many archaeological sites across sub-Saharan Africa have rendered archaeofaunal specimens unidentifiable beyond broad categories, such as “large mammal” or “medium bovid”. Identification of archaeofaunal specimens through Zooarchaeology by Mass Spectrometry (ZooMS), or peptide mass fingerprinting of bone collagen, offers an avenue for identification of morphologically ambiguous or unidentifiable bone fragments from such assemblages. However, application of ZooMS analysis has been hindered by a lack of complete reference peptide markers for African taxa, particularly bovids. Here we present the complete set of confirmed ZooMS peptide markers for members of all African bovid tribes. We also identify two novel peptide markers that can be used to further distinguish between bovid groups. We demonstrate that nearly all African bovid subfamilies are distinguishable using ZooMS methods, and some differences exist between tribes or sub-tribes, as is the case for Bovina (cattle) vs. Bubalina (African buffalo) within the subfamily Bovinae. We use ZooMS analysis to identify specimens from extremely fragmented faunal assemblages from six Late Holocene archaeological sites in Zambia. ZooMS-based identifications reveal greater taxonomic richness than analyses based solely on morphology, and these new identifications illuminate Iron Age subsistence economies c. 2200–500 cal BP. While the Iron Age in Zambia is associated with the transition from hunting and foraging to the development of farming and herding, our results demonstrate the continued reliance on wild bovids among Iron Age communities in central and southwestern Zambia Iron Age and herding focused primarily on cattle. We also outline further potential applications of ZooMS in African archaeology.Introduction Faunal identifications and key research questions ZooMS in African archaeology Materials & methods Collagen extraction and digestion Peptide mass fingerprinting LC-MS/MS Biomarker identification and confirmation Identification of archaeological samples Results and discussion - Data quality control - Distinguishing among bovid groups Comparison with published markers ZooMS analysis and archaeofaunal identifications Herding economies and the persistence of hunting in Iron Age Zambia Conclusio

The Middle to Later Stone Age transition at Panga ya Saidi, in the tropical coastal forest of eastern Africa

The Middle to Later Stone Age transition is a critical period of human behavioral change that has been variously argued to pertain to the emergence of modern cognition, substantial population growth, and major dispersals of Homo sapiens within and beyond Africa. However, there is little consensus about when the transition occurred, the geographic patterning of its emergence, or even how it is manifested in the stone tool technology that is used to define it. Here, we examine a long sequence of lithic technological change at the cave site of Panga ya Saidi, Kenya, that spans the Middle and Later Stone Age and includes human occupations in each of the last five Marine Isotope Stages. In addition to the stone artifact technology, Panga ya Saidi preserves osseous and shell artifacts, enabling broader considerations of the covariation between different spheres of material culture. Several environmental proxies contextualize the artifactual record of human behavior at Panga ya Saidi. We compare technological change between the Middle and Later Stone Age with on-site paleoenvironmental manifestations of wider climatic fluctuations in the Late Pleistocene. The principal distinguishing feature of Middle from Later Stone Age technology at Panga ya Saidi is the preference for fine-grained stone, coupled with the creation of small flakes (miniaturization). Our review of the Middle to Later Stone Age transition elsewhere in eastern Africa and across the continent suggests that this broader distinction between the two periods is in fact widespread. We suggest that the Later Stone Age represents new short use-life and multicomponent ways of using stone tools, in which edge sharpness was prioritized over durability

A Quaternary sequence of terrestrial molluscs from East Africa: a record of diversity, stability, and abundance since Marine Isotope Stage 5 (78,000 BP)

A Quaternary sequence of subfossil terrestrial molluscs from tropical Kenya is described and discussed. It preserves a remarkably complete fauna of the Indian Ocean coastal forest from the surroundings of Panga ya Saidi cave, a site featuring repeated human occupation extending back at least 78,000 years. Mollusc diversity, composition, and abundance are very similar to extant faunas of the coastal forest. They vary relatively little over the period studied (chiefly a 50,000-year sequence from MIS 5 to the start of MIS 2) apart from a short-lived decrease in the dominance of “forest-only” species around 45,800 BP. The fauna of the most recently preserved layers (MIS 1) is likewise similar. Most of the 72 snail (and slug) species found are still extant at the coast, including some narrow-range endemics, but 8 species are now more western in their known distribution. The native African status of Kaliella barrakporensis and 2 other snail species with Asian type localities are confirmed, as is the previously disputed occurrence of native Helicoidea at the coast. Two new subfossil species were identified and are described as Maizania meteor sp. n. (Maizaniidae) and Juventigulella saidii sp. n. (Streptaxidae). No major habitat or faunistic shifts are observed, confirming previous evidence for long-term ecological continuity at the site. The data are the first of their kind from coastal East Africa and provide a new independent proxy of the environmental context to the archaeological sequence, as well as a reference point for future studies of terrestrial molluscs in the region

Ancient DNA and deep population structure in sub-Saharan African foragers

Multiple lines of genetic and archaeological evidence suggest that there were major demographic changes in the terminal Late Pleistocene epoch and early Holocene epoch of sub-Saharan Africa(1-4). Inferences about this period are challenging to make because demographic shifts in the past 5,000 years have obscured the structures of more ancient populations(3,5). Here we present genome-wide ancient DNA data for six individuals from eastern and south-central Africa spanning the past approximately 18,000 years (doubling the time depth of sub-Saharan African ancient DNA), increase the data quality for 15 previously published ancient individuals and analyse these alongside data from 13 other published ancient individuals. The ancestry of the individuals in our study area can be modelled as a geographically structured mixture of three highly divergent source populations, probably reflecting Pleistocene interactions around 80-20 thousand years ago, including deeply diverged eastern and southern African lineages, plus a previously unappreciated ubiquitous distribution of ancestry that occurs in highest proportion today in central African rainforest hunter-gatherers. Once established, this structure remained highly stable, with limited long-range gene flow. These results provide a new line of genetic evidence in support of hypotheses that have emerged from archaeological analyses but remain contested, suggesting increasing regionalization at the end of the Pleistocene epoch. DNA analysis of 6 individuals from eastern and south-central Africa spanning the past approximately 18,000 years, and of 28 previously published ancient individuals, provides genetic evidence supporting hypotheses of increasing regionalization at the end of the Pleistocene.info:eu-repo/semantics/publishedVersio

Hunter-gatherer technological organization and responses to Holocene climate change in coastal, lakeshore, and grassland ecologies of eastern Africa

The Holocene of eastern Africa saw extreme climatic fluctuations between hyper-humid and arid conditions, which manifested differently across the region's lake basins, coastal ecotones, and terrestrial biomes. Changes to resource availability, distribution, and predictability presented different constraints and opportunities to diverse hunter-gatherer communities. Major ongoing questions concern how humans reconfigured economic, social, and technological strategies in different regional settings. The role of more stable coastal environments in these processes remains especially under-studied. Here, we examine and compare relationships between environmental change and the organization of stone tool technology at the site of Panga ya Saidi Cave, eastern Kenya, in strata dating from c. 15-0.2 ka. Located near the Indian Ocean coast, this dataset provides the first insights into Holocene human-environmental relationships in a coastal forest zone of eastern Africa. Integrating the new Panga ya Saidi environmental and archaeological records with other high-resolution records from nearby terrestrial and lacustrine zones, we take a comparative approach to address how climatic fluctuations shaped trajectories of hunter-gatherer adaptations through the Holocene. We argue that lithic technologies deployed within lake basins and coastal zones reflect more stable land-use strategies with less residential mobility compared to those associated with terrestrial foraging. All regions exhibit technological reconfigurations with the arrival of pastoralism, except for the coastal forest which appear largely consistent across the study period. Results inform ongoing debates into the resilience of recent eastern African hunter-gatherers and food-producers and provide an analogical framework for examining human-environmental dynamics deeper in time.1. Introduction 2. Background 2.1. Holocene climatic change in north-eastern Africa 2.2. Lithic technological organization and inferring human-environmental interactions 3. Holocene climatic and technological change across eastern Africa 3.1. Lake Turkana Basin Fisher-foragers 3.2. Hunter-gatherers in the Central Rift valley 3.3. Lake Victoria Fisher-foragers 3.4. Central Tanzania/Lake Eyasi basin hunter-gatherers 3.5. The eastern African coast 3.6. Panga ya Saidi 4. Methods 4.1. Temporal division and sampling 4.2. Lithic analysis 5. Results 5.1. Tool technology 5.2. Core morphology 5.3. Core preparation and repair 5.4. Core reduction strategies 5.4.1. Use of quartz and limestone 5.4.2. Chert technology 6. Discussion 6.1. Terminal Pleistocene-to-Holocene mobility and lithic economy at Panga ya Saidi 6.2. Relationship between technological variation, climate change, and forager resilience in Holocene eastern Africa 6.3. Wider implications for understanding variation in stone tool technology 7. Conclusion