Changes to the Fossil Record of Insects through Fifteen Years of Discovery

The first and last occurrences of hexapod families in the fossil record are compiled from publications up to end-2009. The major features of these data are compared with those of previous datasets (1993 and 1994). About a third of families (>400) are new to the fossil record since 1994, over half of the earlier, existing families have experienced changes in their known stratigraphic range and only about ten percent have unchanged ranges. Despite these significant additions to knowledge, the broad pattern of described richness through time remains similar, with described richness increasing steadily through geological history and a shift in dominant taxa, from Palaeoptera and Polyneoptera to Paraneoptera and Holometabola, after the Palaeozoic. However, after detrending, described richness is not well correlated with the earlier datasets, indicating significant changes in shorter-term patterns. There is reduced Palaeozoic richness, peaking at a different time, and a less pronounced Permian decline. A pronounced Triassic peak and decline is shown, and the plateau from the mid Early Cretaceous to the end of the period remains, albeit at substantially higher richness compared to earlier datasets. Origination and extinction rates are broadly similar to before, with a broad decline in both through time but episodic peaks, including end-Permian turnover. Origination more consistently exceeds extinction compared to previous datasets and exceptions are mainly in the Palaeozoic. These changes suggest that some inferences about causal mechanisms in insect macroevolution are likely to differ as well

New palaeodirections and palaeointensity data from extensive profiles through the Ediacaran section of the Volyn Basalt Province (NW-Ukraine)

Summary Changes of the geomagnetic field over geologic time scales can be used to study the evolution and processes of its sources in Earth's deep interior. As a stark contrast to the geomagnetic field behaviour of the recent past, the field in the late Neoproterozoic is defined by ambiguous polar wander paths, ultra-low field strengths and a period of extreme reversal hyperactivity. Palaeointensity data from this time are still scarce and conclusions made from these data suffer from large uncertainties brought by a low sampling density. In this study, we present new palaeomagnetic and palaeointensity data from quarry outcrops and five deep drill cores, covering the longest and most complete profiles through the Ediacaran units in the Volyn Basalt Province (NW-Ukraine) to date. Palaeodirectional analysis of thermal and alternating field demagnetisation experiments reveal at least six reversals in the profile as well as four palaeopoles that agree with previously published poles. Whole-rock palaeointensity estimates are derived from a multi-method approach that utilises thermal and microwave Thellier experiments in Coe- and IZZI-protocol as well as Wilson and double-heating-Shaw experiments. These produce instantaneous site-mean palaeointensity estimates in the range of 2.1±0.4 µT - 11.1±3.5 µT and virtual dipole moments in the range of 0.31±0.06 ×1022Am2 - 1.67±0.53 ×1022Am2. Small-scale variations of palaeointensities throughout the profile not only show extremely weak field strengths around polarity changes but could also suggest that the field strength never fully recovered between reversals. These new results increase the density of intensity data coverage for Baltica in the Late Ediacaran and, in combination with previously published Ediacaran dipole moments, support an extremely weak field that seems to increase in strength towards the Ediacaran-Cambrian transition.</jats:p