Soil erosion as affected by shrub encroachment in northeastern Patagonia

La erosion de los suelos es la causa principal de las perdidas irreversibles del potencial productivo de los suelos en la mayoria de los pastizales naturales. En el nordeste de Patagonia, el aumento de la erosion de los suelos ha estado estrechamente asociado al aumento de la cobertura de arbustos en las estepas herbaceas o arbustivas-herbaceas. Nosotros empleamos lluvia simulada para determinar la tasa de infiltracion y la produccion de sedimentos en parches de estepas herbaceas, arbustivas-herbaceas y arbustivas del sitio ecologico Punta Ninfas. Las coberturas de suelo desnudo y de gravas fueron mayores y la cobertura de mantillo menor en la estepa arbustiva respecto a las estepas arbustiva- herbacea y herbacea. En los espacios entre arbustos de la estepa arbustiva, la densidad aparente fue mayor y la macroporosidad y la materia organica fueron menores (P 0.05) que en los monticulos debajo de los arbustos y en las estepas arbustivaherbacea y herbacea. La tasa de infiltration fue un 60 y un 65% mas baja en la estepa arbustiva que en las estepas herbacea y arbustiva-herbacea, respectivamente. Por el contrario, la produccion total y la concentracion de sedimentos fueron mas altas (P s 0.05) en la estepa arbustiva comparado con las estepas herbacea y arbustiva-herbacea. La cobertura de gravas fue la variable que mejor predijo la tasa de infiltracion y la produccion de sedimentos. El contenido de materia organica de los sedimentos, mayormente mantillo, fue similar en la estepa arbustiva y la arbustiva-herbacea y en ambas mayores (P 0.05) que en la estepa herbacea. La remocion de mantillo por el escurrimiento superficial posiblemente represente uno de los procesos que provocan la transicion de la estepa arbustiva-herbacea a la estepa arbustiva. Las altas tasas de remocion de sedimentos, principalmente mantillo, de los espacios entre arbustos de la estepa arbustiva pueden limitar la recuperacion natural de las propiedades fisicas a hidrologicas de los suelos. Estos parches degradados no pueden captar las lluvias incidentes, limitando asi las posibilidades de recuperacion de los pastos perennes y favoreciendo la dominancia de los arbustos.Soil erosion is the primary cause of irreversible loss of soil productivity on most rangelands. In northeastern Patagonia, the increase in soil erosion has been closely associated with the increase in shrub cover in the grass or shrub-grass steppes. We used rainfall simulation to compare infiltration and sediment production from patches of grass, shrub-grass, and shrub steppes of the Punta Ninfas range site. Bare soil and gravel covers were higher and litter cover was lower in the shrub steppe than in the shrub-grass and the grass steppes. In the shrub inter-spaces of the shrub steppe, bulk density was greater and macroporosity and soil organic matter were lower (P less than or equal to 0.05) than in the mounds beneath shrubs and in the grass and shrub-grass areas. Infiltration rate was 60 to 65% lower in the shrub steppe than in the grass and shrub-grass steppes, respectively. On the contrary, total sediment production and concentration were higher (P less than or equal to 0.05) in the shrub steppe as compared to the grass and the shrub-grass areas. Gravel cover was the variable that best predicted infiltration and sediment production. The organic matter content of the sediment, mostly litter, in the shrub and the shrub-grass steppes were similar and greater (P less than or equal to 0.05) than in the grass steppe. Runoff litter removal may represent one of the processes that drive the transition from shrub-grass to shrub steppes. High rates of sediment removal, mainly litter, from the shrub interspaces of the shrub steppe may limit the natural recovery of the soil physical and hydrological properties. These degraded patches fail to capture incident rainfall and restrict the possibilities for the recovery of perennial grasses favoring the dominance of shrubs.Fil: Parizek, B.. Universidad Nacional de la Patagonia "San Juan Bosco"; ArgentinaFil: Rostagno, Cesar Mario. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Centro Nacional Patagónico; ArgentinaFil: Sottini, Roberto Raul. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Santa Fe. Instituto Nacional de Limnología. Universidad Nacional del Litoral. Instituto Nacional de Limnología; Argentin

Effects of habitat and land use on breeding season density of male Asian Houbara Chlamydotis macqueenii

Landscape-scale habitat and land-use influences on Asian Houbara Chlamydotis macqueenii (IUCN Vulnerable) remain unstudied, while estimating numbers of this cryptic, low-density, over-hunted species is challenging. In spring 2013, male houbara were recorded at 231 point counts, conducted twice, across a gradient of sheep density and shrub assemblages within 14,300 km² of the Kyzylkum Desert, Uzbekistan. Four sets of models related male abundance to: (1) vegetation structure (shrub height and substrate); (2) shrub assemblage; (3) shrub species composition (multidimensional scaling); (4) remote-sensed derived land-cover (GLOBCOVER, 4 variables). Each set also incorporated measures of landscape rugosity and sheep density. For each set, multi-model inference was applied to generalised linear mixed models of visit-specific counts that included important detectability covariates and point ID as a random effect. Vegetation structure received strongest support, followed by shrub species composition and shrub assemblage, with weakest support for the GLOBCOVER model set. Male houbara numbers were greater with lower mean shrub height, more gravel and flatter surfaces, but were unaffected by sheep density. Male density (mean 0.14 km-2, 95% CI, 0.12‒0.15) estimated by distance analysis differed substantially among shrub assemblages, being highest in vegetation dominated by Salsola rigida (0.22 [CI, 0.20‒0.25]), high in areas of S. arbuscula and Astragalus (0.14 [CI, 0.13‒0.16] and 0.15 [CI, 0.14‒0.17] respectively), lower (0.09 [CI, 0.08‒0.10]) in Artemisia and lowest (0.04 [CI, 0.04‒0.05]) in Calligonum. The study area was estimated to hold 1,824 males (CI: 1,645‒2,030). The spatial distribution of relative male houbara abundance, predicted from vegetation structure models, had the strongest correspondence with observed numbers in both model-calibration and the subsequent year’s data. We found no effect of pastoralism on male distribution but potential effects on nesting females are unknown. Density differences among shrub communities suggest extrapolation to estimate country- or range-wide population size must take account of vegetation composition

Phylogenetic structure and formation mechanism of shrub communities in arid and semiarid areas of the Mongolian Plateau

The mechanisms of species coexistence within a community have always been the focus in ecological research. Community phylogenetic structure reflects the relationship of historical processes, regional environments, and interactions between species, and studying it is imperative to understand the formation and maintenance mechanisms of community composition and biodiversity. We studied the phylogenetic structure of the shrub communities in arid and semiarid areas of the Mongolian Plateau. First, the phylogenetic signals of four plant traits (height, canopy, leaf length, and leaf width) of shrubs and subshrubs were measured to determine the phylogenetic conservation of these traits. Then, the net relatedness index (NRI) of shrub communities was calculated to characterize their phylogenetic structure. Finally, the relationship between the NRI and current climate and paleoclimate (since the Last Glacial Maximum, LGM) factors was analyzed to understand the formation and maintenance mechanisms of these plant communities. We found that desert shrub communities showed a trend toward phylogenetic overdispersion; that is, limiting similarity was predominant in arid and semiarid areas of the Mongolian Plateau despite the phylogenetic structure and formation mechanisms differing across habitats. The typical desert and sandy shrub communities showed a significant phylogenetic overdispersion, while the steppified desert shrub communities showed a weak phylogenetic clustering. It was found that mean winter temperature (i.e., in the driest quarter) was the major factor limiting steppified desert shrub phylogeny distribution. Both cold and drought (despite having opposite consequences) differentiated the typical desert to steppified desert shrub communities. The increase in temperature since the LGM is conducive to the invasion of shrub plants into steppe grassland, and this process may be intensified by global warming

Immediate effects of microclimate modification enhance native shrub encroachment

Shrubs have become more dense and expanded beyond their range all over the world for a variety of reasons including increased temperatures, overgrazing, and alteration of historical fire regime. Native shrubs have been encroaching on Virginia barrier island grasslands for over half a century for unknown reasons. Species composition, soil nutrients, leaf area index (LAI), and ground and air temperature were recorded across the shrub to grass transition and at free-standing shrubs in a coastal grassland in order to determine the effect of shrub encroachment on plant community and microclimate. Species richness was significantly lower inside shrub thickets. Soil water content, organic matter, nitrogen (N), carbon (C), and LAI were higher in shrub thickets and free-standing shrubs compared to grasslands. Summer and fall maximum temperatures were lower and more moderate where shrubs were present. Fall and winter minimum temperatures were highest inside shrub thickets. Native shrubs impact microclimate and species composition immediately upon encroachment. These shrubs lower overall species composition, increase soil nutrients and moisture, moderate summer temperature, and increase winter temperature, which has consequences on a larger scale. As barrier islands are critical for protecting marsh and mainland habitats, understanding this mechanism for shrub expansion is important to predict future encroachment of shrubs and displacement of grassland habitat

FO-Definability of Shrub-Depth

Shrub-depth is a graph invariant often considered as an extension of tree-depth to dense graphs. We show that the model-checking problem of monadic second-order logic on a class of graphs of bounded shrub-depth can be decided by AC^0-circuits after a precomputation on the formula. This generalizes a similar result on graphs of bounded tree-depth [Y. Chen and J. Flum, 2018]. At the core of our proof is the definability in first-order logic of tree-models for graphs of bounded shrub-depth

Influence of topography and moisture and nutrient availability on green alder function on the low arctic tundra, NT

The Arctic has warmed by at least 3°C over the past 50 years and this rapid warming is expected to continue. Climate warming is driving the proliferation of shrubs across the tundra biome with implications for energy balance, climate, hydrology, nutrient cycling, and biodiversity. Changes in tundra plant water use attributable to shrub expansion are predicted to increase evapotranspirative water loss which may amplify local warming and reduce run-off. However, little is known about the extent to which shrubs will enhance evapotranspirative water loss in these systems. Direct measures of shrub water use are needed to accurately predict evapotranspiration rates and the associated hydrological and energetic impacts. In addition, it is crucial that we understand the abiotic factors that drive shrub distribution and physiological function to forecast further changes in tundra ecosystem function. Shrubs are expanding in areas that have a higher potential of accumulating moisture, such as drainage channels and hill slopes. Shrub expansion may be limited by variation in water and nutrient availability across topographic gradients. Nevertheless, the associations between shrub function and abiotic limitations remain understudied. To address these knowledge gaps, we measured sap flow, stem water potential, and a range of functional traits of green alder (Alnus viridis) shrubs and quantified water and nutrient availability in shrub patches on the low arctic tundra of the Northwest Territories. Frost table depth was a significant negative driver of sap flow and underlies decreased surface water availability with thaw. This was further supported through significantly lower stem water potential values as the growing season progressed. Shrubs in upslope locations had significantly lower water potentials relative to shrubs in downslope locations, demonstrating topographic variation in shrub water status. Shrubs in channels and at the tops of patch slopes significantly differed in leaf functional traits representing leaf investment, productivity, and water use efficiency. Channel shrubs reflected traits associated with higher resource availability and productivity whereas shrubs at the tops of patches reflected the opposite. This work provides insight into the abiotic drivers of tall shrub water use and productivity, both of which will be essential for predicting ecosystem function

Shrub-depth: Capturing Height of Dense Graphs

The recent increase of interest in the graph invariant called tree-depth and in its applications in algorithms and logic on graphs led to a natural question: is there an analogously useful "depth" notion also for dense graphs (say; one which is stable under graph complementation)? To this end, in a 2012 conference paper, a new notion of shrub-depth has been introduced, such that it is related to the established notion of clique-width in a similar way as tree-depth is related to tree-width. Since then shrub-depth has been successfully used in several research papers. Here we provide an in-depth review of the definition and basic properties of shrub-depth, and we focus on its logical aspects which turned out to be most useful. In particular, we use shrub-depth to give a characterization of the lower ${\omega}$ levels of the MSO1 transduction hierarchy of simple graphs