Non-Abelian Born-Infeld Action and Type I - Heterotic Duality (II): Nonrenormalization Theorems

Type I - heterotic duality in D=10 predicts various relations and constraints on higher order F^n couplings at different string loop levels on both sides. We prove the vanishing of two-loop corrections to the heterotic F^4 terms, which is one of the basic predictions from this duality. Furthermore, we show that the heterotic F^5 and (CP even) F^6 couplings are not renormalized at one loop. These results strengthen the conjecture that in D=10 any Tr F^(2n) coupling appears only at the disc tree-level on type I side and at (n-1)-loop level on the heterotic side. Our non-renormalization theorems are valid in any heterotic string vacuum with sixteen supercharges.Comment: 35 pages, harvmac; cosmetic changes; final version to appear in NP

New D=4 gauged supergravities from N=4 orientifolds with fluxes

We consider classes of T_6 orientifolds, where the orientifold projection contains an inversion I_{9-p} on 9-p coordinates, transverse to a Dp-brane. In absence of fluxes, the massless sector of these models corresponds to diverse forms of N=4 supergravity, with six bulk vector multiplets coupled to N=4 Yang--Mills theory on the branes. They all differ in the choice of the duality symmetry corresponding to different embeddings of SU(1,1)\times SO(6,6+n) in Sp(24+2n,R), the latter being the full group of duality rotations. Hence, these Lagrangians are not related by local field redefinitions. When fluxes are turned on one can construct new gaugings of N=4 supergravity, where the twelve bulk vectors gauge some nilpotent algebra which, in turn, depends on the choice of fluxes.Comment: 51 pages, 1 figure. Latex. Reference added. Typos corrected. Discussion on gaugings expande

Effective superpotentials for compact D5-brane Calabi-Yau geometries

For compact Calabi-Yau geometries with D5-branes we study N=1 effective superpotentials depending on both open- and closed-string fields. We develop methods to derive the open/closed Picard-Fuchs differential equations, which control D5-brane deformations as well as complex structure deformations of the compact Calabi-Yau space. Their solutions encode the flat open/closed coordinates and the effective superpotential. For two explicit examples of compact D5-brane Calabi-Yau hypersurface geometries we apply our techniques and express the calculated superpotentials in terms of flat open/closed coordinates. By evaluating these superpotentials at their critical points we reproduce the domain wall tensions that have recently appeared in the literature. Finally we extract orbifold disk invariants from the superpotentials, which, up to overall numerical normalizations, correspond to orbifold disk Gromov-Witten invariants in the mirror geometry.Comment: 55 pages; v2: references added, typos correcte

N=4 Supergravity Lagrangian for Type IIB Orientifold on T^6/Z_2 in Presence of Fluxes and D3-Branes

We derive the Lagrangian and the transformation laws of N=4 gauged supergravity coupled to matter multiplets whose sigma-model of the scalars is SU(1,1)/U(1)x SO(6,6+n)/SO(6)xSO(6+n) and which corresponds to the effective Lagrangian of the Type IIB string compactified on the T^6/Z_2 orientifold with fluxes turned on and in presence of n D3-branes. The gauge group is T^12 x G where G is the gauge group on the brane and T^12 is the gauge group on the bulk corresponding to the gauged translations of the R-R scalars coming from the R-R four--form. The N=4 bulk sector of this theory can be obtained as a truncation of the Scherk-Schwarz spontaneously broken N=8 supergravity. Consequently the full bulk spectrum satisfies quadratic and quartic mass sum rules, identical to those encountered in Scherk-Schwarz reduction gauging a flat group. This theory gives rise to a no scale supergravity extended with partial super-Higgs mechanism.Comment: 49 pages, LaTex, 2 figures. Misprints corrected, more comments adde

Hierarchies from Fluxes in String Compactifications

Warped compactifications with significant warping provide one of the few known mechanisms for naturally generating large hierarchies of physical scales. We demonstrate that this mechanism is realizable in string theory, and give examples involving orientifold compactifications of IIB string theory and F-theory compactifications on Calabi-Yau four-folds. In each case, the hierarchy of scales is fixed by a choice of RR and NS fluxes in the compact manifold. Our solutions involve compactifications of the Klebanov-Strassler gravity dual to a confining N=1 supersymmetric gauge theory,and the hierarchy reflects the small scale of chiral symmetry breaking in the dual gauge theory.Comment: 35 pages. v2: minor eqn. and reference change

Size Doesn't Matter: Towards a More Inclusive Philosophy of Biology

notes: As the primary author, O’Malley drafted the paper, and gathered and analysed data (scientific papers and talks). Conceptual analysis was conducted by both authors.publication-status: Publishedtypes: ArticlePhilosophers of biology, along with everyone else, generally perceive life to fall into two broad categories, the microbes and macrobes, and then pay most of their attention to the latter. ‘Macrobe’ is the word we propose for larger life forms, and we use it as part of an argument for microbial equality. We suggest that taking more notice of microbes – the dominant life form on the planet, both now and throughout evolutionary history – will transform some of the philosophy of biology’s standard ideas on ontology, evolution, taxonomy and biodiversity. We set out a number of recent developments in microbiology – including biofilm formation, chemotaxis, quorum sensing and gene transfer – that highlight microbial capacities for cooperation and communication and break down conventional thinking that microbes are solely or primarily single-celled organisms. These insights also bring new perspectives to the levels of selection debate, as well as to discussions of the evolution and nature of multicellularity, and to neo-Darwinian understandings of evolutionary mechanisms. We show how these revisions lead to further complications for microbial classification and the philosophies of systematics and biodiversity. Incorporating microbial insights into the philosophy of biology will challenge many of its assumptions, but also give greater scope and depth to its investigations

Systematics of the Neotropical Genus Leptodactylus Fitzinger, 1826 (Anura: Leptodactylidae): Phylogeny, the Relevance of Non-molecular Evidence, and Species Accounts

A phylogeny of the species-rich clade of the Neotropical frog genus Leptodactylus sensu stricto is presented on the basis of a total evidence analysis of molecular (mitochondrial and nuclear markers) and non-molecular (adult and larval morphological and behavioral characters) sampled from > 80% of the 75 currently recognized species. Our results support the monophyly of Leptodactylus sensu stricto, with Hydrolaetare placed as its sister group. The reciprocal monophyly of Hydrolaetare and Leptodactylus sensu stricto does not require that we consider Hydrolaetare as either a subgenus or synonym of Leptodactylus sensu lato. We recognize Leptodactylus sensu stricto, Hydrolaetare, Adenomera, and Lithodytes as valid monophyletic genera. Our results generally support the traditionally recognized Leptodactylus species groups, with exceptions involving only a few species that are easily accommodated without proposing new groups or significantly altering contents. The four groups form a pectinate tree, with the Leptodactylus fuscus group diverging first, followed by the L. pentadactylus group, which is sister to the L. latrans and L. melanonotus groups. To evaluate the impact of non-molecular evidence on our results, we compared our total evidence results with results obtained from analyses using only molecular data. Although non-molecular evidence comprised only 3.5% of the total evidence matrix, it had a strong impact on our total evidence results. Only one species group was monophyletic in the molecular-only analysis, and support differed in 86% of the 54 Leptodactylus clades that are shared by the results of the two analyses. Even though no non-molecular evidence was included for Hydrolaetare, exclusion of that data partition resulted in that genus being nested within Leptodactylus, demonstrating that the inclusion of a small amount of non-molecular evidence for a subset of species can alter not only the placement of those species, but also species that were not scored for those data. The evolution of several natural history and reproductive traits is considered in the light of our phylogenic framework. Invasion of rocky outcrops, larval oophagy, and use of underground reproductive chambers are restricted to species of the Leptodactylus fuscus and L. pentadactylus groups. In contrast, larval schooling, larval attendance, and more complex parental care are restricted to the L. latrans and L. melanonotus groups. Construction of foam nests is plesiomorphic in Leptodactylus but their placement varies extensively (e.g., underground chambers, surface of waterbodies, natural or excavated basins). Information on species synonymy, etymology, adult and larval morphology, advertisement call, and geographic distribution is summarized in species accounts for the 30 species of the Leptodactylus fuscus group, 17 species of the L. pentadactylus group, eight species of the L. latrans group, and 17 species of the L. melanonotus group, as well as the three species that are currently unassigned to any species group.Se presenta una filogenia del género Leptodactylus, un ciado neotropical rico en especies, basada en análises combinados de datos moleculares (marcadores nuclear y mitocondriales) y no moleculares (caracteres de la morfología de adultos y larvas así como de comportamiento) se muestrearon > 80% de las 75 especies reconocidas. Los resultados apoyan la monofília de Leptodactylus sensu stricto, con Hydrolaetare como su grupo hermano. La monofília recíproca de Hydrolaetare y Leptodactylus no requiere considerar a Hydrolaetare como un subgénero o sinónimo de Leptodactylus sensu lato. Se reconocen Leptodactylus sensu stricto, Hydrolaetare, Adenomera y Lithodytes como géneros monofiléticos válidos. Los resultados en general resuelven los grupos tradicionalmente reconocidos de Leptodactylus, con excepciones de algunas especies que son reasignadas sin la necesidad de proponer nuevos grupos o alterar significativamente el contenido de los grupos tradicionales. Los cuatro grupos de especies forman una topología pectinada donde el grupo de L. fuscus tiene una posición basal, seguido por el grupo de L. pentadactylus que es el grupo hermano al clado formado por los grupo de L. latrans y L. melanonotus. Se estimó el impacto de los datos no moleculares en los resultados, comparándose los resultados de evidencia total con los de los análises de datos moleculares solamente. Los datos no moleculares representan un 3.5% de la matriz de evidencia total, pero estos datos tuvieron un impacto significativo en los resultados del análisis de evidencia total. En el análisis estrictamente molecular solamente un grupo de especies resultó monofilético, y el apoyo difirió en 86% de los 54 ciados de Leptodactylus compartidos entre los dos análises. A pesar que datos no moleculares no fueron incluidos para Hydrolaetare, la exclusión de evidencia no molecular resultó en el género estar dentro de Leptodactylus, demostrando que la inclusión de evidencia no molecular pequeña para un subgrupo de especies altera no solamente la posición topológica de esas especies, sino tambien de las especies para las cuales dichos datos no fueron codificados. La evolución de patrones de historia natural y reprodución se evalúan en el contexto filogenético. La invasión de afloramientos rocosos y la construción de cámaras de reprodución subterraneas está limitada a los grupos de Leptodactylus fuscus y L. pentadactylus, mientras que la oofagia larval está restringida al grupo de L. pentadactylus. Por otro lado, los cárdumenes larvales, la proteción del cárdumen, y otros comportamientos parentales complejos carecterizan al clado formado por los grupos de especies de L. latrans y L. melanonotus. Los resúmenes de especies incluyen información de sinonimias, etimología, morfología de adultos y larvas, cantos, y distribución geográfica para las 30 especies del grupo de Leptodactylus fuscus, 17 especies del grupo L. pentadactylus, ocho especies del grupo de L. latrans, 17 especies del grupo de L. melanonotus, así como para las tres especies que actualmente no se encuentran asociadas a ninguno de los grupos de especies.Taran Grant was supported by Conselho Nacional de Desenvolvimento Científico e Tecnológico Proc. 307001/2011-3 and Fundação de Amparo à Pesquisa do Estado de São Paulo Proc. 2012/10000-5

Complex mosaic of sexual dichromatism and monochromatism in Pacific robins results from both gains and losses of elaborate coloration

Pacific robins exhibit one of the most complex range-wide mosaics of sexual dichromatism and monochromatism. The evolutionary origins of this geographic mosaic remain poorly understood despite long-standing interest from ornithologists, and its influential role in the development of Ernst Mayr’s theories on speciation and the 'Biological Species Concept'. One factor limiting our understanding of the evolution of sexual plumage variation in Pacific robins is a lack of well-resolved taxon boundaries and phylogenetic relationships in the group. Here, we use primarily historical museum specimens to obtain dense sampling of mtDNA, nuclear DNA, plumage color and morphometrics from all named taxa in the radiation in order to infer taxon boundaries and relationships. We use these data to test hypotheses about colonization history, plumage evolution and reduced island dichromatism. Our data show that the Pacific robin radiation comprises four distinct lineages, which warrant recognition as separate species – the previously recognized Norfolk robin P. multicolor and redcapped robin P. goodenovii, and two new species we propose to name: ‘Solomon robin’ P. polymorpha Mayr, 1934 for the populations on Solomon and Bougainville Islands, and ‘Mayr’s robin’ P. pusilla Peale, 1848 (in honor of Ernst Mayr’s detailed work on the southwest Pacific robins) for the populations on Vanuatu, Fiji and Samoa. Our data suggest that the common ancestor of the entire Pacific robin radiation was most likely sexually dichromatic and that the radiation-wide mosaic of sexual plumage color arose via repeated losses of elaborate plumage in males and gains of elaborate plumage in females on separate islands.Anna M. Kearns, Leo Joseph, Jeremy J. Austin, Amy C. Driskell and Kevin E. Omlan