La longue séquence de Marchésieux: reconstitution de paléoenvironnements marins durant le premier cycle glaciaire de l'hémisphère nord

International audienceA l'échelle des cinq derniers millions d'années, les enregistrements paléoclimatiques restitués par les sédiments océaniques illustrent le contrôle des paramètres orbitaux (précession, obliquité et excentricité) sur le climat global. La tendance au refroidissement qui accompagne la fin du Néogène est ainsi marquée, vers -2,4 Ma, par le développement rapide des calottes de glace de l'hémisphère nord (Shackleton et al., 1984). Alors que l'histoire du climat global restituée par les enregistrements océaniques est sans cesse précisée, les données concernant l'évolution des environnements continentaux et côtiers au cours de ces changements restent essentiellement fragmentaires. Le forage effectué à Marchésieux (Manche; Normandie) a permis de réaliser une étude pluridisciplinaire de la signature de ce premier cycle glaciaire de l'hémisphère nord (Prétiglien). Les premiers résultats stratigraphiques, paléoenvironnementaux (eustatisme, température, paléobathymétrie) sont présentés

Occurrence of whale barnacles in Nerja Cave (Málaga, southern Spain): Indirect evidence of whale consumption by humans in the Upper Magdalenian

A total of 167 plates of two whale barnacle species (Tubicinella majorLamarck, 1802 and Cetopirus complanatusMörch, 1853) have been found in the Upper Magdalenian layers of Nerja Cave, Mina Chamber (Maro, Málaga, southern Spain). This is the first occurrence of these species in a prehistoric site. Both species are specific to the southern right whale Eubalena australis, today endemic in the Southern Hemisphere. Because of Antarctic sea-ice expansion during the Last Glacial Period, these whales could have migrated to the Northern Hemisphere, and reached southern Spain. Whale barnacles indicate that maritime-oriented forager human groups found stranded whales on the coast and, because of the size and weight of the large bones, transported only certain pieces (skin, blubber and meat) to the caves where they were consumed

Acasta vesiculosa Carriol, 2008, sp. nov.

Acasta vesiculosa sp. nov. Fig. 3 (3–6) Etymology: Morphologic, an allusion to the calcareous vesicles of the sheath. Occurrence: Ferrière-Larçon and Pauvrelay (Paulmy commune), Indre-et-Loire, France, faluns of Middle Miocene age. Diagnosis: Shell with carinal and lateral sheath vesicular; carinal sheath adhering to the wall in the middle of its basal edge. Material: Type material. Holotype: MNHN A 24752 carina. Paratypes: MNHN A 24753 carino-lateral; MNHN A 24754 lateral; MNHN A 24755 carino-lateral. Additional material. Fragments of 8 carinae, 11 carino-laterals and 6 laterals. Measurements: Paratype MNHN A 24753 height of the middle of the plate 5.9 mm. Paratype MNHN A 24754 height of the middle of this broken plate 3.1 mm. Description: Shell white; parietes externally with knobs, basis of much longer, hollow, upward curving thorns sloping downward, internally slightly ribbed; radii and alae not very wide, upper edge very oblique; sutural edge of radii smooth; sheath broad, transversally striated, with basal edge free from wall in laterals and carino-laterals, not in carinae where central part of basal edge goes down, adhering to wall; sheath of carinae and laterals composed of irregular, thin-walled calcareous vesicles. Whole specimen unavailable, general shape and relative proportional width of parietes of carino-laterals unknown; Opercular plates and rostrum unknown. Affinities: Extinct Acasta may be distinguished from A. vesiculosa sp. nov. in the lack of a vesicular sheath. They also differ as follows: A. sarda is higher with external surface of the parietes with horizontal thorns; A. formae is higher, with upper edge of the radii parallel to the basis; A. undulata, a large species, has compartments rather externally smooth; A. muricata lacks striae on the internal surface of the parietes; A. fischeri has the upper edges of the radii and alae parallel to the basal edge of the plates; A. spongites has edges of the radii slightly crenated, A. cyathus has very wide radii. In light of the absence of the holotype and an incomplete description, the taxonomic placement of A. hebertiana cannot be confirmed.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on pages 44-45, DOI: 10.5281/zenodo.18029

Balanus nodulus Carriol, 2008, sp. nov.

Balanus nodulus sp. nov. Fig. 2 (11) Diagnosis: Paries exteriorly with protuberances. Etymology: Morphological, alluding to protuberances (nod in Latin) of the paries; Occurrence: Pauvrelay (Paulmy commune), Indre-et-Loire, France; faluns of Middle Miocene age. Material: Holotype: MNHN A 25755 right lateral plate. Measurements: Holotype, MNHN A 25755: height of the middle of the plate 1.8 mm. Description: Colour ivory. Paries exteriorly with small protuberances on growth ridges. Parietal tubes in single row. Longitudinal septa of parietal inner lamina with lateral denticles. Sheath with lateral margin not extended. Radius solid with very oblique upper edge, sutural edge with smooth teeth, inner face without longitudinal abutment. Ala with secondary alar increment, summit oblique, sutural edge septed. Affinities: Among the Balanoidea, barnacles with calcareous projections were restricted to the Acastinae Kolbasov, 1993 and the Megabalaninae Newman, 1979. Balanus nodulus sp. nov. is the first Balaninae, found with this pattern.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on page 45, DOI: 10.5281/zenodo.18029

Balanus spongicola subsp. spongicola Brown 1844

Balanus spongicola spongicola Brown, 1844 Fig. 2 (12–13) Balanus spongicola, Brown, 1844: 121, pl. 53 figs 14–16 Other references: B. spongicola, Carriol, 1982: 14, 1992: 146, 1996: 57; De Alessandri, 1906: 277, 1907: 195, 1908: 218; Lecointre, 1910: 137; Menesini, 1984: 298; Menesini & Casella, 1988: 250; Newman & Ross, 1976: 66 (long list of references); Relini, 1980: 73; Stubbings: 1963: 22. Occurrence: Ferrière-Larçon, Pauvrelay (Paulmy commune), Les Bournais (Saint-Laurent-de-Lin commune), Indre-et-Loire, France; faluns of Middle Miocene age. Material: MNHN A 25756: a right scutum; MNHN A 25757: a right tergum. In addition to the figured opercular plates, several unregistered specimens have been observed: complete shells and opercular plates from Saint Laurent-de-Lin commune and fragments of opercular plates from Ferrière-Larçon and Paulmy. Measurements: MNHN A 25756: occludent margin 5.0 mm; MNHN A 25757: scutal margin 2.8 mm. Description: The material studied herein conforms to the description of Stubbings (1963). Shape conical, smooth-walled; orifice toothed owing to oblique margins to radii. Parietal tubes septed. Scutum with prominent growth ridges cut by grooves radiating from apex; internally, adductor pit moderately deep, adductor ridge rather small; depressor muscle pit strongly developed, elongated, with characteristic deep furrow prolonging line of pit towards apex; spur fasciole, broad, truncate, separated by about half its width from basiscutal angle; basal margin on carinal side curves smoothly into spur.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on pages 45-46, DOI: 10.5281/zenodo.18029

Crustacés Cirripèdes actuels du Muséum D'Histoire naturelle de Lyon

Catalogue of the extant Cirripedia Crustacea (Pedunculata : Lepadoidea, Scalpelloidea ; Sessilia : Verrucomorpha, Balanomorpha) giving identity and also qualitative and quantitative state of indexed material (115 batches).Inventaire des Crustacés Cirripèdes actuels (Pedunculata : Lepadoidea, Scalpelloidea ; Sessilia : Verrucomorpha, Balanomorpha) donnant l'identité ainsi que l'état qualitatif et quantitatif du matériel répertorié (115 lots).Carriol René-Pierre. Crustacés Cirripèdes actuels du Muséum D'Histoire naturelle de Lyon. In: Cahiers scientifiques du Muséum d'histoire naturelle de Lyon - Centre de conservation et d'étude des collections, tome 2, 2001. pp. 5-17

Actinobalanus collinsi Zullo & Perreault 1989

Actinobalanus collinsi Zullo & Perreault, 1989 Fig. 3 (1–2) Actinobalanus collinsi Zullo & Perreault, 1989: 6 –8; pl. 1, figs 5–7; pl. 2; pl. 3, figs 1–4. Type locality: Borgerhout (Berchem Formation, Edegem sands, Metro works spoil heaps) Antwerp, Belgium. Occurrence: Ferrière-Larçon and Pauvrelay (Paulmy commune), Indre-et-Loire, France, faluns of Middle Miocene age. Material: MNHN A 24750: left scutum. MNHN A 24751: right tergum. In addition to the figured specimens, numerous unregistered opercular plates have been observed. Measurements: MNHN A 24750: occludent margin 3.2 mm, tergal margin 1.8 mm, basal margin 2.2 mm. MNHN A 24751: scutal margin 2.5 mm, basal margin 2.5 mm. Diagnosis (after Zullo & Perreault, 1989): Parietes usually coarsely costate; radii narrow, disparietal; scutum thick, without external radial ornament; scutal adductor ridge short, prominent, centrally located; scutal articular ridge extending well beyond tergal margin; tergum thick, narrow, with straight scutal margin produced into broad shelf; tergal spur short, acuminate, occupying less than one-third width of basal margin; juncture of tergal spur with carinal side of basal margin arcuate. Remarks: The capitular plates of A. collinsi cannot be distinguished from those of A. bisulcatus. However, the opercular plates conform to the diagnosis. Stratigraphic and geographic range: Lower Miocene, Belgium (known only from the type locality); Middle Miocene, France (faluns of Touraine). The record of A. collinsi from the Middle Miocene of France extends the stratigraphic range and the geographic distribution of this species, which was previously known only in the Lower Miocene of Belgium.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on pages 41-42, DOI: 10.5281/zenodo.18029

Paractinobalanus Carriol, 2008, gen. nov.

Genus Paractinobalanus gen. nov. Diagnosis: Shell small with 6 solid compartmental plates; basis porous; radii transparietal, sutural edges with simple transverse septa. Type species: Paractinobalanus moronii sp. nov. Etymology: from para (Greek) = near + Actinobalanus Discussion: Paractinobalanus differs from all other archaeobalanid genera except Actinobalanus Moroni, 1967 in having a tubiferous basis. It can be distinguished from Actinobalanus by its possession of transparietal radii.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on page 42, DOI: 10.5281/zenodo.18029

Actinobalanus bisulcatus

Actinobalanus bisulcatus (Darwin, 1854 a) Fig. 3 (7) Balanus bisulcatus Darwin, 1854 a: 293; pl. 8, figs 6 a–c Synonymy: Balanus bisulcatus var. plicatus Darwin, 1854 a: 293; 1854 b: 26. Balanus (Armatobalanus) bisulcatus plicatus (Darwin, 1854 a): Davadie, 1963: 74. Actinobalanus bisulcatus plicatus (Darwin, 1854 a): Newman & Ross, 1976: 49. Balanus stellaris (Brocchi, 1814): Prieur & Philippe, 1984: 65. Other references: Balanus bisulcatus Darwin, 1854 a: De Alessandri, 1906 (in part); 1922: 221. Balanus (Armatobalanus) bisulcatus Darwin, 1854 a: Davadie, 1963: 74. Balanus (Actinobalanus) bisulcatus Darwin, 1854 a: Moroni, 1967: 925. Actinobalanus bisulcatus (Darwin, 1854 a): Newman & Ross, 1976: 49; Zullo & Perreault, 1989: 3; Carriol, 1992: 143. Type locality: Corraline Crag, Ramsholt, Gedgrave, Sutton, England. Occurrence: Ferrière-Larçon and Pauvrelay (Paulmy commune), Indre-et-Loire, France, faluns of Middle Miocene age. Material: MNHN A 24749: left scutum. In addition to the figured scutum, two unregistered scuta, from Paulmy, have been observed. Measurements: MNHN A 24749: occludent margin 2.6 mm, tergal margin 1.5 mm. Other scutum: occludent margin 2.9 mm, tergal margin 1.5 mm. Diagnosis (after Zullo & Perreault, 1989): Parietes smooth or markedly costate; radii narrow or broad, disparietal, scutum with two to four external longitudinal furrows; scutal adductor ridge prominent, length greater than one-half length of occludent margin; scutal articular ridge not extending beyond tergal margin; tergal spur very short, less than one-tenth length of plate, broad, occupying one-half width of basal margin, subtruncate basally, nearly confluent with basiscutal angle; juncture of spur and carinal side of basal margin arcuate. Remarks: The capitular plates of A. bisulcatus cannot be distinguished from those of A. collinsi. However, the scuta, characteristic of the species with their external longitudinal furrows (two and three furrows for the specimens from Paulmy and three for the one from Ferrière-Larçon), conform to the diagnosis. Stratigraphic and geographic range: Burdigalian (France: Cestas, Aquitain basin; Faucon-Mollans- Malaucène basin), Middle Miocene (France, faluns of Touraine), Pliocene (England, Coralline Crag, Sutton; France, Rauville and Gourbesville, Cotentin; Belgium, Bolderberg, near Hasselt, and Karrendijke formation). This is the first record of Balanus bisulcatus from the Middle Miocene.Published as part of Carriol, René-Pierre, 2008, New genus and new species of Cirripedia (Chthamalidae, Tetraclitidae, Archaeobalanidae and Balanidae) from the Middle Miocene of the faluns of Touraine (France), pp. 31-48 in Zootaxa 1675 on pages 40-41, DOI: 10.5281/zenodo.18029