Evolutionary Physiology: The extent of C4 and CAM photosynthesis in the Genera Anacampseros and Grahamia of the Portulacaceae

The Portulacaceae is one of the few terrestrial plant families known to have both C(4) and Crassulacean acid metabolism (CAM) species. There may be multiple origins of the evolution of CAM within the Portulacaceae but the only clear evidence of C(4) photosynthesis is found in members of the genus Portulaca. In the Portulaca, CAM succulent tissue is overlaid with the C(4) tissue in a unique fashion where both pathways are operating simultaneously. Earlier reports have shown that the clade containing the genera Anacampseros and Grahamia may also contain C(4) photosynthetic species similar to the Portulaca, which would indicate multiple origins of C(4) photosynthesis within the family. The aim of the present study was to ascertain the true photosynthetic nature of these genera. An initial survey of the carbon isotope composition of the Anacampseros ranged from -12.6 per thousand to -24.0 per thousand, indicating very little CAM activity in some species, with other values close to the C(4) range. Anacampseros (=Grahamia) australiana which had been previously identified as a C(4) species had a carbon isotope composition value of -24.0 per thousand, which is more indicative of a C(3) species with a slight contribution of CAM activity. Other Anacampseros species with C(4)-like values have been shown to be CAM plants. The initial isotope analysis of the Grahamia species gave values in the range of -27.1 per thousand to -23.6 per thousand, placing the Grahamia species well towards the C(3) photosynthetic range. Further physiological studies indicated increased night-time CO(2) uptake with imposition of water stress, associated with a large diurnal acid fluctuation and a marked increased phosphoenolpyruvate carboxylase activity. This showed that the Grahamia species are actually facultative CAM plants despite their C(3)-like carbon isotope values. The results indicate that the Grahamia and Anacampseros species do not utilize the C(4) photosynthetic pathway. This is the first to identify that the Grahamia species are facultative CAM plants where CAM can be induced by water stress. This work supports earlier physiological work that indicates that this clade containing Anacampseros and Grahamia species comprises predominantly facultative CAM plants. This report suggests there may be only one clade which contains C(4) photosynthetic members with CAM-like characteristics

Leptons from Dark Matter Annihilation in Milky Way Subhalos

Numerical simulations of dark matter collapse and structure formation show that in addition to a large halo surrounding the baryonic component of our galaxy, there also exists a significant number of subhalos that extend hundreds of kiloparsecs beyond the edge of the observable Milky Way. We find that for dark matter (DM) annihilation models, galactic subhalos can significantly modify the spectrum of electrons and positrons as measured at our galactic position. Using data from the recent Via Lactea II simulation we include the subhalo contribution of electrons and positrons as boundary source terms for simulations of high energy cosmic ray propagation with a modified version of the publicly available GALPROP code. Focusing on the DM DM -> 4e annihilation channel, we show that including subhalos leads to a better fit to both the Fermi and PAMELA data. The best fit gives a dark matter particle mass of 1.2 TeV, for boost factors of 90 in the main halo and 1950-3800 in the subhalos (depending on assumptions about the background), in contrast to the 0.85 TeV mass that gives the best fit in the main halo-only scenario. These fits suggest that at least a third of the observed electron cosmic rays from DM annihilation could come from subhalos, opening up the possibility of a relaxation of recent stringent constraints from inverse Compton gamma rays originating from the high-energy leptons.Comment: 8 pages, 13 figures; added referenc

Neutrinos from supernovae: experimental status and perspectives

I discuss the state of the art in the search for neutrinos from galactic stellar collapses and the future perspectives of this field. The implications for the neutrino physics of a high statistics supernova neutrino burst detection by the network of detectors operating around the world are also reviewed.Comment: 19 pages, 12 figures. Extended version of talk given at IInd International Workshop on Matter, Anti-Matter and Dark Matter, Trento (Italy), 29-30 October 2001. A reduced version will appear in Int. J. of Mod. Phys.

Radion Induced Spontaneous Baryogenesis

We describe a possible scenario for the baryogenesis arising when matter is added on the branes of a Randall-Sundrum model with a radion stabilizing potential. We show that the radion field can naturally induce spontaneous baryogenesis when the cosmological evolution for the matter on the branes is taken into account.Comment: LaTeX 2e, 8 pages and no figures, minor corrections to match version to appear in MPL

Electroweak phase transition in technicolor

Several phenomenologically viable walking technicolor models have been proposed recently. I demonstrate that these models can have first order electroweak phase transitions, which are sufficiently strong for electroweak baryogenesis. Strong dynamics can also lead to several separate transitions at the electroweak scale, with the possibility of a temporary restoration and an extra breaking of the electroweak symmetry. First order phase transitions will produce gravitational waves, which may be detectable at future experiments.Comment: 6 pages, 4 figures. Talk at PASCOS 2010 conference, Valencia, 19-23 July 201

Metastable dark matter mechanisms for INTEGRAL 511 keV γ\gamma rays and DAMA/CoGeNT events

We explore dark matter mechanisms that can simultaneously explain the galactic 511 keV gamma rays observed by INTEGRAL/SPI, the DAMA/LIBRA annual modulation, and the excess of low-recoil dark matter candidates observed by CoGeNT. It requires three nearly degenerate states of dark matter in the 4-7 GeV mass range, with splittings respectively of order an MeV and a few keV. The top two states have the small mass gap and transitions between them, either exothermic or endothermic, can account for direct detections. Decays from one of the top states to the ground state produce low-energy positrons in the galaxy whose associated 511 keV gamma rays are seen by INTEGRAL. This decay can happen spontaneously, if the excited state is metastable (longer-lived than the age of the universe), or it can be triggered by inelastic scattering of the metastable states into the shorter-lived ones. We focus on a simple model where the DM is a triplet of an SU(2) hidden sector gauge symmetry, broken at the scale of a few GeV, giving masses of order \lsim 1 GeV to the dark gauge bosons, which mix kinetically with the standard model hypercharge. The purely decaying scenario can give the observed angular dependence of the 511 keV signal with no positron diffusion, while the inelastic scattering mechanism requires transport of the positrons over distances \sim 1 kpc before annihilating. We note that an x-ray line of several keV in energy, due to single-photon decays involving the top DM states, could provide an additional component to the diffuse x-ray background. The model is testable by proposed low-energy fixed target experiments.Comment: 27 pp, 19 figures; v2. minor clarification, added refs; v3. corrected observed rate of positron production, added new section responding to criticisms of arXiv:0904.1025; v4. corrected typos in eqs. (6) and (40

A coded aperture imaging system optimized for hard X-ray and gamma ray astronomy

A coded aperture imaging system was designed for the Gamma-Ray imaging spectrometer (GRIS). The system is optimized for imaging 511 keV positron-annihilation photons. For a galactic center 511-keV source strength of 0.001 sq/s, the source location accuracy is expected to be + or - 0.2 deg

The Gamma-Ray Imaging Spectrometer (GRIS): A new balloon-borne experiment for gamma-ray line astronomy

High resolution gamma-ray spectroscopy is a relatively new field that holds great promise for further understanding of high energy astrophysical processes. When the high resolution gamma-ray spectrometer (GRSE) was removed from the GRO payload, a balloon program was initiated to permit continued development and improvement of instrumentation in this field, as well as continued scientific observations. The Gamma-Ray Imaging Spectrometer (GRIS) is one of the experiments selected as part of this program. The instrument contains a number of new and innovative features that are expected to produce a significant improvement in source location accuracy and sensitivity over previous balloon and satellite experiments

High Temperature Matter and Gamma Ray Spectra from Microscopic Black Holes

The relativistic viscous fluid equations describing the outflow of high temperature matter created via Hawking radiation from microscopic black holes are solved numerically for a realistic equation of state. We focus on black holes with initial temperatures greater than 100 GeV and lifetimes less than 6 days. The spectra of direct photons and photons from $\pi^0$ decay are calculated for energies greater than 1 GeV. We calculate the diffuse gamma ray spectrum from black holes distributed in our galactic halo. However, the most promising route for their observation is to search for point sources emitting gamma rays of ever-increasing energy.Comment: 33 pages, 13 figures, to be submitted to PR