Digestive structures in the Middle Ordovician trilobite Prionocheilus Rouault 1847, from the Barrandian area of Czech Republic

Remains of a digestive system from a slightly damaged articulated specimen of the comparatively rare bathycheilid trilobite Prionocheilus vokovicensis (Šnajdr, 1956) are described for the first time. The specimen comes from the Middle Ordovician Šárka Formation of the Prague Basin and contains the midgut region of the digestive system preserved through the axial region of glabella and six anterior thoracic segments. The anterior-most part of the digestive system is unknown as the anterior glabellar lobes are not preserved in the studied specimen. In the cephalic shield, the remains of two pairs of gut diverticulae are seen in the posterior region of the glabella. Remains of five pairs of small cavities developed in the axis of the first six thoracic segments represent the remains of thoracic gut diverticulae. The discussed specimen possess the first undoubted remain of digestive structures established within the family Bathycheilidae (PŘibyl, 1953)

Nomenclature of Cambrian lithostratigraphy of the Skryje-Týřovice Basin

Ecuador Debate es una publicación periódica electrónica dependiente del Centro Andino de Acción Popular (CAAP), que cubre diversos temas de la actualidad local y Latinoamericana. Centrada especialmente en asuntos económicos y políticos del Ecuador, la publicación también incluye importantes secciones dedicadas a la cultura (crítica literaria), el mundo agrario, social-urbano, y los temas de conflictividad social. Estos últimos son analizados desde un punto de vista de sociológico y cuantitat..

YORP and Yarkovsky effects in asteroids (1685) Toro, (2100) Ra-Shalom, (3103) Eger, and (161989) Cacus

The rotation states of small asteroids are affected by a net torque arising from an anisotropic sunlight reflection and thermal radiation from the asteroids' surfaces. On long timescales, this so-called YORP effect can change asteroid spin directions and their rotation periods. We analyzed lightcurves of four selected near-Earth asteroids with the aim of detecting secular changes in their rotation rates that are caused by YORP. We use the lightcurve inversion method to model the observed lightcurves and include the change in the rotation rate $\mathrm{d} \omega / \mathrm{d} t$ as a free parameter of optimization. We collected more than 70 new lightcurves. For asteroids Toro and Cacus, we used thermal infrared data from the WISE spacecraft and estimated their size and thermal inertia. We also used the currently available optical and radar astrometry of Toro, Ra-Shalom, and Cacus to infer the Yarkovsky effect. We detected a YORP acceleration of $\mathrm{d}\omega / \mathrm{d} t = (1.9 \pm 0.3) \times 10^{-8}\,\mathrm{rad}\,\mathrm{d}^{-2}$ for asteroid Cacus. For Toro, we have a tentative ($2\sigma$) detection of YORP from a significant improvement of the lightcurve fit for a nonzero value of $\mathrm{d}\omega / \mathrm{d} t = 3.0 \times 10^{-9}\,\mathrm{rad}\,\mathrm{d}^{-2}$. For asteroid Eger, we confirmed the previously published YORP detection with more data and updated the YORP value to $(1.1 \pm 0.5) \times 10^{-8}\,\mathrm{rad}\,\mathrm{d}^{-2}$. We also updated the shape model of asteroid Ra-Shalom and put an upper limit for the change of the rotation rate to $|\mathrm{d}\omega / \mathrm{d} t| \lesssim 1.5 \times 10^{-8}\,\mathrm{rad}\,\mathrm{d}^{-2}$. Ra-Shalom has a greater than $3\sigma$ Yarkovsky detection with a theoretical value consistent with observations assuming its size and/or density is slightly larger than the nominally expected values

Digestive and appendicular soft-parts, with behavioural implications, in a large Ordovician trilobite from the Fezouata Lagerstätte, Morocco

Trilobites were one of the most successful groups of marine arthropods during the Palaeozoic era, yet their soft-part anatomy is only known from a few exceptionally-preserved specimens found in a handful of localities from the Cambrian to the Devonian. This is because, even if the sclerotized appendages were not destroyed during early taphonomic stages, they are often overprinted by the three-dimensional, mineralised exoskeleton. Inferences about the ventral anatomy and behavioural activities of trilobites can also be derived from the ichnological record, which suggests that most Cruziana and Rusophycus trace fossils were possibly produced by the actions of trilobites. Three specimens of the asaphid trilobite Megistaspis (Ekeraspis) hammondi, have been discovered in the Lower Ordovician Fezouata Konservat-Lagerstätte of southern Morocco, preserving appendages and digestive tract. The digestive structures include a crop with digestive caeca, while the appendages display exopodal setae and slight heteropody (cephalic endopods larger and more spinose than thoracic and pygidial ones). The combination of these digestive structures and the heteropody has never been described together among trilobites, and the latter could assist in the understanding of the production of certain comb-like traces of the Cruziana rugosa group, which are extraordinarily abundant on the shallow marine shelves around Gondwana.This work has been supported by the Spanish Ministry of Economy and Competitiveness, project number CGL2012- 39471/BTE.Peer reviewe

Plated Cambrian Bilaterians Reveal the Earliest Stages of Echinoderm Evolution

Echinoderms are unique in being pentaradiate, having diverged from the ancestral bilaterian body plan more radically than any other animal phylum. This transformation arises during ontogeny, as echinoderm larvae are initially bilateral, then pass through an asymmetric phase, before giving rise to the pentaradiate adult. Many fossil echinoderms are radial and a few are asymmetric, but until now none have been described that show the original bilaterian stage in echinoderm evolution. Here we report new fossils from the early middle Cambrian of southern Europe that are the first echinoderms with a fully bilaterian body plan as adults. Morphologically they are intermediate between two of the most basal classes, the Ctenocystoidea and Cincta. This provides a root for all echinoderms and confirms that the earliest members were deposit feeders not suspension feeders

Animal Behavior Frozen in Time: Gregarious Behavior of Early Jurassic Lobsters within an Ammonoid Body Chamber

Direct animal behavior can be inferred from the fossil record only in exceptional circumstances. The exceptional mode of preservation of ammonoid shells in the Posidonia Shale (Lower Jurassic, lower Toarcian) of Dotternhausen in southern Germany, with only the organic periostracum preserved, provides an excellent opportunity to observe the contents of the ammonoid body chamber because this periostracum is translucent. Here, we report upon three delicate lobsters preserved within a compressed ammonoid specimen of Harpoceras falciferum. We attempt to explain this gregarious behavior. The three lobsters were studied using standard microscopy under low angle light. The lobsters belong to the extinct family of the Eryonidae; further identification was not possible. The organic material of the three small lobsters is preserved more than halfway into the ammonoid body chamber. The lobsters are closely spaced and are positioned with their tails oriented toward each other. The specimens are interpreted to represent corpses rather than molts. The lobsters probably sought shelter in preparation for molting or against predators such as fish that were present in Dotternhausen. Alternatively, the soft tissue of the ammonoid may have been a source of food that attracted the lobsters, or it may have served as a long-term residency for the lobsters (inquilinism). The lobsters represent the oldest known example of gregariousness amongst lobsters and decapods in the fossil record. Gregarious behavior in lobsters, also known for extant lobsters, thus developed earlier in earth's history than previously known. Moreover, this is one of the oldest known examples of decapod crustaceans preserved within cephalopod shells