80 research outputs found

    Diversity dynamics in New Caledonia: towards the end of the museum model?

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    <p>Abstract</p> <p>Background</p> <p>The high diversity of New Caledonia has traditionally been seen as a result of its Gondwanan origin, old age and long isolation under stable climatic conditions (the museum model). Under this scenario, we would expect species diversification to follow a constant rate model. Alternatively, if New Caledonia was completely submerged after its breakup from Gondwana, as geological evidence indicates, we would expect species diversification to show a characteristic slowdown over time according to a diversity-dependent model where species accumulation decreases as space is filled.</p> <p>Results</p> <p>We reanalyze available datasets for New Caledonia and reconstruct the phylogenies using standardized methodologies; we use two ultrametrization alternatives; and we take into account phylogenetic uncertainty as well as incomplete taxon sampling when conducting diversification rate constancy tests. Our results indicate that for 8 of the 9 available phylogenies, there is significant evidence for a diversification slowdown. For the youngest group under investigation, the apparent lack of evidence of a significant slowdown could be because we are still observing the early phase of a logistic growth (i.e. the clade may be too young to exhibit a change in diversification rates).</p> <p>Conclusions</p> <p>Our results are consistent with a diversity-dependent model of diversification in New Caledonia. In opposition to the museum model, our results provide additional evidence that original New Caledonian biodiversity was wiped out during the episode of submersion, providing an open and empty space facilitating evolutionary radiations.</p

    Dyscolus danglesi Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus danglesi&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: 4D9BB587-2E50-4A2F-BD6A-8C693E0C7029&lt;/p&gt; &lt;p&gt;Figs 17&ndash;18&lt;/p&gt; Etymology &lt;p&gt;Noun in the genitive case, dedicated to Olivier Dangles, senior researcher in ecology, companion of Humboldtian adventures.&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt; (only known specimen)&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &male;; Loja Province, Parque Nacional Yacuri, Waypoint 170; 4.781387&ordm; S, 79.386938&ordm; W; 2870 m a.s.l.; 4 Aug. 2016; P. Moret and S. Aguirre leg.; COI voucher PM170-01, BOLD sequence SUM190-18; MNHN.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 17. Wingless. Body length: 12.8 mm. Body entirely black; femora and tibiae nigropiceous, tarsi and palpi reddish brown; antennomeres 1&ndash;4 nigropiceous with the base and the apex testaceous, antennomeres 5&ndash;11 reddish brown. Elytral microsculpture transverse. Head small, distinctly constricted basally; frons with two broad round depressions close to the anterior supraorbital seta; eyes convex and prominent, genae long, slightly convex; mandibles long, acutely ended. Pronotum cordiform, slightly broader than long; sides feebly arcuate apically, sinuate basally; hind angles rounded; two pairs of lateral setae. Elytra elongate-oval, base very narrow, humeri completely effaced; striae weakly impressed, subpunctate, intervals slightly convex near apex; subapical sinuation obsolete. Third elytral interval without discal setae. Last visible abdominal ventrite of the male with three pairs of setae along its apical margin. Legs slender, fourth metatarsomere with one pair of subapical dorsolateral setae, apical lobes asymmetrical with a big outer lobe and no inner lobe. &lt;i&gt;Male genitalia&lt;/i&gt;: Fig. 18. Median lobe arcuate in its basal half, then straight (in lateral view), apex short and blunt, endophallus with seven denticles in a small, slightly sclerotized subapical structure. &lt;i&gt;Female genitalia&lt;/i&gt;: unknown.&lt;/p&gt; Comparisons &lt;p&gt; This species shares with &lt;i&gt;D. caulatus&lt;/i&gt; Moret, 1993 the general form of the body and the absence of setae on the third interval of the elytra, but &lt;i&gt;D. caulatus&lt;/i&gt; lacks the anterior supraorbital seta and the posterior pronotal seta, and has a conspicuous foveate elytral pattern.&lt;/p&gt; Habitat &lt;p&gt;Upper montane forest on the Eastern slope of the Andes, at around 2870 m a.s.l. Collected by pyrethrin insecticide fogging on mossy branches and epiphytes, two meters above ground.&lt;/p&gt; Geographic distribution &lt;p&gt;Only known from the type locality in Southern Ecuador, in Parque Nacional Yacuri. Probably microendemic.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 22-23, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt

    Dyscolus verecundissimus Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus verecundissimus&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: CD4CC133-EF1C-4F71-8F38-0E954A5F94B4&lt;/p&gt; &lt;p&gt;Figs 55, 59&lt;/p&gt; &lt;p&gt; &lt;i&gt;Dyscolus verecundus&lt;/i&gt; Moret, 1998: 20, &lt;i&gt;pro parte.&lt;/i&gt;&lt;/p&gt; &lt;p&gt; &lt;i&gt;Dyscolus&lt;/i&gt; (&lt;i&gt;Dyscolus&lt;/i&gt;) &lt;i&gt;verecundus&lt;/i&gt; &ndash; Moret 2005: 142, &lt;i&gt;pro parte.&lt;/i&gt;&lt;/p&gt; Etymology &lt;p&gt; Superlative form of the Latin adjective &lsquo; &lt;i&gt;verecundus&lt;/i&gt; &rsquo;, which is the name of a related species.&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &male;; Chimborazo Province, Volc&aacute;n Chimborazo Est, moraine; 4470&ndash;4530 m a.s.l.; 4 Aug. 1998; P. Moret leg.; MNHN.&lt;/p&gt; &lt;p&gt; &lt;b&gt;Paratype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; 1 &female;; same collection data as for holotype; CPM.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 55. Very similar in almost every aspect to &lt;i&gt;D. verecundus&lt;/i&gt; and &lt;i&gt;D. verecundior&lt;/i&gt; Moret sp. nov., so the description will be limited to the few distinctive characters of this new species. Body length: 11.5&ndash;11.9 mm, as the biggest specimens of &lt;i&gt;D. verecundus&lt;/i&gt;; head slightly broader with convex eyes; antennae thinner, the second antennomere 1.76 to 1.77 times as long as wide (1.41 to 1.65 times in &lt;i&gt;D. verecundus&lt;/i&gt; and &lt;i&gt;D. verecundior&lt;/i&gt; Moret sp. nov.); pronotum broader, 1.27 to 1.28 times as wide as long (1.13 to 1.24 times in &lt;i&gt;D. verecundus&lt;/i&gt; and &lt;i&gt;D. verecundior&lt;/i&gt; Moret sp. nov.); apical blade of the median lobe broader and more rounded at apex in dorsal view (Fig. 59); endophallus with a large sclerotized area at middle, and a reduced, almost obsolete squamose area near apex (two sclerotized areas of similar size in &lt;i&gt;D. verecundus&lt;/i&gt; and &lt;i&gt;D. verecundior&lt;/i&gt; Moret sp. nov.).&lt;/p&gt; Habitat &lt;p&gt;Humid superp&aacute;ramo at around 4500 m a.s.l.&lt;/p&gt; Geographic distribution &lt;p&gt;Microendemic species, restricted to the Chimborazo volcano in the Western Cordillera.&lt;/p&gt; Comments &lt;p&gt; The isolated population we raise here to species rank was initially assigned to &lt;i&gt;D. verecundus&lt;/i&gt; by Moret (2005). There are more morphological differences between &lt;i&gt;D. verecundissimus&lt;/i&gt; Moret sp. nov. and the pair of sister species &lt;i&gt;D. verecundus&lt;/i&gt; and &lt;i&gt;D. verecundior&lt;/i&gt; Moret sp. nov. than between the two latter, especially in the male aedeagus.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 49-51, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt

    Dyscolus ruizi Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus ruizi&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: 811C080F-1087-49A8-BB75-7B71DA04C593&lt;/p&gt; &lt;p&gt;Figs 34&ndash;35&lt;/p&gt; Etymology &lt;p&gt;Noun in the genitive case, dedicated to Carlos Ruiz Carreira, associate professor at the Universidad de La Laguna (Canarias), who investigated in 2013&ndash;2014 the biodiversity of carabid beetles along an elevational gradient between Loja and Zamora.&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &male;; Loja Province, Parque Nacional Podocarpus, Cajanuma, Waypoint 397; 4&deg;7&prime;2.4&Prime; S, 79&deg;10&prime;6.3&Prime; W; 2900 m a.s.l.; 17 Mar. 2015; P. Moret and C. Ruiz leg.; COI voucher PM392-68, BOLD sequence SUM056-18; QCAZ.&lt;/p&gt; &lt;p&gt; &lt;b&gt;Paratypes&lt;/b&gt; (10 &male;&male;, 5 &female;&female;)&lt;/p&gt; &lt;p&gt; ECUADOR &ndash; &lt;b&gt;Loja Province&lt;/b&gt; &bull; 1 &male;; Cajanuma Sendero; 4.113&ordm; S, 79.174&ordm; W; 2920 m a.s.l.; 24 Sep. 2013; D. Mar&iacute;n and C. Ruiz leg.; CISEC &bull; 1 &male;; same collection data as for preceding; COI voucher CR120, BOLD sequence GBNP 068-17; QCAZ &bull; 1 &male;; El Tiro; 3.993&ordm; S, 79.147&ordm; W; 2838 m a.s.l.; 11 Jul. 2013; D. Mar&iacute;n and C. Ruiz leg.; COI voucher CR008, BOLD sequence GBNP 006-17; CISEC &bull; 1 &male;, 1 &female;; Saraguro, Huashapamba; 2920 m a.s.l.; 10 Dec. 2005; I. Mas leg.; CISEC 0003240, 0003271 &bull; 1 &male;; same collection data as for preceding; CPM. &ndash; &lt;b&gt;Loja / Zamora Province&lt;/b&gt; &bull; 1 &male;; El Tiro&ndash;Cima; 3.993&ordm; S, 79.147&ordm; W; 2838 m a.s.l.; 5 Sep. 2013; D. Mar&iacute;n and C. Ruiz leg.; CISEC &bull; 1 &male;, 1 &female;; same collection data as for preceding; CPM &bull; 1 &male;; LZ03 El Tiro&ndash;Cima; 3.993&ordm; S, 79.147&ordm; W; 2838 m a.s.l.; 11 Jul. 2013; D. Mar&iacute;n and C. Ruiz leg.; MNHN &bull; 1 &female;; LZ03 El Tiro; 2813 m a.s.l.; 26 Mar. 2008; Mar&iacute;n Rosero leg.; CISEC 0002573. &ndash; &lt;b&gt;Zamora-Chinchipe Province&lt;/b&gt; &bull; 1 &female;; Carretera Loja&ndash;Zamora; 3.988&ordm; S, 79.118&ordm; W; 2528 m a.s.l.; 18 Jul. 2013; D. Mar&iacute;n and C. Ruiz leg.; COI voucher CR002, BOLD sequence GBNP 002-17; CISEC &bull; 1 &female;; same collection data as for preceding; COI voucher CR003, BOLD sequence GBNP003-17; CISEC &bull; 1 &male;; same collection data as for preceding; 5 Dec. 2013; D. Mar&iacute;n and C. Ruiz leg.; COI voucher CR001, BOLD sequence GBNP001-17; CISEC &bull; 1 &male;; same collection data as for preceding; COI voucher CR055, BOLD sequence GBNP029-17; CISEC.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 34. Wingless. Body length: 9.2&ndash;11.3 mm. Entirely black or piceous black. Elytral microsculpture finely transverse, weakly impressed. Head broad basally, feebly constricted, eyes small, genae long and convex; antennae short. Pronotum elongate, wider at apex than at base; sides moderately arcuate apically, straight or feebly sinuate basally; hind angles obtuse and blunt; two pairs of lateral setae, the basal one anterad the hind angle. Elytra narrow depressed medially; no subapical sinuation; striae entire, well impressed; intervals flat to subconvex. Third elytral interval with two setae, one near base and one near apex, or with only one subapical seta. Last visible abdominal ventrite with one pair (&male;) or two pairs (&female;) of setae along its apical margin. Legs robust, anterior femora swollen; dorsal surface of tarsi rugose, almost squamose, not sulcate; fourth metatarsomere with one pair of dorsolateral subapical setae, apical lobes short, the external lobe two times longer than the inner lobe; fifth metatarsomere asetose ventrally. &lt;i&gt;Male genitalia&lt;/i&gt;: Fig. 35. Median lobe arcuate, apex robust, acute, endophallus without sclerotized structure. &lt;i&gt;Female genitalia&lt;/i&gt;: unstudied.&lt;/p&gt; Comparisons &lt;p&gt; &lt;i&gt;Dyscolus ruizi&lt;/i&gt; Moret sp. nov. belongs to the same clade as &lt;i&gt;D. marini&lt;/i&gt; Moret sp. nov., &lt;i&gt;D. arborarius&lt;/i&gt; Moret sp. nov., &lt;i&gt;D. moretianus&lt;/i&gt;, &lt;i&gt;D. sulcipedis&lt;/i&gt; Moret sp. nov. and &lt;i&gt;D. rugitarsis&lt;/i&gt; Moret sp. nov. However, by its robust legs, short antennae, small eyes and convex genae, &lt;i&gt;D. ruizi&lt;/i&gt; Moret sp. nov. is more similar to species living at higher elevations in the p&aacute;ramo, as &lt;i&gt;D. denigratus&lt;/i&gt; (Bates 1891), than to these species which share the same forest environment beneath 3000 m. &lt;i&gt;Dyscolus ruizi&lt;/i&gt; Moret sp. nov. differs from &lt;i&gt;D. denigratus&lt;/i&gt; by its elongate pronotum and the absence of the middle seta on the third interval.&lt;/p&gt; Habitat &lt;p&gt;Upper montane forest and subp&aacute;ramo on the eastern slope of the Andes, at around 2800&ndash;3000 m a.s.l. The specimens from Saraguro&ndash;Huashapamba were collected in pitfall traps baited with dung.&lt;/p&gt; Geographic distribution &lt;p&gt;Only known from the Parque Nacional Podocarpus and surroundings, in southern Ecuador. Probably microendemic.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 33-34, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt

    Dyscolus salazarae Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus salazarae&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: CD4CC653-FE0C-466A-A380-4C3361CB8579&lt;/p&gt; &lt;p&gt;Figs 32&ndash;33&lt;/p&gt; Etymology &lt;p&gt;Noun in the genitive case, dedicated to Fernanda Salazar, administrator of the invertebrate collection of the QCAZ Museum (Pontificia Universidad Cat&oacute;lica del Ecuador, Quito).&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt; (only known specimen)&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &male;; Loja Province, Parque Nacional Podocarpus, Cajanuma, Waypoint 392; 4&deg;6&prime;58.4&Prime; S, 79&deg;10&prime;18.6&Prime; W; 2850 m a.s.l.; 17 Mar. 2015; P. Moret and C. Ruiz leg.; COI voucher PM392-09, BOLD sequence SUM059-18; MNHN.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 32. Wingless. Body length: 9.3&ndash;9.6 mm. Head, pronotum and elytra brunneopiceous; legs, antennae and mouthparts reddish brown. Elytral microsculpture isodiametric. Head convex, markedly constricted basally, eyes small, not bulging, genae almost flat in dorsal view, as long as the eyes. Pronotum elongate, convex; sides weakly arcuate anterad, almost straight in basal fourth, with a very short sinuation at level of the basal seta, hind angles obtuse and blunt, anterior angles pronounced, broadly rounded; two pairs of lateral setae. Elytra elongate, subparallel; base relatively narrow, shoulders rounded; sides arcuate apically without subapical sinuation; striae entire, well impressed, not punctate; intervals slightly convex. Third elytral interval asetose. Legs rather short, meso- and metatarsomeres 1&ndash;3 with a lateral sulcus on each side; fourth metatarsomere with one pair of dorsolateral subapical setae, its apical lobes very short, the outer lobe twice as large as the inner lobe; fifth metatarsomere asetose ventrally. Last visible abdominal ventrite of the male with one pair of setae along its apical margin. &lt;i&gt;Male genitalia&lt;/i&gt;: Fig. 33. Median lobe almost straight in its basal half, shortly arcuate before apex; apex slightly reflexed upward, narrow and acute in lateral view; endophallus without sclerotized structure. &lt;i&gt;Female genitalia&lt;/i&gt;: unknown.&lt;/p&gt; Comparisons &lt;p&gt; Closely related to &lt;i&gt;D. moreti&lt;/i&gt; Perrault, 1993, but smaller (most specimens of &lt;i&gt;D. moreti&lt;/i&gt; range from 10 to 11 mm), the lateral margin of the pronotum narrower, the elytra narrower and less convex, the apex of the aedeagus shorter, more acute, and not bent ventrally as in &lt;i&gt;D. moreti&lt;/i&gt;.&lt;/p&gt; Habitat &lt;p&gt;Upper montane forest on the Eastern slope of the Andes, at around 2850 m a.s.l.&lt;/p&gt; Geographic distribution &lt;p&gt;Only known from the type locality in Southern Ecuador. Probably microendemic.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 34-36, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt

    Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes

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    International audienceThe genus Dyscolus Dejean, 1831 is a highly speciose taxon of neotropical Carabidae and the major component of high-altitude ground beetle communities in the tropical Andes. The aim of this study is threefold: (i) refine the taxonomic position of the equatorial members of Dyscolus using molecular data, (ii) provide a delimitation of the species found in Ecuador in páramo and montane forest environments based on a robust combination of molecular and morphological data, (iii) describe the new species and take the nomenclatural decisions made necessary by the results of this study. The seclusion of Dyscolus from more basal platynine clades including Platynus, Batenus and Glyptolenus, is supported by a phylogenetic analysis of the COI marker. Twenty-five new species of Dyscolus, most of them microendemic, are described and illustrated: D. aquator Moret sp. nov. (Tandayapa, Pichincha), D. arauzae Moret sp. nov. (Mt Cayambe, Pichincha), D. arborarius Moret sp. nov. (Parque Nacional Yacuri, Loja), D. barragani Moret sp. nov. (Mt Ayapungu, Chimborazo), D. crespoae Moret sp. nov. (Parque Nacional Podocarpus, Zamora-Chinchipe), D. danglesi Moret sp. nov. (Parque Nacional Yacuri, Loja), D. donosoi Moret sp. nov. (Parque Nacional Podocarpus, Zamora-Chinchipe), D. eleonorae Moret sp. nov. (Cotopaxi and Pichincha provinces), D. famelicus Moret sp. nov. (Papallacta, Napo), D. giselae Moret sp. nov. (Reserva Otonga, Cotopaxi), D. globoculus Moret sp. nov. (Parque Nacional Podocarpus, Zamora-Chinchipe), D. gobbii Moret sp. nov. (Guamaní and Mt Antisana, Pichincha), D. incommunis Moret sp. nov. (Tandayapa, Pichincha), D. marini Moret sp. nov. (Parque Nacional Podocarpus, Loja), D. piscator Moret sp. nov. (Guamaní, Napo), D. placitus Moret sp. nov. (Guamaní, Napo), D. ravidus Moret sp. nov. (Parque Nacional Podocarpus, Loja), D. rivinus Moret sp. nov. (Reserva Otonga, Cotopaxi), D. rugitarsis Moret sp. nov. (Parque Nacional Yacuri, Loja), D. ruizi Moret sp. nov. (Parque Nacional Podocarpus, Loja and Zamora), D. salazarae Moret sp. nov. (Parque Nacional Podocarpus, Loja), D. silvestris Moret sp. nov. (Papallacta, Napo), D. sulcipedis Moret sp. nov. (Parque Nacional Yacuri, Loja), D. verecundior Moret sp. nov. (Mt Illiniza and Mt Corazón, Pichincha) and D. verecundissimus Moret sp. nov. (Mt Chimborazo, Chimborazo). Dyscolus palatus Moret, 1998 is newly synonymized with D. denigratus (Bates, 1891). We demonstrate the subgenus Hydrodyscolus Moret, 1996 to be polyphyletic and therefore consider it a junior synonym of Dyscolus Dejean, 1831

    Origine de la biodiversité en Nouvelle-Calédonie (analyse phylogénétique de l'endémisme chez les insectes dictyoptères)

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    Dans le but d analyser l origine de l endémisme en Nouvelle-Calédonie, l histoire évolutive des Insectes Blattidae (Hexapodes, Dictyoptères) a été reconstruite au niveau moléculaire par l application de différentes méthodes phylogénétiques. Les différents résultats obtenus, aux niveaux régional et local, permettent de modifier fortement les scénarios de l origine de la biodiversité en Nouvelle-Calédonie. Loin d être une relique purement Gondwanienne et une région dont la diversification a été progressive et ancienne ( Modèle Muséum ), la Nouvelle-Calédonie montre les caractéristiques nuancées de toutes les faunes tropicales, avec plusieurs strates de peuplement et une combinaison d événements anciens et récents. Ce scénario se révèle être aussi plus cohérent avec l histoire géologique mouvementée de la Nouvelle-Calédonie, notamment la submersion totale du territoire.PARIS-BIUSJ-Thèses (751052125) / SudocPARIS-BIUSJ-Physique recherche (751052113) / SudocSudocFranceF

    Dyscolus crespoae Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus crespoae&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: AEA12495-C2BD-4A65-AACF-C54DB6CAFBA7&lt;/p&gt; &lt;p&gt;Fig. 29&lt;/p&gt; Etymology &lt;p&gt;Noun in the genitive case, dedicated to Ver&oacute;nica Crespo, professor at the Pontificia Universidad Cat&oacute;lica del Ecuador, limnologist and specialist of the ecology of Andean tropical streams.&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &female;; Zamora-Chinchipe Province, Parque Nacional Podocarpus, Estaci&oacute;n El Colibr&iacute;; 3&deg;59&prime;16.1&Prime; S, 79&deg;5&prime;39.0&Prime; W; 2110 m a.s.l.; 18 Mar. 2015; P. Moret and C. Ruiz leg.; by night 19.30&ndash; 21 pm; on the ground; QCAZ.&lt;/p&gt; &lt;p&gt; &lt;b&gt;Paratype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; 1 &female;; same collection data as for holotype; COI voucher PM402-03, BOLD sequence SUM188-18; CPM.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 29. Wingless. Body length: 10.6 mm (paratype) to 11.3 mm (holotype). Head and pronotum black, elytra nigropiceous; legs, antennae and mouthparts brownish to reddish brown. Dorsal integuments shiny, elytral microsculpture transverse. Head relatively small, elongate, eyes prominent, genae long, almost flat. Pronotum cordiform; sides markedly sinuate basally, hind angles obtuse and sharp; laterobasal impressions deep, lateral margins reflexed; two pairs of lateral setae. Elytra oval, convex; striae entire, shallowly impressed, distinctly punctate; intervals flat; preapical sinuation weakly marked. Third elytral interval without discal setae. Last visible abdominal ventrite with two pairs of setae along its apical margin. Legs moderately slender, all tarsi densely pubescent ventrally; metatarsomeres 1&ndash;3 bisulcate; external lobe of the fourth metatarsomere 2.5 times longer than inner lobe; fifth metatarsomere asetose ventrally. Male unknown. &lt;i&gt;Female genitalia&lt;/i&gt;: unstudied.&lt;/p&gt; Comparisons &lt;p&gt; This species belongs to the same clade as &lt;i&gt;D. ravidus&lt;/i&gt; Moret sp. nov. Both have the same general aspect and share the character of an asetose third interval, but in &lt;i&gt;D. crespoae&lt;/i&gt; Moret sp. nov. the dorsal surface is shiny with a transverse microsculpture on the elytra, the elytra are more convex, the hind angles of&lt;/p&gt; &lt;p&gt;the pronotum more obtuse, the ventral pubescence of the tarsi denser, the apical lobes of the fourth metatarsomere more asymmetrical.&lt;/p&gt; Habitat &lt;p&gt;Montane forest on the Eastern slope of the Andes, at around 2100 m a.s.l. Active at the beginning of the night (7.30&ndash;9 pm) on the surface of the leaf litter above ground.&lt;/p&gt; Geographic distribution &lt;p&gt;Only known from the type locality in Southern Ecuador, in the Parque Nacional Podocarpus. Probably microendemic.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 30-32, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt

    Dyscolus gobbii Moret 2020, sp. nov.

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    &lt;i&gt;Dyscolus gobbii&lt;/i&gt; Moret sp. nov. &lt;p&gt;urn:lsid:zoobank.org:act: 787DC50E-8874-4E7F-BCAD-FE33230508A1&lt;/p&gt; &lt;p&gt;Fig. 42&lt;/p&gt; Etymology &lt;p&gt;Noun in the genitive case, dedicated to Mauro Gobbi, specialist of the ecology of glacier-foreland invertebrates, who took part in the discovery of this species.&lt;/p&gt; Type material &lt;p&gt; &lt;b&gt;Holotype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; &female;; Pichincha Province, Guaman&iacute;, Paso de la Virgen, Waypoint 210; 0&deg;19&prime;18.3&Prime; S, 78&deg;11&prime;54.8&Prime; W; 4230 m a.s.l.; 27 Feb. 2017; P. Moret and M. Gobbi leg.; COI voucher PM210-16, BOLD sequence SUM228-18; QCAZ.&lt;/p&gt; &lt;p&gt; &lt;b&gt;Paratype&lt;/b&gt;&lt;/p&gt; &lt;p&gt;ECUADOR &bull; 1 &female;; Pichincha Province, Volc&aacute;n Antisana NW, B1C3; 0.457780&ordm; S, 78.167970&ordm; W; 4401 m a.s.l.; 22 Apr. 2016; E. Moreno leg.; COI voucher in ethanol PM188-01, BOLD sequence SUM183-18; CPM.&lt;/p&gt; Diagnostic description &lt;p&gt; Habitus: Fig. 42. Wingless. Body length: 10.2 mm (holotype) to 12.3 mm (paratype). Body piceous black, legs brownish with reddish-brown femora. Elytral microsculpture made of oblong sculpticells, shallowly impressed. Head big, convex dorsally; mandibles long and sharp, as long as the length of the head from base to apex of the labium; eyes moderately bulging, genae oblique, slightly convex in dorsal view. Pronotum transverse, subquadrate, narrower at base than at apex; lateroapical lobes broadly rounded; hind angles obtusely rounded; two pairs of lateral setae. Elytra oval-shaped, convex, with effaced humeri. Striae weakly impressed, with traces of punctuation; intervals 1&ndash;4 slightly convex, 5&ndash;8 flat; third interval with five or six discal setae. Last visible abdominal ventrite of the female with two pairs of setae along its apical margin. Legs slender but short; fourth metatarsomere with convex sides, without subapical dorsolateral setae (holotype) or with weak and short subapical dorsolateral setae (paratype), apical lobes ovoid, the outer lobe slightly longer than the inner lobe. Male genitalia: unknown. &lt;i&gt;Female genitalia&lt;/i&gt;: unstudied.&lt;/p&gt; Comparisons &lt;p&gt; This isolated species remotely resembles &lt;i&gt;Dyscolus segnipes&lt;/i&gt; Moret, 1990, with which it coexists in the Guaman&iacute; p&aacute;ramo. &lt;i&gt;Dyscolus segnipes&lt;/i&gt; has much shorter mandibles and well developed subapical dorsolateral setae on the fourth metatarsomere. The COI tree places &lt;i&gt;gobbii&lt;/i&gt; and &lt;i&gt;segnipes&lt;/i&gt; in two clades far apart from each other.&lt;/p&gt; Habitat &lt;p&gt;Humid superp&aacute;ramo, from 4200 to 4400 m a.s.l. The Guaman&iacute; specimen was collected under a stone on the shore of a small lake; the Antisana specimen was collected by pitfall trapping in a mesic environment, 20 metres from a glacier-fed stream.&lt;/p&gt; Geographic distribution &lt;p&gt;Microendemic species, restricted to the Antisana and Guaman&iacute; areas in the Eastern Cordillera.&lt;/p&gt;Published as part of &lt;i&gt;Moret, Pierre &amp; Murienne, JĂ©rĂ´me, 2020, Integrative taxonomy of the genus Dyscolus (Coleoptera, Carabidae, Platynini) in Ecuadorian Andes, pp. 1-55 in European Journal of Taxonomy 646&lt;/i&gt; on pages 40-42, DOI: 10.5852/ejt.2020.646, &lt;a href="http://zenodo.org/record/3829682"&gt;http://zenodo.org/record/3829682&lt;/a&gt
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