Testing Speculative Behavior in Farmland Demand

Substantial increases in farmland demand in sub-urbanization have had profound effects on agriculture and produced a surge in farmland values. With escalating land values, farmland can take on the characteristics of a speculative asset and farmland owners may be more responsive to the investment value of farmland than the productive value. Speculation has been shown to have a significant impact on the agricultural production decisions of farms, and may encourage farmers to curtail capital investments and prematurely idle productive farmland. This paper investigates the effects of farmland value appreciation on agriculture and isolates the speculative component of land use demand, using New Jersey as a case study. A conceptual model of land use and investment behavior is developed and estimated. Two empirical models are used in the analysis; one that accounts for speculation and one that does not. The former is found to be superior. An inverse relationship is estimated between the rate of appreciation and the demand for farmland, suggesting a direct relationship between appreciation and land supplied to development. The relationship, however, is found to be positive at rates of farmland value appreciation in excess of the risk free rate of return. This suggests an identifiable speculative demand component whereby farmland owners retain farmland at high rates of appreciation. Results also support the conjecture that when the rate of appreciation is lower than the risk free rate, the speculative behavior of farmland owners is to keep less land in agriculture.Land Economics/Use,

Stability and response of polygenic traits to stabilizing selection and mutation

When polygenic traits are under stabilizing selection, many different combinations of alleles allow close adaptation to the optimum. If alleles have equal effects, all combinations that result in the same deviation from the optimum are equivalent. Furthermore, the genetic variance that is maintained by mutation-selection balance is $2 \mu/S$ per locus, where $\mu$ is the mutation rate and $S$ the strength of stabilizing selection. In reality, alleles vary in their effects, making the fitness landscape asymmetric, and complicating analysis of the equilibria. We show that that the resulting genetic variance depends on the fraction of alleles near fixation, which contribute by $2 \mu/S$, and on the total mutational effects of alleles that are at intermediate frequency. The interplay between stabilizing selection and mutation leads to a sharp transition: alleles with effects smaller than a threshold value of $2\sqrt{\mu / S}$ remain polymorphic, whereas those with larger effects are fixed. The genetic load in equilibrium is less than for traits of equal effects, and the fitness equilibria are more similar. We find that if the optimum is displaced, alleles with effects close to the threshold value sweep first, and their rate of increase is bounded by $\sqrt{\mu S}$. Long term response leads in general to well-adapted traits, unlike the case of equal effects that often end up at a sub-optimal fitness peak. However, the particular peaks to which the populations converge are extremely sensitive to the initial states, and to the speed of the shift of the optimum trait value.Comment: Accepted in Genetic

Rock Slope Design using Q-slope and Geophysical Survey Data

The Q-slope method for rock slope engineering provides an empirical means of assessing the stability of excavated rock slopes in the field. It enables rock engineers and engineering geologists to make potential adjustments to slope angles as rock mass conditions become apparent during the construction of reinforcement-free road or railway cuttings and in open cast mines. Q-slope was developed by supplementing the Q-system which has been extensively used for characterizing rock exposures, drill core and underground mines and tunnels under construction for over 40 years. The Q׳ parameters (RQD, Jn ,Jr and Ja) have remained unchanged in Q-slope, although a new method for applying Jr /Ja ratios to both sides of a potential wedge is used, with relative orientation weightings for each side. The term Jw has been replaced with the more comprehensive term Jwice , which takes into account long-term exposure to various climatic and environmental conditions such as intense erosive rainfall and ice-wedging effects. SRF categories have been developed for slope surface conditions, stress-strength ratios and major discontinuities such as faults, weakness zones or joint swarms. Through case studies across Europe, Australia, Asia, and Central America, a simple relationship between Q-slope and long-term stable slope angles was established. The Q-slope method is designed such that it suggests stable, maintenance-free, bench face slope angles of, for instance, 40–45°, 60–65° and 80–85° with respective Q-slope values of approximately 0.1, 1.0 and 10. Q-slope has also been found to be compatible with P-wave velocity and acoustic and optical televiewer data obtained from borehole and surface-based geophysical surveys to determine appropriate rock slope angles

Were rivers flowing across the Sahara during the last interglacial? Implications for human migration through Africa.

Human migration north through Africa is contentious. This paper uses a novel palaeohydrological and hydraulic modelling approach to test the hypothesis that under wetter climates c.100,000 years ago major river systems ran north across the Sahara to the Mediterranean, creating viable migration routes. We confirm that three of these now buried palaeo river systems could have been active at the key time of human migration across the Sahara. Unexpectedly, it is the most western of these three rivers, the Irharhar river, that represents the most likely route for human migration. The Irharhar river flows directly south to north, uniquely linking the mountain areas experiencing monsoon climates at these times to temperate Mediterranean environments where food and resources would have been abundant. The findings have major implications for our understanding of how humans migrated north through Africa, for the first time providing a quantitative perspective on the probabilities that these routes were viable for human habitation at these times

Similarity Selection and the Evolution of Sex: Revisiting the Red Queen

For over 25 years, many evolutionary ecologists have believed that sexual reproduction occurs because it allows hosts to change genotypes each generation and thereby evade their coevolving parasites. However, recent influential theoretical analyses suggest that, though parasites can select for sex under some conditions, they often select against it. These models assume that encounters between hosts and parasites are completely random. Because of this assumption, the fitness of a host depends only on its own genotype (“genotypic selection”). If a host is even slightly more likely to encounter a parasite transmitted by its mother than expected by random chance, then the fitness of a host also depends on its genetic similarity to its mother (“similarity selection”). A population genetic model is presented here that includes both genotypic and similarity selection, allowing them to be directly compared in the same framework. It is shown that similarity selection is a much more potent force with respect to the evolution of sex than is genotypic selection. Consequently, similarity selection can drive the evolution of sex even if it is much weaker than genotypic selection with respect to fitness. Examination of explicit coevolutionary models reveals that even a small degree of mother–offspring parasite transmission can cause parasites to favor sex rather than oppose it. In contrast to previous predictions, the model shows that weakly virulent parasites are more likely to favor sex than are highly virulent ones. Parasites have figured prominently in discussions of the evolution of sex, but recent models suggest that parasites often select against sex rather than for it. With the inclusion of small and realistic exposure biases, parasites are much more likely to favor sex. Though parasites alone may not provide a complete explanation for sex, the results presented here expand the potential for parasites to contribute to the maintenance of sex rather than act against it

Chromosomal speciation and molecular divergence - Accelerated evolution in rearranged chromosomes

Humans and their closest evolutionary relatives, the chimpanzees, differ in ~1.24% of their genomic DNA sequences. The fraction of these changes accumulated during the speciation processes that have separated the two lineages may be of special relevance in understanding the basis of their differences. We analyzed human and chimpanzee sequence data to search for the patterns of divergence and polymorphism predicted by a theoretical model of speciation. According to the model, positively selected changes should accumulate in chromosomes that present fixed structural differences, such as inversions, between the two species. Protein evolution was more than 2.2 times faster in chromosomes that had undergone structural rearrangements compared with colinear chromosomes. Also, nucleotide variability is slightly lower in rearranged chromosomes. These patterns of divergence and polymorphism may be, at least in part, the molecular footprint of speciation events in the human and chimpanzee lineages

Mapping Mendelian traits in asexual progeny using changes in marker allele frequency

Linkage between markers and genes that affect a phenotype of interest may be determined by examining differences in marker allele frequency in the extreme progeny of a cross between two inbred lines. This strategy is usually employed when pooling is used to reduce genotyping costs. When the cross progeny are asexual, the extreme progeny may be selected by multiple generations of asexual reproduction and selection. We analyse this method of measuring phenotype in asexual progeny and examine the changes in marker allele frequency due to selection over many generations. Stochasticity in marker frequency in the selected population arises due to the finite initial population size. We derive the distribution of marker frequency as a result of selection at a single major locus, and show that in order to avoid spurious changes in marker allele frequency in the selected population, the initial population size should be in the low to mid hundreds

Looking beyond tasks to develop flexible leadership.

There is a determination within health and social care to make leaders and leadership better and more able to face the challenges within an ever-changing social, political and organisational landscape. However, this presents difficulties, especially as workplace culture and climate may make people unlikely to take different leadership approaches. This paper reports on the findings of an evaluation of a leadership development programme delivered to senior leaders in health and social care. It uses qualitative data to demonstrate that, even in a safe, risk-free environment, health and social care leaders approach a hypothetical scenario in a defensive, hierarchical and expert-led manner; only after they were given ‘expert’ permission to change leadership style, did they act collectively and seek to adopt a transformational style of leadership

Maladaptation and the paradox of robustness in evolution

Background. Organisms use a variety of mechanisms to protect themselves against perturbations. For example, repair mechanisms fix damage, feedback loops keep homeostatic systems at their setpoints, and biochemical filters distinguish signal from noise. Such buffering mechanisms are often discussed in terms of robustness, which may be measured by reduced sensitivity of performance to perturbations. Methodology/Principal Findings. I use a mathematical model to analyze the evolutionary dynamics of robustness in order to understand aspects of organismal design by natural selection. I focus on two characters: one character performs an adaptive task; the other character buffers the performance of the first character against perturbations. Increased perturbations favor enhanced buffering and robustness, which in turn decreases sensitivity and reduces the intensity of natural selection on the adaptive character. Reduced selective pressure on the adaptive character often leads to a less costly, lower performance trait. Conclusions/Significance. The paradox of robustness arises from evolutionary dynamics: enhanced robustness causes an evolutionary reduction in the adaptive performance of the target character, leading to a degree of maladaptation compared to what could be achieved by natural selection in the absence of robustness mechanisms. Over evolutionary time, buffering traits may become layered on top of each other, while the underlying adaptive traits become replaced by cheaper, lower performance components. The paradox of robustness has widespread implications for understanding organismal design

The statistical mechanics of a polygenic characterunder stabilizing selection, mutation and drift

By exploiting an analogy between population genetics and statistical mechanics, we study the evolution of a polygenic trait under stabilizing selection, mutation, and genetic drift. This requires us to track only four macroscopic variables, instead of the distribution of all the allele frequencies that influence the trait. These macroscopic variables are the expectations of: the trait mean and its square, the genetic variance, and of a measure of heterozygosity, and are derived from a generating function that is in turn derived by maximizing an entropy measure. These four macroscopics are enough to accurately describe the dynamics of the trait mean and of its genetic variance (and in principle of any other quantity). Unlike previous approaches that were based on an infinite series of moments or cumulants, which had to be truncated arbitrarily, our calculations provide a well-defined approximation procedure. We apply the framework to abrupt and gradual changes in the optimum, as well as to changes in the strength of stabilizing selection. Our approximations are surprisingly accurate, even for systems with as few as 5 loci. We find that when the effects of drift are included, the expected genetic variance is hardly altered by directional selection, even though it fluctuates in any particular instance. We also find hysteresis, showing that even after averaging over the microscopic variables, the macroscopic trajectories retain a memory of the underlying genetic states.Comment: 35 pages, 8 figure