Adaptation and enslavement in endosymbiont-host associations

The evolutionary persistence of symbiotic associations is a puzzle. Adaptation should eliminate cooperative traits if it is possible to enjoy the advantages of cooperation without reciprocating - a facet of cooperation known in game theory as the Prisoner's Dilemma. Despite this barrier, symbioses are widespread, and may have been necessary for the evolution of complex life. The discovery of strategies such as tit-for-tat has been presented as a general solution to the problem of cooperation. However, this only holds for within-species cooperation, where a single strategy will come to dominate the population. In a symbiotic association each species may have a different strategy, and the theoretical analysis of the single species problem is no guide to the outcome. We present basic analysis of two-species cooperation and show that a species with a fast adaptation rate is enslaved by a slowly evolving one. Paradoxically, the rapidly evolving species becomes highly cooperative, whereas the slowly evolving one gives little in return. This helps understand the occurrence of endosymbioses where the host benefits, but the symbionts appear to gain little from the association.Comment: v2: Correction made to equations 5 & 6 v3: Revised version accepted in Phys. Rev. E; New figure adde

Fast outflows in compact radio sources: evidence for AGN-induced feedback in the early stages of radio source evolution

We present intermediate resolution, wide wavelength coverage spectra for a complete sample of 14 compact radio sources taken with the aim of investigating the impact of the nuclear activity on the circumnuclear (ISM) in the early stages of radio source evolution. We observe spatially extended line emission (up to 20 kpc) in the majority of sources which is consistent with a quiescent halo. In the nuclear apertures we observe broad, highly complex emission line profiles. Multiple Gaussian modelling of the [O III]5007 line reveals 2-4 components which can have FWHM and blueshifts relative to the halo of up to 2000 km/s. We interpret these broad, blueshifted components as material in outflow and discuss the kinematical evidence for jet-driven outflows. Comparisons with samples in the literature show that compact radio sources harbour more extreme nuclear kinematics than their extended counterparts, a trend seen within our sample with larger velocities in the smaller sources. The observed velocities are also likely to be influenced by source orientation with respect to the observer's line of sight. Nine sources have associated HI absorption. In common with the optical emission line gas, the HI profiles are often highly complex with the majority of the detected components significantly blueshifted, tracing outflows in the neutral gas. The sample has been tested for stratification in the ISM (FWHM/ionisation potential/critical density) as suggested by Holt et al. (2003) for PKS1345+12 but we find no significant trends within the sample using a Spearman Rank analysis. This study supports the idea that compact radio sources are young radio loud AGN observed during the early stages of their evolution and currently shedding their natal cocoons through extreme circumnuclear outflows.Comment: Accepted for publication in MNRAS, 24 pages, 7 figure

New X-ray Constraints on Starburst and Seyfert Activity in the Barred Spiral Galaxy NGC 1672

The nearby barred spiral galaxy NGC 1672 shows dramatic starburst activity and may also host a Seyfert 2 nucleus. We present new X-ray observations that set constraints on starburst and Seyfert activity in NGC 1672. Two ROSAT HRI exposures, taken in 1992 and 1997, are used to investigate long-term variability of the known X-ray sources and to search for new sources of X-ray emission. We find large-amplitude (about 69%) variability from X-3, one of the off-nuclear sources located near an end of the galactic bar. X-3 has a peak observed 0.2-2.0 keV luminosity of about 2.5 x 10^{39} erg/s, and it is probably a luminous X-ray binary or young supernova remnant. We do not observe variability of the nuclear source X-1 or the strong off-nuclear source X-2. Our analyses also reveal two new off-nuclear sources, one of which is associated with a bright region along a spiral arm, and we find evidence for large-scale diffuse X-ray emission throughout part of the disk of NGC 1672. Furthermore, we use ASCA data taken in 1995 to constrain the hard X-ray properties of NGC 1672. While the nuclear source X-1 is the dominant soft X-ray source in NGC 1672, we find that the bulk of the 2-10 keV and 5-10 keV emission is spatially coincident with the off-nuclear source X-3, giving it an apparent 0.2-8 keV luminosity of about 6 x 10^{39} erg/s. A power-law plus Raymond-Smith model provides an acceptable fit to the full-band ASCA spectra. We do not find any evidence for a luminous but absorbed nuclear X-ray source. If there is a luminous Seyfert 2 nucleus in NGC 1672, it must be obscured by a `Compton-thick' torus with a column density of > 2 x 10^{24} cm^{-2}.Comment: 19 pages, AJ, accepted, also available from http://www.astro.psu.edu/users/niel/papers/papers.htm

Fecundity and the demographic strategies of coral morphologies

JM, AB and SC were supported by fellowships from the Australian Research Council (FT110100609, FT0990652 and DP0880544 respectively). MD was supported by the ERC (BioTIME 250189) and the Scottish Funding Council (MASTS - HR09011).Understanding species differences in demographic strategies is a fundamental goal of ecology. In scleractinian corals, colony morphology is tightly linked with many demographic traits, such as size-specific growth and morality. Here, we test how well morphology predicts the colony size-fecundity relationship in eight species of broadcast-spawning corals. Variation in colony fecundity is greater among morphologies than between species with a similar morphology, demonstrating that colony morphology can be used as a quantitative proxy for demographic strategies. Additionally, we examine the relationship between size-specific colony fecundity and mechanical vulnerability (i.e. vulnerability to colony dislodgment). Interestingly, the relationship between size-specific fecundity and mechanical vulnerability varied among morphologies. For tabular species, the most fecund colonies are the most mechanically vulnerable, while the opposite is true for massive species. For corymbose and digitate colonies, mechanical vulnerability remains relatively constant as fecundity increases. These results reveal strong differences in the demographic trade-offs among species of different morphologies. Using colony morphology as a quantitative proxy for demographic strategies can help predict coral community dynamics and responses to anthropogenic change.Publisher PDFPeer reviewe

The host galaxies of three radio-loud quasars: 3C 48, 3C 345, and B2 1425+267

Observations with the Wide-Field/Planetary Camera-2 of the Hubble Space Telescope (HST) are presented for three radio-loud quasars: 3C 48 (z=0.367), B2 1425+267 (z=0.366), and 3C 345 (z=0.594). All three quasars have luminous (~4 L^*) galaxies as hosts, which are either elliptical (B2 1425+267 and 3C 345) or interacting (3C 48), and all hosts are 0.5 - 1.0 mag bluer in (V-I) than other galaxies with the same overall morphology at similar redshifts to the quasars. The host of 3C 48 has many H II regions and a very extended tidal tail. All nine of the radio-loud quasars studied here and in Bahcall et al. (1997) either have bright elliptical hosts or occur in interacting systems. There is a robust correlation between the radio emission of the quasar and the luminosity of host galaxy; the radio-loud quasars reside in galaxies that are on average about 1 mag brighter than hosts of the radio-quiet quasars.Comment: Accepted for publication in ApJ. 3 postscript and 3 jpeg figures. Original figures may be found in ftp://eku.sns.ias.edu/pub/sofia/RadioLoud

The Nature of LINERs

We present $J$-band ($1.15-1.35 \mu$m) spectroscopy of a sample of nine galaxies showing some degree of LINER activity (classical LINERs, weak-[O {\sc i}] LINERs and transition objects), together with $H$-band spectroscopy for some of them. A careful subtraction of the stellar continuum allows us to obtain reliable [Fe {\sc ii}]$1.2567 \mu$m/Pa$\beta$ line ratios. We conclude that different types of LINERs (i.e., photoionized by a stellar continuum or by an AGN) cannot be easily distinguished based solely on the [Fe {\sc ii}]$1.2567 \mu$m/Pa$\beta$ line ratio. The emission line properties of many LINERs can be explained in terms of an aging starburst. The optical line ratios of these LINERs are reproduced by a model with a metal-rich H {\sc ii} region component photoionized with a single stellar temperature $T_* = 38,000$K, plus a supernova remnant (SNR) component. The [Fe {\sc ii}] line is predominantly excited by shocks produced by SNRs in starbursts and starburst-dominated LINERs, while Pa$\beta$ tracks H {\sc ii} regions ionized by massive young stars. The contribution from SNRs to the overall emission line spectrum is constrained by the [Fe {\sc ii}]$1.2567 \mu$m/Pa$\beta$ line ratio. Although our models for aging starbursts are constrained only by these infrared lines, they consistently explain the optical spectra of the galaxies also. The LINER-starburst connection is tested by predicting the time dependence of the ratio of the ionizing luminosity ($L_{\rm ion}$) to the supernova rate (SNr), $L_{\rm ion}$/(SNr). We predict the relative number of starbursts to starburst-dominated LINERs (aging starbursts) and show that it is in approximate agreement with survey findings for nearby galaxies.Comment: Accepted in ApJ (19 pages, 8 figures, uses emulateapj.sty

Spitzer Space Telescope Infrared Imaging and Spectroscopy of the Crab Nebula

We present 3.6, 4.5, 5.8, 8.0, 24, and 70 micron images of the Crab Nebula obtained with the Spitzer Space Telescope IRAC and MIPS cameras, Low- and High-resolution Spitzer IRS spectra of selected positions within the nebula, and a near-infrared ground-based image made in the light of [Fe II]1.644 micron. The 8.0 micron image, made with a bandpass that includes [Ar II]7.0 micron, resembles the general morphology of visible H-alpha and near-IR [Fe II] line emission, while the 3.6 and 4.5 micron images are dominated by continuum synchrotron emission. The 24 micron and 70 micron images show enhanced emission that may be due to line emission or the presence of a small amount of warm dust in the nebula on the order of less than 1% of a solar mass. The ratio of the 3.6 and 4.5 micron images reveals a spatial variation in the synchrotron power law index ranging from approximately 0.3 to 0.8 across the nebula. Combining this information with optical and X-ray synchrotron images, we derive a broadband spectrum that reflects the superposition of the flatter spectrum jet and torus with the steeper diffuse nebula, and suggestions of the expected pileup of relativistic electrons just before the exponential cutoff in the X-ray. The pulsar, and the associated equatorial toroid and polar jet structures seen in Chandra and HST images (Hester et al. 2002) can be identified in all of the IRAC images. We present the IR photometry of the pulsar. The forbidden lines identified in the high resolution IR spectra are all double due to Doppler shifts from the front and back of the expanding nebula and give an expansion velocity of approximately 1264 km/s.Comment: 21 pages, 4 tables, 16 figure

Elusive Active Galactic Nuclei

A fraction of active galactic nuclei do not show the classical Seyfert-type signatures in their optical spectra, i.e. they are optically "elusive". X-ray observations are an optimal tool to identify this class of objects. We combine new Chandra observations with archival X-ray data in order to obtain a first estimate of the fraction of elusive AGN in local galaxies and to constrain their nature. Our results suggest that elusive AGN have a local density comparable to or even higher than optically classified Seyfert nuclei. Most elusive AGN are heavily absorbed in the X-rays, with gas column densities exceeding 10^24 cm^-2, suggesting that their peculiar nature is associated with obscuration. It is likely that in elusive AGN, the nuclear UV source is completely embedded and the ionizing photons cannot escape, which prevents the formation of a classical Narrow Line Region. Elusive AGN may contribute significantly to the 30 keV bump of the X-ray background.Comment: accepted for publication in MNRAS Letters, 6 pages, 3 figures, typos and references correcte

Supercooled liquid water cloud observed, analysed, and modelled at the top of the planetary boundary layer above Dome C, Antarctica

Abstract. A comprehensive analysis of the water budget over the Dome C (Concordia, Antarctica) station has been performed during the austral summer 2018–2019 as part of the Year of Polar Prediction (YOPP) international campaign. Thin (∼100 m deep) supercooled liquid water (SLW) clouds have been detected and analysed using remotely sensed observations at the station (tropospheric depolarization lidar, the H2O Antarctica Microwave Stratospheric and Tropospheric Radiometer (HAMSTRAD), net surface radiation from the Baseline Surface Radiation Network (BSRN)), radiosondes, and satellite observations (CALIOP, Cloud-Aerosol LIdar with Orthogonal Polarization/CALIPSO, Cloud Aerosol Lidar and Infrared Pathfinder Satellite Observations) combined with a specific configuration of the numerical weather prediction model: ARPEGE-SH (Action de Recherche Petite Echelle Grande Echelle – Southern Hemisphere). The analysis shows that SLW clouds were present from November to March, with the greatest frequency occurring in December and January when ∼50 % of the days in summer time exhibited SLW clouds for at least 1 h. Two case studies are used to illustrate this phenomenon. On 24 December 2018, the atmospheric planetary boundary layer (PBL) evolved following a typical diurnal variation, which is to say with a warm and dry mixing layer at local noon thicker than the cold and dry stable layer at local midnight. Our study showed that the SLW clouds were observed at Dome C within the entrainment and the capping inversion zones at the top of the PBL. ARPEGE-SH was not able to correctly estimate the ratio between liquid and solid water inside the clouds with the liquid water path (LWP) strongly underestimated by a factor of 1000 compared to observations. The lack of simulated SLW in the model impacted the net surface radiation that was 20–30 W m−2 higher in the BSRN observations than in the ARPEGE-SH calculations, mainly attributable to the BSRN longwave downward surface radiation being 50 W m−2 greater than that of ARPEGE-SH. The second case study took place on 20 December 2018, when a warm and wet episode impacted the PBL with no clear diurnal cycle of the PBL top. SLW cloud appearance within the entrainment and capping inversion zones coincided with the warm and wet event. The amount of liquid water measured by HAMSTRAD was ∼20 times greater in this perturbed PBL than in the typical PBL. Since ARPEGE-SH was not able to accurately reproduce these SLW clouds, the discrepancy between the observed and calculated net surface radiation was even greater than in the typical PBL case, reaching +50 W m−2, mainly attributable to the downwelling longwave surface radiation from BSRN being 100 W m−2 greater than that of ARPEGE-SH. The model was then run with a new partition function favouring liquid water for temperatures below −20 down to −40 ∘C. In this test mode, ARPEGE-SH has been able to generate SLW clouds with modelled LWP and net surface radiation consistent with observations during the typical case, whereas, during the perturbed case, the modelled LWP was 10 times less than the observations and the modelled net surface radiation remained lower than the observations by ∼50 W m−2. Accurately modelling the presence of SLW clouds appears crucial to correctly simulate the surface energy budget over the Antarctic Plateau

A comparative HST imaging study of the host galaxies of radio-quiet quasars, radio-loud quasars and radio galaxies: Paper I

We present the first results from a major HST WFPC2 imaging study aimed at providing the first statistically meaningful comparison of the morphologies, luminosities, scalelengths and colours of the host galaxies of radio-quiet quasars, radio-loud quasars, and radio galaxies. We describe the design of this study and present the images which have been obtained for the first half of our 33-source sample. We find that the hosts of all three classes of luminous AGN are massive elliptical galaxies, with scalelengths ~=10 kpc, and R-K colours consistent with mature stellar populations. Most importantly this is the the first unambiguous evidence that, just like radio-loud quasars, essentially all radio-quiet quasars brighter than M_R = -24 reside in massive ellipticals. This result removes the possibility that radio `loudness' is directly linked to host galaxy morphology, but is however in excellent accord with the black-hole/spheroid mass correlation recently highlighted by Magorrian et al. (1998). We apply the relations given by Magorrian et al. to infer the expected Eddington luminosity of the putative black hole at the centre of each of the spheroidal host galaxies we have uncovered. Comparison with the actual nuclear R-band luminosities suggests that the black holes in most of these galaxies are radiating at a few percent of the Eddington luminosity; the brightest host galaxies in our low-z sample are capable of hosting quasars with M_R = -28, comparable to the most luminous quasars at z = 3. Finally we discuss our host-derived black-hole masses in the context of the radio-luminosity:black-hole mass correlation recently uncovered for nearby galaxies by Franceschini et al. (1998), and the resulting implications for the physical origin of radio loudness.Comment: Submitted for publication in the Astrophysical Journal, 55 pages of latex, plus 12 postscript figures (Figures 1a-1s (greyscales of images and model fits, and Figures 2a-2g (luminosity profiles and model fits) can be downloaded from http://www.roe.ac.uk/astronomy/html/rjm1.shtml