52 research outputs found

    The structure of feeding behavior in a phytophagous insect (Hylobius abietis)

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    Analysis of the feeding behavior of animals using such a high temporal resolution that meals can be defined may improve our understanding of the mechanisms regulating feeding. Meals can be distinguished in an ethologically meaningful manner by using the ‘meal criterion’, the shortest non-feeding interval between feeding bouts recognized as meals. However, such a criterion has only been determined for a few insect species. Applying a recent method developed for assessing meal criteria for vertebrates, we determined the meal criterion for Hylobius abietis (L.) (Coleoptera: Curculionidae) based on data from video recordings of single individuals feeding on seedlings of Norway spruce, Picea abies (L.) Karst. (Pinaceae). The pine weevil is an economically important pest insect because it feeds on the stem bark of planted conifer seedlings. Weevils had 4-5 meals per day. Each meal lasted about 24 min during which about 13 mm2 of bark per meal were removed. Females had longer total meal durations and longer non-feeding intervals within meals than males. Girdling seedlings did not affect the weevils’ feeding properties. The size of meals was significantly correlated to the duration of non-feeding intervals before and after them. This study is one of few describing the feeding behavior of an insect at a temporal resolution that allows individual meals to be distinguished. With more meal-related data from insects available, differences in meal properties may be interpreted based on phylogeny, ecology, and physiology. Our results may also assist in the setup and interpretation of studies of plant-insect interactions, and facilitate the evaluation and development of methods to protect plants against herbivores

    Generation of Frequency Tunable and Low Phase Noise Micro- and Millimeter-Wave Signals using Photonic Technologies

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    The concept of generating micro- and millimeter-wave signals by optical means offers a variety of unique features compared to purely electronics such as high frequency tunability, ultra-wideband operation and the possibility to distribute micro- and millimeter-wave signals over kilometers of optical fiber to a remote site. These features make the photonic synthesizer concept a very interesting alternative for several applications in the micro- and millimeter-wave regime. This thesis focuses on the realization and characterization of different photonic synthesizer concepts for the optical generation of frequency tunable and low phase noise micro- and millimeter-wave signals. Advanced microwave photonic approaches utilizing external optical modulation and optical multiplication will be presented, offering high frequency optical millimeter-wave generation up to 110 GHz with superior performances in terms of maximum frequency tuning ranges and phase noise characteristics. In addition, the concept of a novel dual-loop optoelectronic oscillator will be presented that enables optical millimeter-wave signal generation without the need of any electronic reference oscillator. By using the developed dual-loop optoelectronic oscillator, microwave signal generation with tuning ranges in the gigahertz regime has been experimentally demonstrated for the first time.Das Konzept der optischen Mikro- und Millimeterwellen-Generation bietet gegenüber rein elektronischen Konzepten eine Vielzahl einzigartiger Möglichkeiten, bedingt durch die hohe Frequenzabstimmbarkeit, die extrem hohe Bandbreite sowie die Möglichkeit, Mikro- und Millimeterwellen-Signale über optische Fasern kilometerweit zu einer entfernten Station zu übertragen. Diese Eigenschaften machen das Konzept des photonischen Synthesizers zu einer sehr interessanten Alternative für viele Applikationen im Mikro- und Millimeterwellen-Bereich. Diese Arbeit beschäftigt sich mit der Realisierung und Charakterisierung verschiedener photonischer Synthesizer-Konzepte zur optischen Generation von frequenzabstimmbaren Mikro- und Millimeterwellen-Signalen mit geringem Phasenrauschen. Fortschrittliche photonische Konzepte unter Ausnutzung externer optischer Modulation sowie optischer Multiplikation werden vorgestellt. Diese Konzepte ermöglichen die optische Generierung hochfrequenter Millimeterwellen bis zu 110 GHz mit ausgezeichneter Performance in Bezug auf maximale Frequenzabstimmbarkeit sowie Phasenrauschen. Des Weiteren wurde ein neuartiges Konzept des optoelektronischen Oszillators, bestehend aus zwei Faserringen, vorgestellt, welches die Generierung von Millimeterwellen-Signalen ohne die Notwendigkeit eines elektronischen Referenzoszillators ermöglicht. Mit Hilfe des entwickelten optoelektronischen Oszillators wurde erstmals ein Mikrowellen-Signal mit einer Frequenzabstimmbarkeit im Gigahertz-Bereich experimentell erreicht

    Diel behaviour and time budget of the adult pine weevil, Hylobius abietis

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    The pine weevil (Hylobius abietis (L.); Coleoptera: Curculionidae) has a high economic impact on forest regeneration in Europe. The general biology of the pine weevil has received considerable attention, but there is insufficient knowledge about its diel behaviour and time budget. Therefore, the feeding and locomotion behaviour of individual adult weevils on Norway spruce (Picea abies (L.) Karst.) seedlings is observed for 24 hour-periods in the laboratory. Both girdled and non-girdled seedlings are used to assess how the weevils’ behaviour is influenced by the plants physiological response to the girdling. The locomotion pattern shows a distinct maximum during the beginning of the dark phase whereas most feeding occurs during the second half of the dark phase and the first hours of the subsequent light phase. The girdling treatment increase the time that weevils spend on the seedlings during the first part of an observation session, but has no effect on their feeding pattern. The weevils’ time budgets on girdled and non-girdled seedlings are similar. On average, they spend 34 % of their time in locomotion and 6 % on feeding. Females spend more time feeding than males (7.1 % vs. 4.2 %), possibly because they have higher food requirements, e.g. for egg production. Females also spend more time in total on the seedlings than males (26.3 % vs. 7.0 %). This study reveals, in high temporal resolution, the diel feeding and locomotion behaviour and time budget of male and female pine weevils

    Seasonal timing and recurrence of methyl jasmonate treatment influence pine weevil damage to Norway spruce seedlings

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    Defense can be induced in conifer seedlings to reduce pine weevil (Hylobius abietis) damage, by treatment with the plant hormone methyl jasmonate (MJ). Few studies have addressed important practical issues regarding the use of MJ such as treatment incidence and timing, seedling age and its compatibility with plant nursery practices. We examined if levels of pine weevil damage depend on seasonal timing and recurrence of MJ treatment, and if the observed effects depend on plant age. Norway spruce (Picea abies) seedlings from two age cohorts (1 year and 1.5 years old) received four MJ treatments: MJ application before winter storage in the previous year, after winter storage but before spring/summer planting, repeated MJ application (both before winter storage, and before planting) or no MJ application at all. Pine weevil damage was evaluated in a lab and field experiment. We found that the timing and recurrence of MJ treatment affected the amount of damage inflicted by pine weevils in different ways, but these effects were consistent among age cohorts. MJ application before winter storage provided the most effective protection, and this reduction in damage was comparable to that provided by a currently used physical protection method against pine weevil feeding. Our results indicated that MJ can be applied in line with nursery practices (before winter storage) and provides adequate protection for two growing seasons

    Bottom-Up Growth of Monolayer Honeycomb SiC

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    The long theorized two-dimensional allotrope of SiC has remained elusive amid the exploration of graphenelike honeycomb structured monolayers. It is anticipated to possess a large direct band gap (2.5 eV), ambient stability, and chemical versatility. While sp2 bonding between silicon and carbon is energetically favorable, only disordered nanoflakes have been reported to date. Here we demonstrate large-area, bottom-up synthesis of monocrystalline, epitaxial monolayer honeycomb SiC atop ultrathin transition metal carbide films on SiC substrates. We find the 2D phase of SiC to be almost planar and stable at high temperatures, up to 1200 \ub0C in vacuum. Interactions between the 2D-SiC and the transition metal carbide surface result in a Dirac-like feature in the electronic band structure, which in the case of a TaC substrate is strongly spin-split. Our findings represent the first step towards routine and tailored synthesis of 2D-SiC monolayers, and this novel heteroepitaxial system may find diverse applications ranging from photovoltaics to topological superconductivity

    Magnetic topological insulator MnBi6Te10 with zero-field ferromagnetic state and gapped Dirac surface states

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    Magnetic topological insulators (TIs) with nontrivial topological electronic structure and broken time-reversal symmetry exhibit various exotic topological quantum phenomena. The realization of such exotic phenomena at high temperature is one of central topics in this area. We reveal that MnBi6Te10 is a magnetic TI with an antiferromagnetic ground state below 10.8 K whose nontrivial topology is manifested by Dirac-like surface states. The ferromagnetic axion insulator state with Z4 = 2 emerges once spins polarized at field as low as 0.1 T, accompanied with saturated anomalous Hall resistivity up to 10 K. Such a ferromagnetic state is preserved even external field down to zero at 2 K. Theoretical calculations indicate that the few-layer ferromagnetic MnBi6Te10 is also topologically nontrivial with a non-zero Chern number. Angle-resolved photoemission spectroscopy experiments further reveal three types of Dirac surface states arising from different terminations on the cleavage surfaces, one of which has insulating behavior with an energy gap of ~ 28 meV at the Dirac point. These outstanding features suggest that MnBi6Te10 is a promising system to realize various topological quantum effects at zero field and high temperature.Comment: 18 pages, 4 figures and 1 tabl

    Origin of the π -band replicas in the electronic structure of graphene grown on 4H -SiC(0001)

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    The calculated electronic band structure of graphene is relatively simple, with a Fermi surface consisting only of six Dirac cones in the first Brillouin zone-one at each (K) over bar. In contrast, angle-resolved photoemission measurements of graphene grown on SiC(0001) often show six satellite Dirac cones surrounding each primary Dirac cone. Recent studies have reported two further Dirac cones along the (Gamma) over bar-(K) over bar line, and argue that these are not photoelectron diffraction artifacts but real bands deriving from a modulation of the ionic potential in the graphene layer. Here we present measurements using linearly polarized synchrotron light which show all of these replicas as well as several additional ones. Using information obtained from dark corridor orientations and angular warping, we demonstrate that all but one of these additional features-including those previously assigned as real initial-state bands-are possible to explain by simple final-state photoelectron diffraction

    On the Filter Narrowing Issues in Elastic Optical Networks

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    This paper describes the problematic filter narrowing effect in the context of next-generation elastic optical networks. First, three possible scenarios are introduced: the transition from an actual fixed-grid to a flexigrid network, the generic full flexi-grid network, and a proposal for a filterless optical network. Next, we investigate different transmission techniques and evaluate the penalty introduced by the filtering effect when considering Nyquist wavelength division multiplexing, single side-band direct-detection orthogonal frequency division multiplexing, and symbol-rate variable dual polarization quadrature amplitude modulation. Also, different approaches to compensate for the filter narrowing effect are discussed. Results show that the specific needs per each scenario can be fulfilled by the aforementioned technologies and techniques or a combination of them, when balancing performance, network reach, and cost

    Spin-orbit coupled spin-polarised hole gas at the CrSe2-terminated surface of AgCrSe2

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    Funding: We gratefully acknowledge support from the European Research Council (through the QUESTDO project, 714193), the Engineering and Physical Sciences Research Council (Grant No. EP/T02108X/1), and the Leverhulme Trust (Grant No. RL-2016-006). S.-J.K., E.A.M., A.Z., and I.M. gratefully acknowledge studentship support from the International Max-Planck Research School for Chemistry and Physics of Quantum Materials. The research leading to this result has been supported by the project CALIPSOplus under the Grant Agreement 730872 from the EU Framework Programme for Research and Innovation HORIZON 2020.In half-metallic systems, electronic conduction is mediated by a single spin species, offering enormous potential for spintronic devices. Here, using microscopic-area angle-resolved photoemission, we show that a spin-polarised two-dimensional hole gas is naturally realised in the polar magnetic semiconductor AgCrSe2 by an intrinsic self-doping at its CrSe2-terminated surface. Through comparison with first-principles calculations, we unveil a striking role of spin-orbit coupling for the surface hole gas, unlocked by both bulk and surface inversion symmetry breaking, suggesting routes for stabilising complex magnetic textures in the surface layer of AgCrSe2.Publisher PDFPeer reviewe
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