A reassessment of the problematic Ediacaran genus Orbisiana Sokolov 1976

The genus Orbisiana was established in 1976 by B.S. Sokolov based on a collection of seven specimens within core material from a borehole drilled through the Ediacaran-age Gavrilov Yam Formation of the Moscow Basin, Russia. Here we reassess the original material for the type species Orbisiana simplexSokolov 1976, which was long considered to be lost; fix the holotype of the type species; and revise the original diagnosis of the genus and species. Pyritisation of the fossils, which are preserved in finely laminated shales, allows three-dimensional morphological characterisation of this taxon using X-ray microtomography (μCT). Morphological and taphonomic analyses of the type material and additional three-dimensionally preserved specimens from the Verkhovka Formation (Vendian of the White Sea area) suggest that Orbisiana simplex consisted of submillimetric to millimetric globular chambers arranged in compact, grape-like clusters, or forming sinuous to linear aggregates. Occasionally, aggregates can bifurcate, with no appreciable change in chamber dimensions or shape. The phylogenetic affinity of Orbisiana remains uncertain, but its chambered construction, putative agglutinated structure of the chamber walls, and compact, occasionally branching chamber arrangement are shared with agglutinated tests of the Ediacaran genus Palaeopascichnus. Our reassessment and systematic study of the genus Orbisiana sheds new light on one of the least studied members of the late Ediacaran macroscopic biota.NER

PALAEONTOLOGICAL (RADIOLARIAN) LATE JURASSIC AGE CONSTRAINT FOR THE STEPANAVAN OPHIOLITE (LESSER CAUCASUS, ARMENIA)

Η χρονολόγηση με βάση μικροπαλαιοντολογικά δεδομένα, των ιζηματογενών καλυμμάτων των οφιολίθων είναι ιδιαίτερης σημασίας για την κατανόηση της παλαιογεωγραφικής και γεωδυναμικής εξέλιξης των περιοχών της Τηθύος. Η οφιολιθική ακολουθία του Stepanavan στη Βόρεια Αρμενία συνίσταται από περιδοτίτες, γάββρους, πλαγιογρανίτες και λάβες και ιζηματογενή καλύματα ραδιολαριτών. Θεωρείται ως η βόρεια επέκταση της οφιολιθικής ζώνης Sevan Akera και η ανατολική επέκταση της ζώνης Izmir-Ankara. Αντιπροσωπεύει κατάλοιπο μιας βραδέως εκτεινόμενης μεσο-ωκεάνιας ράχης, που ήταν ενεργή μεταξύ της Ευρασίας και του νότιου Αρμενικού τεμάχους γκοντβανικής προέλευσης. Τα ραδιολάρια που εξετάστηκαν από τους ραδιολαρίτες στην περιοχή της οφιολιθικής ακολουθίας του Stepanavan πιστοποιούν για πρώτη φορά ηλικία Ανώτερου Ιουρασικού (ανώτερο Κιμμερίδιο έως κατώτερο Τιθώνιο) προσδίδοντας ηλικία για αυτό το τμήμα του ωκεάνιου φλοιού της Τηθύος που παρατηρείται στο Lesser Caucasus.Micropalaeontological age evidence for the sedimentary cover of ophiolites is important to understand the palaeogeographic and geodynamic evolution of Tethyan realms. The Stepanavan ophiolitic suite of Northern Armenia consists of peridotites, gabbros, plagiogranite and lavas with a radiolarite sedimentary cover. It is regarded as the northern extension of the Sevan Akera ophiolitic zone and may be considered as the eastern extension of the Izmir-Ankara suture zone. It represents the relics of a slow-spreading mid oceanic ridge that was active between Eurasia and the South-Armenian Block of Gondwanian origin. Radiolaria extracted from radiolarites of the Stepanavan ophiolite provide for the first time a Late Jurassic (late Kimmeridgian to early Tithonian) age constraint for this part of Tethyan oceanic crust preserved in Lesser Caucasus

Palaeoceanographic implications of new and revised bio-chronostratigraphic constraints from the Profitis Ilias Unit (Rhodes, Greece)

Middle Oxfordian-early Kimmeridgian radiolaria, extracted from the top of radiolarites of Profitis Ilias unit (Rhodes island), suggest that the latter are essentially Middle Jurassic in age and the overlying siliceous shales Late Jurassic. The previously identified Calpionellid horizon at the top of Profitis Ilias siliceous marls is now regarded as early Valanginian in age. The above chronostratigraphic constraints allow tentative correlations to be made between Profitis Ilias and Pindos-Olonos sedimentary units. Finally, the palaeoceanographic significance of the studied series in Rhodes and potentially similar pelagic sequences in the Marmaris area of Turkey are discussed

New ophthalmosaurid ichthyosaurs from the European lower cretaceous demonstrate extensive ichthyosaur survival across the Jurassic–Cretaceous boundary

Background Ichthyosauria is a diverse clade of marine amniotes that spanned most of the Mesozoic. Until recently, most authors interpreted the fossil record as showing that three major extinction events affected this group during its history: one during the latest Triassic, one at the Jurassic–Cretaceous boundary (JCB), and one (resulting in total extinction) at the Cenomanian-Turonian boundary. The JCB was believed to eradicate most of the peculiar morphotypes found in the Late Jurassic, in favor of apparently less specialized forms in the Cretaceous. However, the record of ichthyosaurs from the Berriasian–Barremian interval is extremely limited, and the effects of the end-Jurassic extinction event on ichthyosaurs remains poorly understood. Methodology/Principal Findings Based on new material from the Hauterivian of England and Germany and on abundant material from the Cambridge Greensand Formation, we name a new ophthalmosaurid, Acamptonectes densus gen. et sp. nov. This taxon shares numerous features with Ophthalmosaurus, a genus now restricted to the Callovian–Berriasian interval. Our phylogenetic analysis indicates that Ophthalmosauridae diverged early in its history into two markedly distinct clades, Ophthalmosaurinae and Platypterygiinae, both of which cross the JCB and persist to the late Albian at least. To evaluate the effect of the JCB extinction event on ichthyosaurs, we calculated cladogenesis, extinction, and survival rates for each stage of the Oxfordian–Barremian interval, under different scenarios. The extinction rate during the JCB never surpasses the background extinction rate for the Oxfordian–Barremian interval and the JCB records one of the highest survival rates of the interval. Conclusions/Significance There is currently no evidence that ichthyosaurs were affected by the JCB extinction event, in contrast to many other marine groups. Ophthalmosaurid ichthyosaurs remained diverse from their rapid radiation in the Middle Jurassic to their total extinction at the beginning of the Late Cretaceous

Editorial: Siliceous plankton biomineralization, biodiversity and evolution

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Radiolarian evidence for Early Cretaceous (Late barremian-Early Aptian) submarine activity in the Tethyan oceanic realm preserved in Karabagh (Lesser Caucasus),

International audienceDating radiolarites in the ophiolitic sequences of Alpine mountain belts is important for the geodynamic reconstruction of Neo-Tethys in the Lesser Caucasus.However, radiolarian data from Karabagh are very rare. A moderately well-preserved radiolarian assemblage was obtained from radiolarites that overliepillow lavas cropping out west of the village of Vank in the mountainous region of Karabagh. The presence of the easily recognisable subspecies Aurisaturnaliscarinatus perforatus is of particular significance because it is the end member of a well studied lineage that went extinct in the early Aptian. Indeed, themiddle late Barremian - early early Aptian age range of this subspecies allows to suggest that submarine volcanic activity took place during this interval in theTethyan oceanic realm preserved in Karabagh. Given that this submarine volcanic event is more or less contemporaneous to the Aptian alkaline OIB-typelavas from the Vedi ophiolite, it is likely that the Karabagh lavas are also the expression of a volcanic activity associated to the emplacement of the sameoceanic volcanic plateau