Tomographic reconstruction of neopterous Carboniferous insect nymphs

Two new polyneopteran insect nymphs from the Montceau-les-Mines Lagerstätte of France are presented. Both are preserved in three dimensions, and are imaged with the aid of X-ray micro-tomography, allowing their morphology to be recovered in unprecedented detail. One–Anebos phrixos gen. et sp. nov.–is of uncertain affinities, and preserves portions of the antennae and eyes, coupled with a heavily spined habitus. The other is a roachoid with long antennae and chewing mouthparts very similar in form to the most generalized mandibulate mouthparts of extant orthopteroid insects. Computer reconstructions reveal limbs in both specimens, allowing identification of the segments and annulation in the tarsus, while poorly developed thoracic wing pads suggest both are young instars. This work describes the morphologically best-known Palaeozoic insect nymphs, allowing a better understanding of the juveniles’ palaeobiology and palaeoecology. We also consider the validity of evidence from Palaeozoic juvenile insects in wing origin theories. The study of juvenile Palaeozoic insects is currently a neglected field, yet these fossils provide direct evidence on the evolution of insect development. It is hoped this study will stimulate a renewed interest in such work

Carboniferous Onychophora from Montceau‐les‐Mines, France, and onychophoran terrestrialization

Abstract The geological age of the onychophoran crown‐group, and when the group came onto land, have been sources of debate. Although stem‐group Onychophora have been identified from as early as the Cambrian, the sparse record of terrestrial taxa from before the Cretaceous is subject to contradictory interpretations. A Late Carboniferous species from the Mazon Creek biota of the USA, Helenodora inopinata, originally interpreted as a crown‐group onychophoran, has recently been allied to early Cambrian stem‐group taxa. Here we describe a fossil species from the Late Carboniferous Montceau‐les‐Mines Lagerstätte, France, informally referred to as an onychophoran for more than 30 years. The onychophoran affinities of Antennipatus montceauensis gen. nov., sp. nov. are indicated by the form of the trunk plicae and the shape and spacing of their papillae, details of antennal annuli, and the presence of putative slime papillae. The poor preservation of several key systematic characters for extant Onychophora, however, prohibits the precise placement of the Carboniferous fossil in the stem or crown of the two extant families, or the onychophoran stem‐group as a whole. Nevertheless, A. montceauensis is the most compelling candidate to date for a terrestrial Paleozoic onychophoran

The euthycarcinoid arthropods from Montceau-les-Mines, France: functional morphology and affinities

International audienceNew three-dimensionally preserved specimens of two euthycarcinoid arthropods, namely Schramixerxes gerem and Sottyxerxes multiplex, allow complete description of both the dorsal and ventral sides of the exoskeleton. The functional morphology is tentatively interpreted for the first time. In S. gerem, the ‘thirteenth somite', or ‘monosomite' is fully described and re-interpreted as the main articulatory process of the body, between the cephalic region and the preabdomen. The morphology and arrangement of the two parts of the process clearly indicate that the anterior cephalic region of the body could move laterally and bend ventrally, while posterior somites could only move ventrally. Unlike several other euthycarcinoid species, the ventral side of the head area exhibits one or two plates instead of mandibles; such distinct morphologies are indicative of different feeding mechanisms and behaviours among euthycarcinoid arthropods. Possible homologies with the labrum of Hexapoda support the hypothesis that euthycarcinoids have hexapod affinities

Diversity and paleoenvironment of the flora from the nodules of the Montceau-Les-Mines Biota (Late Carboniferous, France)

A-08-13International audienceThe flora from the Montceau-les-Mines Lagerstatte (Massif Central, France, Late Pennsylvanian) is preserved in small sideritic concretions and was studied in three locations in the (1) Saint-Louis, (2) Saint-Francois, and (3) Sainte-Helene opencast mines. Qualitative and quantitative analyses of plant diversity and floristic composition in 6812 nodules indicate substantial variations in the floral composition of these opencast mines. More than 50 taxa are recognized and belong to groups typical of the Late Pennsylvanian flora (lycopsids, sphenopsids, tree ferns, and pteridosperms). Arborescent sphenopsids and tree ferns were the major components at Saint-Louis, whereas the flora from Saint-Francois consisted mainly of pteridosperms; the one from Sainte-Helene has a more balanced composition. Taphonomic and sedimentological data show that the flora contained in the nodules was hypoautochthonous to parautochthonous. The Montceau Basin displayed a mosaic of paleoenvironments (e.g., deltaic lacustrine, paludal to fluvial) which favored colonization by plants and animals

SYNCARID CRUSTACEANS FROM THE MONTCEAU LAGERSTÄTTE (UPPER CARBONIFEROUS; FRANCE)

International audienceKey aspects of the morphology, autecology, systematics and taphonomy of the crustacean syncarids from the Montceau Lagerstätte (Upper Carboniferous, Stephanian B; France) are presented. Palaeocaris secretanae is the most abundant faunal element of the Montceau biota and shows striking morphological similarities with Palaeocaris typus from the Mazon Creek Lagersta¨tte (Westphalian D; Illinois, USA). Palaeocaris secretanae was a shrimp-like animal with a short head (no head shield), large mandibles, 14 trunk segments (the first one being reduced) and a fan-like caudal termination. Both the body and the appendage design indicate abilities for crawling on the substratum (slender endopods) and for escape reaction (uropodal fan, pleonal flexibility), although swimming activities may have been reduced (trunk appendages with small flap-like exopods). Details of the appendages involved in feeding, e.g. mandibles and maxillipeds, indicate poor ability for predation but point to an omnivorous detritus feeding mode. Poorly developed respiratory organs (small cylindrical epipods) suggest a relatively low level of locomotory activity. The field of vision may have been large and panoramic (stalked eyes). Rows of pores on 12 trunk segments are interpreted as possible sensory organs used for current detection. Females were brooding eggs (clusters of eggs preserved along anteroventral trunk). Microprobe analysis indicates that siderite is the major component of the nodules. Four events played a key-role in the three-dimensional preservation of syncarids: (1) rapid burial, (2) minimal decomposition, (3) phosphatic mineralization shortly after the animal's death and (4) nodule formation around the carcass. Palaeocaris secretanae is morphologically close to Recent syncarids such Anaspides tasmaniae (freshwater streams, Australia) in its general body plan and detailed anatomy, e.g. mouth parts, indicating morphological stasis in syncarids over more than 200 million years

The insect nymph <i>Anebos phrixos</i> gen. et sp. nov. MNHN.F.SOT005630 from the Montceau-les-Mines Lagerstätte, France.

<p>A. Dorsal view, showing wingpads. B. Lateral aspect, of note is the orientation of the head and limbs. C. Ventral abdomen demonstrating trilobite underside. D. Anterior view, showing head, antennae and possible maxillary palps, in addition to leg segmentation. Abbreviations: an = antenna; ca = central axis; ce = cerci; e = eye; fe = femur; L1–3 = legs 1–3; lz = lateral zone; mp = maxillary palps (?); ms = mesonotum; mt = metanotum; pn = pronotum; ta = tarsus; ti = tibia; wp = wing pad. All scale bars equal 5 mm.</p

γ -ray spectroscopy of low-lying excited states and shape competition in Os 194

The properties of excited states in the neutron-rich nucleus 194Os have been investigated using the 192Os(18O,16O)194Os reaction with an 80 MeV beam provided by the IFIN-HH Laboratory, Bucharest. Discrete γ -ray decays from excited states have been measured using the hybrid HPGe-LaBr3(Ce) array RoSPHERE. The current work identifies a number of previously unreported low-lying nonyrast states in 194Os as well as the first measurement of the half-life of the yrast 2+ state of 302(50) ps. This is equivalent to a B(E2 : 2+ → 0+) = 45(16) W.u. and intrinsic quadrupole deformation of βeff = 0.14(1). The experimental results are compared with Hartree-Fock-Bogoliubov–interacting-boson-model calculations and are consistent with a reduction in a quadrupole collectivity in Os isotopes with increasing neutron number

γ -ray spectroscopy of low-lying excited states and shape competition in Os 194

The properties of excited states in the neutron-rich nucleus 194Os have been investigated using the 192Os(18O,16O)194Os reaction with an 80 MeV beam provided by the IFIN-HH Laboratory, Bucharest. Discrete γ -ray decays from excited states have been measured using the hybrid HPGe-LaBr3(Ce) array RoSPHERE. The current work identifies a number of previously unreported low-lying nonyrast states in 194Os as well as the first measurement of the half-life of the yrast 2+ state of 302(50) ps. This is equivalent to a B(E2 : 2+ → 0+) = 45(16) W.u. and intrinsic quadrupole deformation of βeff = 0.14(1). The experimental results are compared with Hartree-Fock-Bogoliubov–interacting-boson-model calculations and are consistent with a reduction in a quadrupole collectivity in Os isotopes with increasing neutron number

Preliminary results of proton inelastic scattering on 57^{57}Fe

International audienceA proton inelastic scattering experiment on a 57Fe target was performed at the 9 MV TANDEM facility of Horia Hulubei-National Institute of Physics and Nuclear Engineering, Bucharest-Măgurele, Romania. The purpose was to determine the γ-production cross sections of the transitions observed in the 57Fe(p, p’γ)57Fe reaction. The detection system consisted of 4 HPGe detectors for detection and a Faraday cup for beam integration. The proton energy was varied from 5 to 16 MeV, in 1-MeV steps. In the current analysis, experimental cross sections were determined only for the high-volume detectors, placed at 110◦ and 150◦ with respect to the beam axis. This work presents the data analysis procedure and the preliminary experimental cross sections for the most intense transitions decaying from the first seven excited states in 57Fe. The results are compared with TALYS 1.9 theoretical calculations performed using default input parameters