2,149 research outputs found

    Are damselfish sexually dimorphic in ultraviolet?

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    Evolution of the rates of mass wasting and fluvial sediment transfer from the epicentral area of the 1999, Mw 7.6 earthquake

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    The 1999 Chichi earthquake (Mw=7.6) triggered more than 20,000 landslides in the epicentral area in central west Taiwan, and subsequent typhoons have caused an even larger number of slope failures. As a result, the suspended sediment load of the epi- central Choshui River has increased dramatically. Measurements of suspended sedi- ment at a downstream gauging station indicate that the unit sediment concentration increased about six times due to the earthquake, and decreased exponentially due to flushing by subsequent typhoons. The e-folding time scale of the seismic perturbation of sediment transfer in the Choshui River is 3-5 years. Based on this estimate of the de- cay of the erosional response to the earthquake, a mass balance can be calculated for the earthquake, including co-seismic uplift and subsidence, post-seismic relaxation, and erosion. This mass balance shows that the Chi-Chi earthquake has acted to build ridge topography in the hanging wall of the fault, but in the far field, some destruc- tion of topography has occurred. However, our estimate of seismically-driven erosion may be incomplete. A detailed analysis of landsliding in the Chenyoulan tributary of the Choshui River indicates that most co-and post seismic landslide debris remains on hillslopes within the catchment. Recent typhoons have continued to cause high rates of landsliding high in the landscape, but rates of mass wasting near the stream net- work have decreased. The full geomorphic response to the Chi-Chi earthquake may be much larger, and more protracted than indicated by river gauging data

    Exploring the origin of ice-filled craters in the north polar region of Mars

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    We investigate the origins of enigmatic ice-filled craters in the north polar region of Mars. We test several explanations for their origin, namely: (1) as polar cap remnants (2) accumulation independently of the polar cap, and (3) upwelling of subsurface water, analogous to either aufice or pingo formation on Earth. Each of these hypotheses has a significant impact on our understanding of Mars’ recent geological and climatic history and the behaviour of water and water ice at high latitudes. We used several lines of evidence to assess the most likely formation mechanism. We first performed a crater survey based on THEMIS visual data and MOLA elevation data to identify any craters that had domal central lumps which were different from normal central peaks. From this survey we identified 17 craters for further study. These include Louth, Korolev, Dokka and other unnamed craters. Using data from orbiting spectrometers; OMEGA on ESA’s Mars Express and CRISM on NASA’s Mars Reconnaissance Orbiter; we verified that the composition of the exposed central domes was predominantly water ice. We found the domes fell into three groups: (1) those completely covered by dunes, (2) those partially covered by dunes and (3) those with no dunes. We investigated the morphology and the relative position of the domes using MOLA elevation data. We found that the domes are always asymmetrically placed within the craters. However, this asymmetry could not easily be linked to wind directions as revealed by dune slip-faces [2]. The domes often have a moat-like structure and in some cases do not cover the entire crater floor, e.g. Louth Crater. From image data, we identified six craters which possessed internal stratigraphy, in the form of regularly spaced layers, and of these we have inspected three in detail. We found that the layers possess both strong sinuosity and high angle unconformities. We interpret the internal stratigraphy as representing a sequence of regular cyclic accumulations, which produced the layers, followed by asymmetric ablation and subsequent resumption of accumulation, to produce the unconformities. Hence, the present-day shape of the domes indicates that they are in a phase of ablation.We attribute the colour contrasts between layers to different levels of dust, or particulate content. This could form a source for the dunes, which are often located on the summits of these domes. We find that this sequence is best explained by a model of atmospheric condensation. Our measurements of internal layer spacing and observations of layer stratigraphy argues that these deposits are not linked directly to a former, more extensive polar cap

    A lifelong study of a pack Rhodesian ridgeback dogs reveals subclinical and clinical tick-borne Anaplasma phagocytophilum infections with possible reinfection or persistence.

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    Various tick-borne infections often occur without specific clinical signs and are therefore notoriously hard to diagnose separately in veterinary practice. Longitudinal studies over multiple tick seasons performing clinical, serological and molecular investigations in parallel, may elucidate the relationship between infection and disease. In this regard, six related Rhodesian Ridgeback dogs living as a pack became subject of lifetime studies due to ongoing tick infestations and recurring clinical problems. Blood samples for diagnostic tests were obtained throughout the years 2000 to 2009

    Functional asymmetry of transmembrane segments in nicotinic acetylcholine receptors

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    Nicotinic acetylcholine receptors are heteropentameric ion channels that open upon activation to a single conducting state. The second transmembrane segments of each subunit were identified as channel-forming elements, but their respective contribution in the gating process remains unclear. Moreover, the detailed impact of variations of the membrane potential, such as occurring during an action potential, on the transmembrane domains, is unknown. Residues at the 12′ position, close to the center of each second transmembrane segment, play a key role in channel gating. We examined their functional symmetry by substituting a lysine to that position of each subunit and measuring the electrical activity of single channels. For 12′ lysines in the α, γ and δ subunits rapid transitions between an intermediate and large conductance appeared, which are interpreted as single lysine protonation events. From the kinetics of these transitions we calculated the pK a values of respective lysines and showed that they vary differently with membrane hyperpolarization. Respective mutations in β or ε subunits gave receptors with openings of either intermediate or large conductance, suggesting extreme pK a values in two open state conformations. The results demonstrate that these parts of the highly homologous transmembrane domains, as probed by the 12′ lysines, sense unequal microenvironments and are differently affected by physiologically relevant voltage changes. Moreover, observation of various gating events for mutants of α subunits suggests that the open channel pore exists in multiple conformations, which in turn supports the notion of functional asymmetry of the channe

    Cenozoic exhumation history of the Alborz Mountains, Iran: New constraints from low-temperature chronometry

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    [1] The Alborz Mountains in the north of the Turkish-Iranian Plateau mainly developed in the Cenozoic as a consequence of the closure of Neo-Tethys and continental collision between Arabia and Eurasia. Cenozoic growth of the fold-and-thrust belt exploited an older Palaezoic-Mesozoic crustal fabric but the extent to which this governed the overall form of the mountain belt is unclear. To determine when and how the Alborz mountain belt has grown, apatite fission track (AFT) and (U-Th)/He (AHe) thermochronometry were performed on 46 bedrock samples collected along 8 transects across the range. AFT central ages range from 157 ± 24 Ma to 10 ± 1 Ma with most ages falling between 40 Ma and 10 Ma. AHe ages range from 17 ± 2 Ma to 6 ± 1 Ma. The data revealed enhanced exhumation ca. 35 ± 5 Ma, minor exhumation between ca. 30–20 Ma and an increase in exhumation thereafter. This pattern matches the tripartite Mid-Cenozoic stratigraphy of Central Iran, where Upper Oligocene-Lower Miocene carbonates are sandwiched between terrestrial clastic formations. The most intriguing thermochronometric signal found is a major acceleration of exhumation that initiated in the late Miocene to early Pliocene, recorded by the AHe data. There does not appear to be a direct tectonic cause for Pliocene intensified erosion, as convergence rates between Arabia and Eurasia have remained relatively constant. Enhanced exhumation at this time may reflect a climatically induced intensification of erosion during Caspian isolation and base level fall, or a regional tectonic re-organization of the Arabia-Eurasia collision, or both

    Temporal response of mountain drainage basins in Taiwan to earthquake and typhoon perturbation.

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    In tectonically-active mountain belts, earthquake-triggered landslides deliver large amounts of sediment to rivers. In previous work, we have quantified the geomorphic impact of the 1999 Mw 7.6 Chi-Chi earthquake in Taiwan, which triggered >20,000 landslides and elevated suspended sediment loads in rivers by up to a factor of four. At the time, many coseismic landslides remained confined to hillslopes and, on the basis of four years of hydrometric data, we predicted that downslope transport of sediment would continue to occur during later storms. During the seven years since the Chi- Chi earthquake, several major typhoons storms have hit Taiwan (e.g., Typhoons Bilis, Toraji, Nari, Mindulle, Aere) and the Water Resources Agency of Taiwan has contin- ued to monitor water discharge and suspended sediment concentration. Here we use these new data to refine the spatial and temporal pattern of the decaying geomorphic response to the Chi-Chi earthquake in the face of several large typhoons. Our results indicate that the broad pattern of exponential decay in sediment concentration for a given river discharge (prevalent in winter seasons without typhoons) is punctuated by markedly elevated periods associated with typhoon storms. However, our analyses show that the change in unit sediment concentration (i.e., suspended sediment concen- tration for a unit water discharge) associated with each storm depends more strongly on the length of time elapsed since the earthquake than it does on the magnitude of the storm itself

    Gebiedsmanagement in ontwikkeling

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