A Middle Jurassic (Bajocian) ophthalmosaurid (Reptilia, Ichthyosauria) from the Tuxedni Formation, Alaska and the early diversification of the clade

Ophthalmosauridae is a clade of derived thunniform ichthyosaurs that are best known from Callovian (late Middle Jurassic) to Cenomanian-aged (Late Cretaceous) deposits in both the Northern and Southern Hemispheres. Ophthalmosaurids arose prior to the Early-Middle Jurassic boundary, however, very little is known about their diversity and distribution in the earliest phase of their evolutionary history during the Aalenian-Bathonian (Middle Jurassic) interval. Here we describe new diagnostic ophthalmosaurid material from the Early Bajocian (Middle Jurassic) of Alaska. The specimen, UAMES 3411, is a partial disarticulated skull that was discovered in the Middle Jurassic Tuxedni Formation, which was deposited in shallow marine settings outboard of the then-accreting Wrangellia composite terrane. The new material is significant in that it is the first Jurassic ichthyosaur described from Alaska, one of the oldest ophthalmosaurids known and the only Middle Jurassic ophthalmosaurid described from the Northern Hemisphere. The new material adds to a rapidly growing data set on ophthalmosaurid diversity and suggests that the clade was geographically widespread by the Early Bajocian, very early in its evolutionary histor

The Slottsmøya marine reptile Lagerstätte: depositional environments, taphonomy and diagenesis

The Late Jurassic Slottsmøya Member Lagerstätte on Spitsbergen offers a unique opportunity to study the relationships between vertebrate fossil preservation, invertebrate occurrences and depositional environment. In this study, 21 plesiosaurian and 17 ichthyosaur specimens are described with respect to articulation, landing mode, preservation, and possible predation and scavenging. The stratigraphic distribution of marine reptiles in the Slottsmøya Member is analysed, and a correlation between high total organic content, low oxygen levels, few benthic invertebrates and optimal reptile preservation is observed. A new model for 3D preservation of vertebrates in highly compacted organic shales is explained

First records of a plesiosaurian (Reptilia: Sauropterygia) and an ichthyosaur (Reptilia: Ichthyosauria) from Yukon, Canada

An isolated centrum collected ex situ from marine shales of the Lower Cretaceous (Albian) Arctic Red Formation along the Road River represents the first documented occurrence of a plesiosaurian from Yukon. This centrum represents the northernmost occurrence of plesiosaurians in the Western Interior Sea of North America prior to the establishment of the first continuous seaway (Western Interior Seaway) connecting the Boreal and Tethyan seas. Additionally, this centrum is potentially the secondoldest elasmosaurid specimen known from North America. A second centrum, collected along the Beaver River, is likely derived from the Lower Cretaceous (Lower Albian) Garbutt Formation exposed farther upstream. It represents the first report of an ichthyosaur from Yukon. Additionally, six associated ribs collected from the Arctic Re

The Alaska Arctic Vegetation Archive (AVA-AK)

The Alaska Arctic Vegetation Archive (AVA-AK, GIVD-ID: NA-US-014) is a free, publically available database archive of vegetation-plot data from the Arctic tundra region of northern Alaska. The archive currently contains 24 datasets with 3,026 non-overlapping plots. Of these, 74% have geolocation data with 25-m or better precision. Species cover data and header data are stored in a Turboveg database. A standardized Pan Arctic Species List provides a consistent nomenclature for vascular plants, bryophytes, and lichens in the archive. A web-based online Alaska Arctic Geoecological Atlas (AGA-AK) allows viewing and downloading the species data in a variety of formats, and provides access to a wide variety of ancillary data. We conducted a preliminary cluster analysis of the first 16 datasets (1,613 plots) to examine how the spectrum of derived clusters is related to the suite of datasets, habitat types, and environmental gradients. Here, we present the contents of the archive, assess its strengths and weaknesses, and provide three supplementary files that include the data dictionary, a list of habitat types, an overview of the datasets, and details of the cluster analysis

High Diversity, Low Disparity and Small Body Size in Plesiosaurs (Reptilia, Sauropterygia) from the Triassic–Jurassic Boundary

Invasion of the open ocean by tetrapods represents a major evolutionary transition that occurred independently in cetaceans, mosasauroids, chelonioids (sea turtles), ichthyosaurs and plesiosaurs. Plesiosaurian reptiles invaded pelagic ocean environments immediately following the Late Triassic extinctions. This diversification is recorded by three intensively-sampled European fossil faunas, spanning 20 million years (Ma). These provide an unparalleled opportunity to document changes in key macroevolutionary parameters associated with secondary adaptation to pelagic life in tetrapods. A comprehensive assessment focuses on the oldest fauna, from the Blue Lias Formation of Street, and nearby localities, in Somerset, UK (Earliest Jurassic: 200 Ma), identifying three new species representing two small-bodied rhomaleosaurids (Stratesaurus taylori gen et sp. nov.; Avalonnectes arturi gen. et sp. nov) and the most basal plesiosauroid, Eoplesiosaurus antiquior gen. et sp. nov. The initial radiation of plesiosaurs was characterised by high, but short-lived, diversity of an archaic clade, Rhomaleosauridae. Representatives of this initial radiation were replaced by derived, neoplesiosaurian plesiosaurs at small-medium body sizes during a more gradual accumulation of morphological disparity. This gradualistic modality suggests that adaptive radiations within tetrapod subclades are not always characterised by the initially high levels of disparity observed in the Paleozoic origins of major metazoan body plans, or in the origin of tetrapods. High rhomaleosaurid diversity immediately following the Triassic-Jurassic boundary supports the gradual model of Late Triassic extinctions, mostly predating the boundary itself. Increase in both maximum and minimum body length early in plesiosaurian history suggests a driven evolutionary trend. However, Maximum-likelihood models suggest only passive expansion into higher body size categories

A Middle Jurassic (Bajocian) ophthalmosaurid (Reptilia, Ichthyosauria) from the Tuxedni Formation, Alaska and the early diversification of the clade

Ophthalmosauridae is a clade of derived thunniform ichthyosaurs that are best known from Callovian (late Middle Jurassic) to Cenomanian-aged (Late Cretaceous) deposits in both the Northern and Southern Hemispheres. Ophthalmosaurids arose prior to the Early-Middle Jurassic boundary, however, very little is known about their diversity and distribution in the earliest phase of their evolutionary history during the Aalenian-Bathonian (Middle Jurassic) interval. Here we describe new diagnostic ophthalmosaurid material from the Early Bajocian (Middle Jurassic) of Alaska. The specimen, UAMES 3411, is a partial disarticulated skull that was discovered in the Middle Jurassic Tuxedni Formation, which was deposited in shallow marine settings outboard of the then-accreting Wrangellia composite terrane. The new material is significant in that it is the first Jurassic ichthyosaur described from Alaska, one of the oldest ophthalmosaurids known and the only Middle Jurassic ophthalmosaurid described from the Northern Hemisphere. The new material adds to a rapidly growing data set on ophthalmosaurid diversity and suggests that the clade was geographically widespread by the Early Bajocian, very early in its evolutionary histor

Two new ophthalmosaurids (Reptilia: Ichthyosauria) from the Agardhfjellet Formation (Upper Jurassic: Volgian/Tithonian), Svalbard, Norway

Ichthyosaur diversity near the Jurassic-Cretaceous boundary, particularly from high paleolatitudes, is poorly known. Two recently collected specimens of medium- to large-bodied ichthyosaurs from the Slottsmøya Member of the Agardhfjellet, Svalbard, Norway represent two new taxa of ophthalmosaurids. The holotype specimen of Cryopterygius kristiansenae gen et sp. nov., PMO 214.578, is a nearly complete and largely articulated skeleton. The specimen consists of a nearly complete skull, the entire presacral and preflexural vertebral series, numerous dorsal ribs and gastralia, an articulated pectoral girdle and nearly complete forelimb, and an articulated left pelvic girdle and hindlimb. The new taxon is diagnosed on a unique suite of features, including a robust and moderately elongate rostrum, a reduced supranarial process, an elongate maxilla that bears a high number of teeth, the absence of a lacrimal-external naris contact, and an anteroposteriorly broad postorbital bar possessing an unidentified element (supratemporal?) that lies posterior to the quadratojugal. Cryopterygius has 52 presacral vertebrae, a distinctive forelimb, including a humerus that bears only two facets at its distal end, and an articulated left pelvic girdle and hindlimb, which facilitates the unequivocal orientation of the ophthalmosaurid femur. The holotype specimen of Palvennia hoybergeti, SVB 1451, includes a nearly complete skull and fragmented postcranial remains. It is diagnosed on its relatively short rostrum, greatly enlarged orbit, narrow postorbital bar, very large pineal foramen, basioccipital with broad extracondylar area laterally, and a gracile stapedial shaft. The Slottsmøya Lagerstätte is established as one of the most productive horizons for Upper Jurassic ichthyosaurs and considerably expands our knowledge of ophthalmosaurid diversity and distribution in the latest Jurassic

Redescription and taxonomic clarification of 'Tricleidus' svalbardensis based on new material from the Agardhfjellet Formation (Middle Volgian)

A partial postcranial plesiosaurian skeleton uncovered on Svalbard in the winter of 1930-31 was described and named Tricleidus svalbardensis Persson, 1962. However, the precise geographic location and stratigraphic unit in which the skeleton was found were not published. Recent fieldwork on Svalbard has uncovered two more plesiosaur skeletons that are demonstrated herein to be conspecific with the 1931 taxon. These new specimens, along with recently discovered historic documents produced by the excavation members of the 1930-31 expedition, reveal that the holotype specimen, PMO A 27745, was recovered from the Upper Jurassic Slottsmøya Member of the Agardhfjellet Formation, which is Middle Volgian in age. Collectively, the Svalbard material is neither morphologically nor stratigraphically consistent with the Callovian taxon Tricleidus Andrews, 1909 and is here referred to Colymbosaurus Seeley, 1874 from the Kimmeridge Clay Formation (Kimmeridgian to Tithonian) of the UK