Estudo taxonômico das espécies de Pipunculidae (Diptera) do Nordeste brasileiro

We conducted a taxonomic study of Pipunculidae species from northeastern Brazil, with specimens from the states of Maranhão, Piauí, Ceará, Paraíba, Rio Grande do Norte and Bahia. The identification was made with the aid of dichotomous keys, original descriptions, redescriptions and by direct comparison with types. For specimen’s identification and illustration it was made dissection of male terminalia. It was examined 614 specimens, belonging to 44 species, distributed in nine known genera and in a new genus propose here. Tomosvaryella scopulata Hardy is recorded for the first time for Brazil. Eighteen species represent new records for the northeastern Brazil; 24 new species are being described and illustrated with the addition of new characters. The species Cephalosphaera miriamae Rafael and Cephalosphaera semispiralis Rafael & Rosa had their male terminalias redescribed based on type specimens and on specimens collected in the Northeastern. Illustrated identification keys are provided for genera and species recorded from Northeastern Brazil, based on external morphological characters and especially based on male terminalia. Diagnosis, comments and maps with geographic records of all species were included.Foi realizado um estudo taxonômico das espécies de Pipunculidae do Nordeste brasileiro, com espécimes oriundos dos estados do Maranhão, Piauí, Ceará, Paraíba, Rio Grande do Norte e Bahia. Para identificação e ilustração dos espécimes foram dissecadas as terminálias masculinas. A identificação das espécies foi feita com auxílio de chaves dicotômicas, descrições originais, redescrições e quando possível, por comparação direta com tipos. Foram examinados 614 espécimes, pertencentes a 44 espécies, distribuídas em nove gêneros já conhecidos e um gênero novo proposto neste trabalho. Tomosvaryella scopulata Hardy é registrada pela primeira vez para o Brasil. Dezoito espécies representam novos registros para o Nordeste brasileiro; 24 espécies novas são descritas e ilustradas com a adição de novos caracteres. As espécies Cephalosphaera miriamae Rafael e Cephalosphaera semispiralis Rafael & Rosa tiveram suas terminálias masculinas redescritas baseadas nos tipos e em exemplares coletados no Nordeste. São fornecidas chaves de identificação ilustradas para os gêneros e espécies registradas para o Nordeste brasileiro, baseadas em caracteres morfológicos externos e principalmente das terminálias masculinas. Foram incluídos diagnoses e comentários, além de mapas com os registros geográficos das espécies

Tabanidae (Diptera) of Maranhão state, Brazil. V. Description of Protosilvius gurupi sp. n. (Pangoniinae, Pangoniini) and key to Protosilvius species

Protosilvius gurupi sp. n. (Tabanidae, Pangoniinae) is described and illustrated based on seven female and 53 male specimens collected in the Amazonian region at Reserva Biológica Gurupi, Centro Novo do Maranhão municipality, northwest Maranhão, Brazil. This is the first record of Protosilvius in northern Brazil and in the Amazon Basin. An illustrated key to all Protosilvius species is also presented. © José Albertino Rafael et al

Inpauema, a new genus of Odiniidae (Diptera) from Brazil, with description of five new species

A new genus and five new species of odiniids (Odiniidae: Traginopinae) are described from the Brazilian Amazon and Cerrado biomes: Inpauema mirador gen. nov. et sp. nov. (type species), I. catarinae sp. nov., I. gaimarii sp. nov., I. raimundoluizi sp. nov., and I. xavieri sp. nov. The genus is being characterized by a unique combination of diagnostic characters: body predominantly dark brown to black, with silvery-gray pruinose spots on inner margin of eyes, longitudinally along middle of lunule and face, on notopleuron and mesopleuron; postcranium concave from dorsal view; one pair of stout proclinate ocellar setae; postocellar setae absent; lunule shorter than frons; gena lacking upturned seta; antennae separated by a maximum distance of 2X the diameter of a single antennal socket and gonocoxal apodemes directed upward, forming an arch. A key to separate Helgreelia Gaimari, 2007 from Inpauema gen. nov. and for the new species is provided. Copyright © 2017 Magnolia Press

Inpauema, a new genus of Odiniidae (Diptera) from Brazil, with description of five new species

A new genus and five new species of odiniids (Odiniidae: Traginopinae) are described from the Brazilian Amazon and Cerrado biomes: Inpauema mirador gen. nov. et sp. nov. (type species), I. catarinae sp. nov., I. gaimarii sp. nov., I. raimundoluizi sp. nov., and I. xavieri sp. nov. The genus is being characterized by a unique combination of diagnostic characters: body predominantly dark brown to black, with silvery-gray pruinose spots on inner margin of eyes, longitudinally along middle of lunule and face, on notopleuron and mesopleuron; postcranium concave from dorsal view; one pair of stout proclinate ocellar setae; postocellar setae absent; lunule shorter than frons; gena lacking upturned seta; antennae separated by a maximum distance of 2X the diameter of a single antennal socket and gonocoxal apodemes directed upward, forming an arch. A key to separate Helgreelia Gaimari, 2007 from Inpauema gen. nov. and for the new species is provided. Copyright © 2017 Magnolia Press

Hexapoda Yearbook (Arthropoda: Mandibulata: Pancrustacea) Brazil 2020: the first annual production survey of new Brazilian species

This paper provided a list of all new Brazilian Hexapoda species described in 2020. Furthermore, based on the information extracted by this list, we tackled additional questions regarding the taxa, the specialists involved in the species descriptions as well as the journals in which those papers have been published. We recorded a total of 680 new Brazilian species of Hexapoda described in 2020, classified in 245 genera, 112 families and 18 orders. These 680 species were published in a total of 219 articles comprising 423 different authors residing in 27 countries. Only 30% of these authors are women, which demonstrates an inequality regarding sexes. In relation to the number of authors by species, the majority of the new species had two authors and the maximum of authors by species was five. We also found inequalities in the production of described species regarding the regions of Brazil, with Southeast and South leading. The top 10 institutions regarding productions of new species have four in the Southeast, two at South and with one ate North Region being the outlier of this pattern. Out of the total 219 published articles, Zootaxa dominated with 322 described species in 95 articles. The average impact factor was of 1.4 with only seven articles being published in Impact Factors above 3, indicating a hardship on publishing taxonomic articles in high-impact journals.The highlight of this paper is that it is unprecedent, as no annual record of Hexapoda species described was ever made in previous years to Brazil.Fil: Silva Neto, Alberto Moreira. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Lopes Falaschi, Rafaela. Universidade Estadual do Ponta Grossa; BrasilFil: Zacca, Thamara. Universidade Federal Do Rio de Janeiro. Museu Nacional; BrasilFil: Hipólito, Juliana. Universidade Federal da Bahia; BrasilFil: Costa Lima Pequeno, Pedro Aurélio. Universidade Federal de Roraima; BrasilFil: Alves Oliveira, João Rafael. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Oliveira Dos Santos, Roberto. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Heleodoro, Raphael Aquino. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Jacobina, Adaiane Catarina Marcondes. Universidade Federal do Paraná; BrasilFil: Somavilla, Alexandre. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Camargo, Alexssandro. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: de Oliveira Lira, Aline. Universidad Federal Rural Pernambuco; BrasilFil: Sampaio, Aline Amanda. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: da Silva Ferreira, André. Universidad Federal Rural Pernambuco; BrasilFil: Martins, André Luis. Universidade Federal do Paraná; BrasilFil: Figueiredo de Oliveira, Andressa. Universidade Federal do Mato Grosso do Sul; BrasilFil: Gonçalves da Silva Wengrat , Ana Paula. Universidade do Sao Paulo. Escola Superior de Agricultura Luiz de Queiroz; BrasilFil: Batista Rosa, Augusto Henrique. Universidade Estadual de Campinas; BrasilFil: Dias Corrêa, Caio Cezar. Universidade Federal Do Rio de Janeiro. Museu Nacional; BrasilFil: Costa De-Souza, Caroline. Museu Paraense Emilio Goeldi; BrasilFil: Anjos Dos Santos, Danielle. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Patagonia Norte. Centro de Investigación Esquel de Montaña y Estepa Patagónica. Universidad Nacional de la Patagonia "San Juan Bosco". Centro de Investigación Esquel de Montaña y Estepa Patagónica; ArgentinaFil: Pacheco Cordeiro, Danilo. Instituto Nacional Da Mata Atlantica; BrasilFil: Silva Nogueira, David. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Almeida Marques, Dayse Willkenia. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Nunes Barbosa, Diego. Universidade Federal do Paraná; BrasilFil: Mello Mendes, Diego Matheus. Instituto de Desenvolvimento Sustentável Mamirauá; BrasilFil: Galvão de Pádua, Diego. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Silva Vilela, Diogo. Universidade Estadual Paulista Julio de Mesquita Filho; BrasilFil: Gomes Viegas, Eduarda Fernanda. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; BrasilFil: Carneiro dos Santos, Eduardo. Universidade Federal do Paraná; BrasilFil: Rodrigues Fernandes, Daniell Rodrigo. Ministério da Ciência, Tecnologia, Inovações. Instituto Nacional de Pesquisas da Amazônia; Brasi

Checklist of the dipterofauna (Insecta) from Roraima, Brazil, with special reference to the Brazilian Ecological Station of Maracá

Roraima is a Brazilian state located in the northern portion of the Amazon basin, with few studies regarding its biodiversity. The Ecological Station of Maracá (Brazil, state of Roraima) harbors the third largest Brazilian pluvial island and is composed of a transitional landscape of savanna and Amazon rainforest components. Despite its ecological importance and strategic localization, few studies covered the dipterofauna of this locality. An updated checklist addressing 41 families of true flies (Diptera) occurring in Roraima is presented based on the literature and the specimens collected during a field expedition that occurred in 2015. This checklist brings several improvements such as new records of 165 taxa to the state of Roraima, 29 taxa to Brazil, and 259 morphotypes, mostly likely representing undescribed species

Macrostomus arcucinctus

Macrostomus arcucinctus (Bezzi, 1909) (Figs 1 A–D) Rhamphomyia arcucincta Bezzi, 1909: 331 (syntypes). Melander, 1928: 186; Collin, 1933: 211 (cit). Macrostomus arcucinctus: Smith, 1961: 53; 1967: 29 (cat.); Rafael & Cumming, 2004: 446. Diagnosis. Females of this species are distinguished by the thorax and abdomen subshiny dark brown to black with blue reflections; mesonotum densely gray pruinose. Legs mainly yellow to dark yellow. Mid femur with small posteroventral pennate setae on distal half. Apex of the wing infuscate; anal lobe narrow. Halter white. Abdomen shiny brown to black. Redescription. Lectotype female. Body length from original description: “ 3.2 mm; wing length 3.0 mm”. Head narrowly dichoptic. Inferior facets slightly larger than superior ones. Frons shiny black. One pair of proclinate ocellar setae. Face gray pruinose, slightly narrower than frons. Postcranium narrow, gray pruinose, with blue reflection in certain light. Inner vertical seta stouter than outer vertical seta, latter subequal in length to uniseriate postocular row of setae; remaining occipital setae slightly shorter than postoculars and restricted to dorsal half. Antenna dark brown, gold pruinose, short pubescent. Proboscis yellow, as long as head height. Thorax subshiny dark brown to black with blue reflection, mesonotum densely gray pruinose, less pruinose medially on scutum. Thoracic chaetotaxy: 4 antepronotals weak; 1 postpronotal stout, 2–3 weak; 4 dorsocentrals; 1 presutural supra-alar; 0 postsutural supra-alar; 1 postalar; 2 small proepisternals; 3 notopleural, anterior seta weaker; 2 pairs of scutellars (broken); 8 laterotergitals. Legs yellow to dark yellow except distal tarsomeres darker. Legs longer setae: mid tibia with 1 dorsal at middle slightly stouter and 1 anteroventral, 1 posteroventral somewhat inconspicuous distally. Mid femur with posteroventral pennate setae on distal half, thinner than in other species with this kind of setae (originally described as “non pennatis”). Hind legs lost. Wings lost, described as: “ alis brevibus hyalinis, macula stigmatica at area apicali fuscis, angulo axillari valde aperto, cellula discoidali magna nervos duos primos praeter basin decolores emittente ” [wing hyaline slightly brown infuscated on pterostigma and apex of veins R 2+3 and R 4+5; anal lobe narrow, distinct, with axillary angle obtuse; veins M 1 and M 2 evanescent]. Halter white. Abdomen shiny brown to black, somewhat blue depending on angle of light incidence, with small setae. Sternites concolorous with tergites. Tergite 8 (Figs 1 A–B) subrectangular in dorsal view, with shallow basal and deeper distal sinus, basally with narrow sclerotized sub-basal band. Sternite 8 (Figs 1A, C) longer than respective tergite, higher at basal half in lateral view, slightly more sclerotized medially, with apex somewhat acute. Tergite 9+10, sternite 10 and cerci lost. Genital fork (Fig. 1D) short, Y-shaped. Genital chamber membranous. Male. Unknown. Geographical records. Bolivia (La Paz). Type material examined. LECTOTYPE ♀ (here designated): “ BOLIVIA [La Paz], Mapiri, Sarampioni, 31.iii. 1903, 700 m. ” (SMTD). Lectotype condition. Wings and hind legs lost. Abdomen in microvial with glycerin. Remarks. The single remaining syntype specimen in the SMTD collection is designated lectotype accordingly to fix and stabilize the current concept of the name.Published as part of Rafael, José Albertino & Marques, Dayse Willkenia Almeida, 2019, Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia, pp. 251-275 in Zootaxa 4567 (2) on pages 252-253, DOI: 10.11646/zootaxa.4567.2.3, http://zenodo.org/record/259493

Macrostomus montanus Rafael & Marques 2019, sp. nov.

Macrostomus montanus sp. nov. (Figs 6 A–N, 7A–E) urn:lsid:zoobank.org:act: 9EB41EDC-134E-4C71-802C-6A6E9ACDB185 Diagnosis. Gena and postgena with setae somewhat stout; wing almost entirely hyaline, slightly brown infuscate at apex of costal cell, pterostigma and apex of cell r 2+3; last section of vein M 4 longer than vein dm-m; cell dm somewhat acute; tergite 8 short with distal sinus; epandrium with posterodorsal lobe longer than anterodorsal lobe, with longer setae more concentrated posteriorly on posterodorsal lobe; posterior cercus inwardly curved, trifid, apparently glabrous; hypandrium well sclerotized, slightly wider medially at level of posterior longer setae. Description. Holotype ♂ (Fig. 6A). Body length 3.5 mm; wing length 3.2 mm. Head dichoptic. Frons shiny black, slightly wider than anterior ocellus width. Two pairs of proclinate ocellar setae, posterior pair weaker. Face narrower than frons width, same as anterior ocellus width. Postcranium shiny black, entirely and sparsely gray pruinose. Inner vertical seta stouter than outer vertical seta, latter subequal in length to uniseriate postocular row of setae; 2–3 pairs of dorsal occipital setae. Gena with 3 setae and postgena with 2 somewhat stout setae. Antenna velvety black with postpedicel short pubescent. Proboscis shiny, brown basally, yellow apically. Palpus black. Thorax shiny black, gray pruinose. Thoracic chaetotaxy: 4 antepronotals; 1 postpronotal stout and 2 minutes; 4 dorsocentrals (right row with 5); 1 presutural supra-alar; 1 postsutural supra-alar; 1 postalar stout, 1 minute; 4 proepisternals weak; 1 notopleural stout and 1 weak; 2 pairs of scutellars, outer pair slightly weaker; 7 laterotergitals. Legs yellow with distal 3/4 of fore tibia, apex of mid femur, entire mid tibia, distal half of hind femur, entire hind tibia, all tarsi brown to black. Legs longer setae: fore tibia with 1 sub-basal and 1 median anterodorsal slightly longer; mid femur with 1 subapical anterior slightly longer; mid tibia with alternating anterodorsal and posterodorsal equally spaced; hind femur with 1 anterior subapical; hind tibia (Fig. 6A) with 4–5 slender dorsal and hind tarsomere 1 with 1 longer median dorsal seta. Wing (Fig. 6A) almost entirely hyaline, slightly brown infuscate at apex of costal cell, pterostigma and apex of cell r 2+3. Veins M 1 and M 2 weaker and apex of CuA+CuP evanescent. Last section of vein M 4 longer than vein dm-m. Cell dm somewhat acute. Halter light yellow. Abdomen (Fig. 6B) shiny black; tergites 1–3 with longer setae posteriorly. Tergite 8 (Fig. 6C) with distal sinus. Sternite 8 (Fig. 6D) with distinct saddle-shaped projection posterodorsally and deep basal sinus. Tergite 8 articulated with sternite 8 somewhat horizontally (Fig. 6E). Epandrium (Fig. 6F) with posterodorsal lobe longer than anterodorsal lobe, with longer setae more concentrated posteriorly on posterodorsal lobe; anterodorsal cleft minute, inconspicuous (smaller seta on Fig. 6F); anteroventral cleft distinct (stronger seta on Fig. 6F). Anterior cercus (Figs 6G, J) small in lateral view, well sclerotized, with longer setae dorsally and posteriorly; when viewed anteriorly (Fig. 6H) emitting mesially large and flat cercal bridge connected medially; cercal bridge expanded posteriorly and fused with upper hypoproctal lobe when viewed dorsally (Fig. 6I). Median cercus (Fig. 6J) somewhat cup-shaped with longer setae dorsally. Posterior cercus (Fig. 6J) inwardly curved, trifid, apparently glabrous. Ventral projection of cercus (Fig. 6K) horizontally placed, distally expanded and setose. Dorsal lobe of hypoproct somewhat inconspicuous, fused with cercal bridge expansion of anterior cercus; ventral lobe of hypoproct distinctly setose, placed between setose lobes of ventral projection of cercus and cercal bridge of anterior cercus. Hypandrium (Figs 6L, M) well sclerotized, almost same width throughout, slightly wider medially at level of posterior longer setae; distally with two pairs of curved hooks, distal pair stouter. Ejaculatory apodeme (Fig. 6L) tetralamellar, all lamella subequal. Phallus (Fig. 6L) slightly longer than hypandrium. Female. Frons wider than width of anterior ocellus, wider than in male. Face slightly narrower than frons, approximately as wide as width of anterior ocellus. Wing distinctly more infuscated at costal and distal margins (Fig. 7A). Tergite 8 (Figs 7B, C) wider basally in lateral view; with shallow basal sinus and deep distal sinus in dorsal view. Sternite 8 (Figs 7B, D, E) highly modified; in lateral and ventral views widened distally and with ventral subapical small translucid protuberance. Genital fork (Figs 7D, E) T-shaped with arms upwardly directed. Genital chamber (Fig. 7D, E) well sclerotized, wide, U-shaped, with basal protuberance upwardly directed. Geographical records. Bolivia (La Paz). Type material examined. HOLOTYPE ♂, paratype ♀, on same pin: “ BOLIVIA, La Paz, Chulumani, cloud forest, above 2000 m [eters]”; “ In copula, Chulumani, 5.iv. [19]79”; “QR code: NHMUK 010664427 ”; “Holot ♂ Parat ♀ Macrostomus montanus Rafael & Marques ” (Fig. 6N) (NHMUK). PARATYPES: Bolivia: same data as holotype, except 2.iv. [19]79, QR code: NHMUK 0 10864440 (1♂, NHMUK); NHMUK 0 10664353 (1♀, INPA); NHMUK 0 10664429 (1♀, NHMUK); NHMUK 0 10664436 (1♀, NHMUK); NHMUK 0 10664441 (1♀, NHMUK); NHMUK 0 10664442 (1♀, INPA); same data, except 25.iii. [19]79, NHMUK 0 10664430 (1♀, NHMUK, dissected); La Paz, Sarampionim [sic], 23.ii.2005, Arm. Malaise, J. Rodriguez (1♀, CBF). Holotype condition. Good. Left hind leg lost; right hind leg glued on piece of white card; abdomen not dissected. Etymology. From the Latin montanus = mountain and refers to the high elevation where the specimens were collected. Variation. One specimen (QR code: NHMUK 010864440) with three pairs of ocellar setae; the additional anterior pair slightly longer than posterior pair. Remarks. Macrostomus montanus sp. nov. is tentatively included in the M. limbipennis species-group, however, it differs mainly by having epandrium with posterodorsal lobe longer than the anterodorsal lobe. In the key presented by Rafael & Cumming (2012), this species runs to couplet 3 separating M. limbipennis (Bezzi) from M. penai Rafael & Cumming, but it differs from the first species by male tergite 8 not humped and from the second species by all cercal lobes shortened (versus elongate and upwardly directed in M. penai), posterior lobe of epandrium longer than anterior lobe (versus same length) and by female sternite 8 with a ventral subapical small translucid protuberance (versus absent), genital fork T-shaped with arms upwardly directed (versus small rod, without lateral arms), genital chamber wide, U-shaped, well sclerotized (versus membranous).Published as part of Rafael, José Albertino & Marques, Dayse Willkenia Almeida, 2019, Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia, pp. 251-275 in Zootaxa 4567 (2) on pages 259-261, DOI: 10.11646/zootaxa.4567.2.3, http://zenodo.org/record/259493

FIGURE 2 in Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia

FIGURE 2. Macrostomus argyrotarsis (Bezzi, 1909). A. Wing, holotype ♀; B. Left leg, posterior view, peruvian specimen; C. Peruvian specimen pinned, lateral view. Scale bars, figures A–C = 1 mm.Published as part of Rafael, José Albertino & Marques, Dayse Willkenia Almeida, 2019, Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia, pp. 251-275 in Zootaxa 4567 (2) on page 254, DOI: 10.11646/zootaxa.4567.2.3, http://zenodo.org/record/259493

Macrostomus argyrotarsis

Macrostomus argyrotarsis (Bezzi, 1909) (Figs 2 A–C). Rhamphomyia argyrotarsis Bezzi, 1909: 324 (holotype ♀). Melander, 1928: 186; Collin, 1933: 211 (cit). Macrostomus argyrotarsis: Smith, 1961: 53; 1967: 30 (cat.); Rafael & Cumming, 2004: 446. Diagnosis. Females of this species are slender, shiny, thorax and legs mainly yellow; postcranium with only a pair of occipital setae behind inner vertical seta; legs without pennate setae; hind tarsomere 1 whitish to silvery, elongate. Wing widely brown infuscate mainly in costal and apical margins, base narrowed and anal lobe very narrow. Tergites 1 and 2 reddish-brown, shiny. Redescription. Holotype female. Body length from original description: “ 4.5 mm ”; wing length 5.0 mm. Head longer than higher. Inferior facets slightly larger than superior ones. Frons shiny brown to black, slightly wider than anterior ocellus width. One pair of proclinate ocellar setae. Face parallel sided, slightly narrower than frons. Postcranium wide, shinning brown to black, entirely slightly gray pruinose. Inner vertical seta stouter than outer vertical seta, latter subequal in length to uniseriate postocular row of setae; only 1 pair of occipital setae behind inner vertical seta. Antenna brown with pedicel dark yellow. Proboscis yellow. Thorax shiny yellow to brown, darker medially on scutum and mediotergite; scutellum concolorous with scutum; mesopleuron yellow pruinose. Thoracic chaetotaxy: 2–3 antepronotals weak; 1 postpronotal stout, 1 weak; 4 dorsocentrals; 1 presutural supraalar; 0 postsutural supra-alar; 1 postalar; 2 small proepisternals; 1 notopleural; 2 pairs of scutellars, inner pair stouter, convergent; 4 laterotergitals with two anterior setae stouter. Legs shiny yellow, except apex of hind femur, entire hind tibia brown to black; hind tarsomere 1 whitish to silvery. Hind femur slightly clavate. Hind tarsomere 1 as long as remaining tarsomeres combined. Legs longer setae: hind tibia with 2–3 longer dorsal setae at distal fifth. Wing (Fig. 2A) widely brown infuscated mainly in costal and apical margins; pterostigma inconspicuous; veins M 1 and M 2 somewhat indistinct distally; vein CuA+CuP evanescent; anal lobe very narrow. Halter dark brown. Abdomen lost beyond tergite 3. Originally described as “tergites 1–2 and respective sternites yellow, remaining with blue/black reflection; postabdomen peculiar, black, with black setae, wider at base and with apical lamellae dark, narrow”. Male. Unknown. Geographical records. Bolivia (La Paz); Peru (Madre de Dios), new record. Type material examined. HOLOTYPE ♀: “ BOLIVIA [La Paz], San Carlos, Mapiri, i. 1903, 800 m” (SMTD). Additional material. PERU: Madre de Dios, Mazuko, 13°02′51.1″S– 70°20′45.9″W, 382 m, 18.viii.2012, sweep, J.A. Rafael, R.R. Cavichioli, D.M. Takiya (1♀, MUSM). Holotype condition. Distal half of postpedicel lost; right wing mounted on microslide; abdomen lost beyond tergite 3. Variation. One female from Peru (Fig. 2C) with mesonotum darker, brown; hind leg (Fig. 2B) with 2 longer posterodorsal and 1 anterodorsal setae at distal third and tarsomeres 1–4 whitish to silvery, tarsomere 5 brown; wing with last section of vein M 4 longer than in holotype (Figs 2C and 2A, respectively) and tergites 1–2 reddish brown, without blue reflection.Published as part of Rafael, José Albertino & Marques, Dayse Willkenia Almeida, 2019, Five new species of Macrostomus Wiedemann and a checklist of Empididae (s. str.) (Diptera) from Bolivia, pp. 251-275 in Zootaxa 4567 (2) on pages 253-254, DOI: 10.11646/zootaxa.4567.2.3, http://zenodo.org/record/259493