24 research outputs found

    Pleistocene non-marine molluscs from Cova de Ca Na Reia, Eivissa

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    Els mol·luscs terrestres fòssils de les Illes Pitiüses

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    Enodicalix (Diploporita, Aristocystitidae): A new echinoderm genus from the Middle Ordovician of Spain

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    Restudy of an Ordovician diploporite specimen from the Middle Ordovician of central Spain reveals that the species ‘Calix’ inornatus MELÉNDEZ has four ambulacra, each with two equal facets, an oval hydropore and a smooth thecal surface. In contrast, the type species of Calix has four ambulacra, each with four facets added in a clockwise direction during growth, a trilobed hydropore, and a theca with spiny plates. Other aristocystitid genera with four ambulacra have one facet per ambulacrum. ‘Calix’ inornatus differs from all other aristocystitid genera and warrants a new generic name, for which we propose Enodicalix.

    Lateral variations in the topmost part of the Blue Lias and basal Charmouth Mudstone Formations (Lower Jurassic) on the Devon and Dorset coast

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    The beds adjacent to the junction of the Lower Jurassic Blue Lias and Charmouth Mudstone formations are intermittently exposed in cliff and foreshore sections over a distance of 8 km on the east Devon and west Dorset coast on either side of Lyme Regis. Comparison of the successions in the highest part of the Blue Lias shows little lateral variation in thickness or lithology, with the exception of minor thickness changes in the two highest limestone beds. In contrast, the basal beds of the Shales-with-Beef Member, the lowest part of the Charmouth Mudstone, are laterally variable. Up to five beds of limestone that are present in the most westerly exposure in Devon are absent at the more easterly exposures in Dorset. This lateral variation does not appear to be related to contemporaneous fault activity. It is largely due to an unconformity at the base of the Shales-with-Beef which cuts out successively more of the basal beds when traced from west to east. The strict application of the definition of the Blue Lias Formation, currently taken at the top of the highest limestone in an interbedded mudstone-limestone succession, would include beds previously classified as Shales-with-Beef in east Devon

    New insights into the origin and relationships of blastoid echinoderms

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    “Pan-dichoporites” (new informal term) is proposed to unite Cambrian blastozoans, such as Cambrocrinus, Ridersia, and Sanducystis, glyptocystitoid and hemicosmitoid rhombiferans, coronates, blastoids, and Lysocystites. Pan-dichoporite ambulacra have double biserial main axes with brachiole facets shared by pairs of floor (glyptocystitoids), side (blastoid) or trunk (hemicosmitoids, coronates) plates. These axial plates are the first two brachiolar plates modified to form the ambulacral axes. In glyptocystitoids the first brachiole facet in each ambulacrum is shared by an oral and another plate. Hence, these are also two modified brachiolar plates and part of the axial skeleton under the Extraxial Axial Theory (EAT). Pan-dichoporites are also characterized by thecae composed of homologous plate circlets. The unique glyptocystitoid genus Rhombifera bears ambulacral facets on five radial plates, which alternate with five orals. The oral area of Lysocystites (blastoid sensu lato) is very similar, which suggests that rhombiferan radials are homologous with “ambulacrals” of Lysocystites and hence with blastoid lancet plates. This implies derivation of blastoids from glyptocystitoids and suggests that blastoid and coronate radials and deltoids are homologous with rhombiferan infralaterals and laterals. Thus, homologous plate circlets occur in all pan-dichoporites, which strengthens the validity of a pan-dichoporite clade. Under Universal Elemental Homology (UEH), deltoids were homologized with rhombiferan orals, but this is inconsistent with the EAT. Deltoids bear respiratory pore structures and so are perforate extraxial skeletal plates, whereas rhombiferan orals are axial skeleton. The new plate homologies also explain why only five plates form the oral frames of coronates, blastoids and Lysocystites, whereas glyptocystitoids (except Rhombifera) have six oral frame plates; all glyptocystitoids have only five laterals. Hemicosmitoids arose by paedomorphic ambulacral reduction, but the paedomorphosis also affected the thecal plates and stem. Paedomorphosis poses special problems for cladistic character analysis, since the new characters often appear to be plesiomorphic

    Trace fossils

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    Book synopsis: The coastline of Dorset is spectacular and beautiful. It exposes a long squence of Jurassic age sedimentary rocks which, with its wealth of invertebrate, vertebrate and plant fossils, has long attracted palaeontologists amateur, professional, interested holiday–maker, dedicated collector and its international importance has been recognised by the designation as a UNESCO World Heritage Site. From the pioneering work of early collectors like Mary Anning onwards, the area has been a cradle of palaeontology. The Lower Lias is particularly fossiliferous and this lavishly illustrated Guide covers all the fossil groups to be found by the collector in the field

    Pentaradiate diploporites (Echinodermata) from the Spanish Middle Ordovician and their taxonomic significance

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    New Middle Ordovician specimens of pentaradiate diploporite blastozoans from the southern Central Iberian Zone, Spain, show that Oretanocalix is an aristocystitid diploporite with five ambulacra; all other aristocystitid genera have 2?4 ambulacra. The laterally elongate oral frame is composed of eight plates, with five, facet-bearing, circumorals (COO) and three, non-facet-bearing, periorals (POO). This necessitates a new interpretation of the aristocystitid oral area and diagnosis of the family Aristocystitidae. Previous interpretations assumed four COO and four POO. Weathered examples lack oral cover plates and show that plates CO1 and CO4, which support ambulacra D and B, respectively, do not reach the inner margin of the peristome. They also reveal abundant diplopores, which were covered by a thin epistereom in life. Diplopores grew at plate sutures and were incorporated into both adjacent plates as the plates enlarged. Oretanocalix julioi n. sp. extends the stratigraphic range of Oretanocalix back to the basal Dobrotivian ?low in the Upper Darriwilian. The genus Batalleria is a sphaeronitid diploporite with food grooves that branched within the peristome. The branching pattern reflects Lovén’s law and is: AR, BL, CR, DL, ER, where A?E denote Carpenter’s ambulacra and L (left) and R (right) denote on which side of the ambulacrum the first branch lies. Each branch leaves the peristome by a separate ambulacral orifice. No other sphaeronitids have food grooves that branch within the peristome and all other sphaeronitids have one ambulacral orifice per ambulacrum. Thus, a new diagnosis of sphaeronitids is necessary
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