New data about the suspensor of succulent angiosperms : ultrastructure and cytochemical study of the embryo-suspensor of Sempervivum arachnoideum L. and Jovibarba sobolifera (Sims) Opiz

The development of the suspensor in two species - Sempervivum arachnoideum and Jovibarba sobolifera - was investigated using cytochemical methods, light and electron microscopy. Cytological processes of differentiation in the embryo-suspensor were compared with the development of embryo-proper. The mature differentiated suspensor consists of a large basal cell and three to four chalazal cells. The basal cell produces haustorial branched invading ovular tissues. The walls of the haustorium and the micropylar part of the basal cell form the wall ingrowths typical for a transfer cells. The ingrowths also partially cover the lateral wall and the chalazal wall separating the basal cell from the other embryo cells. The dense cytoplasm filling the basal cell is rich in: numerous polysomes lying free or covering rough endoplasmic reticulum (RER), active dictyosomes, microtubules, bundles of microfilaments, microbodies, mitochondria, plastids and lipid droplets. Cytochemical tests (including proteins, insoluble polysaccharides and lipids are distributed in the suspensor during different stages of embryo development) showed the presence of high amounts of macromolecules in the suspensor cells, particularly during the globular and heart-shaped phases of embryo development. The protein bodies and lipid droplets are the main storage products in the cells of the embryo-proper. The results of Auramine 0 indicate that a cuticular material is present only on the surface walls of the embryo-proper, but is absent from the suspensor cell wall. The ultrastructural features and cytochemical tests indicate that in the two species - S. arachnoideum and J. sobolifera - the embryo-suspensor is mainly involved in the absorption and transport of metabolites from the ovular tissues to the developing embryo-proper

The battle over Syria's reconstruction

Reconstruction is becoming the new battleground in the Syrian conflict—its continuation by other means. It is instrumentalized by the regime as a way to reconsolidate its control over the country and by rival regional and international powers to shape the internal balance of power and establish spheres of influence in the country. The paper examines the Asad regime’s practices, including co-optation of militia leaders via reconstruction concessions and use of reconstruction to clear strategic areas of opposition-dominated urban settlements. The paper then surveys how the geopolitical struggle in Syria has produced an asymmetry as regards reconstruction: those powers that lost the geo-political contest on the ground seek to use geo-economic superiority to reverse the geo-political outcome. Then the impact of proxy wars and spheres of influence in the country on the security context for reconstruction is examined. Finally, the reconstruction initiatives of the various external parties are assessed, including Russia, Iran and Turkey as well as the spoiler role by which the US seeks to obstruct reconstruction that would spell victory in Syria for its Russian and Iranian rivals.PostprintPeer reviewe

A multiscale method for optimising surface topography in elastohydrodynamic lubrication (EHL) using metamodels

The frictional performance of a bearing is of significant interest in any mechanical system where there are lubricated surfaces under load and in relative motion. Surface topography plays a major role in determining the coefficient of friction for the bearing because the size of the fluid film and topography are of a comparable order. The problem of optimising topography for such a system is complicated by the separation in scales between the size of the lubricated domain and that of the topography, which is of at least one order of magnitude or more smaller. This paper introduces a multiscale method for optimising the small scale topography for improved frictional performance of the large scale bearing. The approach fully couples the elastohydrodynamic lubrication at both scales between pressure generated in the lubricant and deformation of the bounding surfaces. Homogenised small scale data is used to inform the large scale model and is represented using Moving Least Squares metamodels calibrated by cross validation. An optimal topography for a minimum coefficient of friction for the bearing is identified and comparisons made of local minima in the response, where very different topographies with similar frictional performance are observed. Comparisons of the optimal topography with the smooth surface model demonstrated the complexity of capturing the non-linear effect of topography and the necessity of the multiscale method in capturing this. Deviations from the smooth surface model were quantified by the metamodel coefficients and showed how topographies with a similar frictional performance have very different characteristics

On the Laminar Flow Characteristics of Conical Bearings. Part I—Analytical Approach

An analysis has been made of the characteristics of externally pressurized central recess conical bearings with nonconstant film thickness under the assumption of isothermal laminar flow of a viscous incompressible fluid. The flow in the narrow gaps that may be convergent, constant, or divergent has been approximately determined on the basis of the lubrication theory with convective inertia neglected except for that part which is due to rotation. The pressure distribution along the gap, the load capacity, and the torque of the bearings, were theoretically predicted. It was found that, while the gap is very narrow, both the load capacity and the torque for the convergent case decrease moderately, and for the divergent case increase significantly with respect to the constant gap bearing, provided with flow rate, rotational velocity and the film thickness at the outlet of the gap are the same.</jats:p

On the Laminar Flow Characteristics of Conical Bearings. Part II—Experimental Verification

The laminar flow characteristics of the externally pressurized central recess conical bearings with narrow gaps, predicted theoretically in Part I of this paper, have been verified experimentally. Two main effects on the performance of conical bearings have been investigated, namely: the effect of nonconstant film thickness along the gap of the bearing and the effect of rotation. It was found that experimental pressure distributions and load capacity of the constant and divergent gap bearings were in close agreement with the theoretical data for the low rotational velocities however, with the increased rotation, the experimental values were higher than theoretically predicted. The experimental values of pressure, load capacity, and the torque of convergent gap bearings were found higher than theoretical.</jats:p