Stability Analysis of a Ratio-Dependent Predator-Prey Model Incorporating a Prey Refuge

A ratio-dependent predator-prey model incorporating a prey refuge with disease in the prey population is formulated and analyzed. The effects of time delay due to the gestation of the predator and stage structure for the predator on the dynamics of the system are concerned. By analyzing the corresponding characteristic equations, the local stability of a predator-extinction equilibrium and a coexistence equilibrium of the system is discussed, respectively. Further, it is proved that the system undergoes a Hopf bifurcation at the coexistence equilibrium, when τ=τ0. By comparison arguments, sufficient conditions are obtained for the global stability of the predator-extinction equilibrium. By using an iteration technique, sufficient conditions are derived for the global attractivity of the coexistence equilibrium of the proposed system

The Dynamic Complexity of a Holling Type-IV Predator-Prey System with Stage Structure and Double Delays

We invest a predator-prey model of Holling type-IV functional response with stage structure and double delays due to maturation time for both prey and predator. The dynamical behavior of the system is investigated from the point of view of stability switches aspects. We assume that the immature and mature individuals of each species are divided by a fixed age, and the mature predator only attacks the mature prey. Based on some comparison arguments, sharp threshold conditions which are both necessary and sufficient for the global stability of the equilibrium point of predator extinction are obtained. The most important outcome of this paper is that the variation of predator stage structure can affect the existence of the interior equilibrium point and drive the predator into extinction by changing the maturation (through-stage) time delay. Our linear stability work and numerical results show that if the resource is dynamic, as in nature, there is a window in maturation time delay parameters that generate sustainable oscillatory dynamics

Moving forward in circles: challenges and opportunities in modelling population cycles

Population cycling is a widespread phenomenon, observed across a multitude of taxa in both laboratory and natural conditions. Historically, the theory associated with population cycles was tightly linked to pairwise consumer–resource interactions and studied via deterministic models, but current empirical and theoretical research reveals a much richer basis for ecological cycles. Stochasticity and seasonality can modulate or create cyclic behaviour in non-intuitive ways, the high-dimensionality in ecological systems can profoundly influence cycling, and so can demographic structure and eco-evolutionary dynamics. An inclusive theory for population cycles, ranging from ecosystem-level to demographic modelling, grounded in observational or experimental data, is therefore necessary to better understand observed cyclical patterns. In turn, by gaining better insight into the drivers of population cycles, we can begin to understand the causes of cycle gain and loss, how biodiversity interacts with population cycling, and how to effectively manage wildly fluctuating populations, all of which are growing domains of ecological research

Theoretical Study of Pest Control Using Stage Structured Natural Enemies with Maturation Delay: A Crop-Pest-Natural Enemy Model

In the natural world, there are many insect species whose individual members have a life history that takes them through two stages, immature and mature. Moreover, the rates of survival, development, and reproduction almost always depend on age, size, or development stage. Keeping this in mind, in this paper, a three species crop-pest-natural enemy food chain model with two stages for natural enemies is investigated. Using characteristic equations, a set of sufficient conditions for local asymptotic stability of all the feasible equilibria is obtained. Moreover, using approach as in (Beretta and Kuang, 2002), the possibility of the existence of a Hopf bifurcation for the interior equilibrium with respect to maturation delay is explored, which shows that the maturation delay plays an important role in the dynamical behavior of three species system. Also obtain some threshold values of maturation delay for the stability-switching of the particular system. In succession, using the normal form theory and center manifold argument, we derive the explicit formulas which determine the stability and direction of bifurcating periodic solutions. Finally, a numerical simulation for supporting the theoretical analysis is given.Comment: 28 pages, 9 figure

Robust permanence for ecological equations with internal and external feedbacks.

Species experience both internal feedbacks with endogenous factors such as trait evolution and external feedbacks with exogenous factors such as weather. These feedbacks can play an important role in determining whether populations persist or communities of species coexist. To provide a general mathematical framework for studying these effects, we develop a theorem for coexistence for ecological models accounting for internal and external feedbacks. Specifically, we use average Lyapunov functions and Morse decompositions to develop sufficient and necessary conditions for robust permanence, a form of coexistence robust to large perturbations of the population densities and small structural perturbations of the models. We illustrate how our results can be applied to verify permanence in non-autonomous models, structured population models, including those with frequency-dependent feedbacks, and models of eco-evolutionary dynamics. In these applications, we discuss how our results relate to previous results for models with particular types of feedbacks

Evolution of predator dispersal in relation to spatio-temporal prey dynamics : how not to get stuck in the wrong place!

Peer reviewedPublisher PD